The Ephemeroptera Species of North America and Index to Their Complete Nomenclature

Transcription

1 Transactions of the American Entomological Society 122 (1) pp. 1-54, 1996 The Ephemeroptera Species of North America and Index to Their Complete Nomenclature w. P. MCCAFFERTY Department of Entomology, Purdue University, West Lafayette, Indiana 47907, U.S.A. ABSTRACT A total of 672 species and subspecies of mayflies (Ephemeroptera) are listed for North America (Canada, Mexico, and the continental United States). Changes to the list are as follows: Callibaetis ferrugineus hageni Eaton, 1885, new status [ =C. americanus Banks, 1900, new synonym] and Ca/libaetis ferrugineus ferrugineus (Walsh), 1862, new status, are western and eastern subspecies, respectively; Camelobaetidius arriaga (Traver & Edmunds), 1968 [=C. chiapas (Edmunds & Traver), 1968, new synonym, =C. jenseni (Edmunds & Traver), 1968, new synonym]; Isonychia sicca (Walsh), 1862 [=I. edmundsi Kondratieff & Voshell, 1984, new synonym]; Tortopus pue/la (Pictet); 1843 [=T. incertus (Traver), 1935, new synonym]; and Lachlania saskatchewanensis Ide, 1941 [=L. powelli Edmunds, 1951, new synonym]. Ten species are listed as nomina dubia. A utilitarian, alphabetical index to 1,428 different species names historically used for the North American fauna is provided. The checklist and index serve as an initial basis for a continually updated electronic version available over the internet. The last accounting of the mayfly species of North America was contained in the now out-of-print book by Edmunds et al. (1976). Species and their general distribution were tabulated under general treatments of each genus known at that time, however, no other nomenclatural annotations were provided. The last comprehensive treatment of the North American species, i.e. with species synonymies, was by Traver (1935). That work was essentially limited to species known from north of Mexico, and synonymies were not always complete. The quantity of revisionary work on North American Ephemeroptera since 1976 (e.g., Allen 1980, Bednarik & McCafferty 1979, Flowers 1980, Funk & Sweeney 1994, Kondratieff & Voshell 1984, McCafferty & Waltz 1990, Morihara & McCafferty 1979) and the lack of a recent general account of the entire fauna and its historical nomenclature, have made tracking of the mayfly fauna and its updated classification particularly difficult. This is a serious problem for the many workers who are not mayfly specialists but must deal with mayflies species, whether they be involved in limnological or geographic survey work, freshwater ecological studies, or environmental quality analyses. It is also a special concern of curators of general entomological and zoological collections who must 1

2 2 NORTH AMERICA EPHEMEROPTERA SPECIES manage their Ephemeroptera holdings. The need for modern checklists of all orders of insects and its endorsement by the Entomological Collections Network has been discussed by Morse (1993). The list and index provided here is intended to alleviate the problems discussed above. In addition, and more importantly for the future, it will serve as the initial basis of an electronic version available on the World Wide Web that will be continually updated and will allow all of the convenient search capabilities associated with that medium. The World Wide Web version of the list may be accessed via the Mayfly Central home page. The checklist below is a consecutively numbered accounting of all recognized species and subspecies of mayflies known from Canada, Mexico, and the continental United States of America. The list is formatted alphabetically by genus and species name under families that are arranged phylogenetically under suborders, and infraorders when appropriate, from McCafferty (1991) with modifications from McCafferty & Wang (1994) and Wang & McCafferty (1995). Equivalents of family names beginning with Edmunds et al. (1976) are given in parentheses. To facilitate tracking of all names that have ever been associated with the mayflies of North America, a comprehensive index for all such names follows the checklist. It is a completely alphabetical list of all names, including synonyms, homonyms, genus I species combinations, misspellings, etc., that historically have been used to refer to the North American Ephemeroptera species. Currently accepted names appear in italics, whereas all others are not italicized. All names in the index have a species number from the checklist associated with them that indicates to what currently valid name in the checklist they refer. Authors and dates of publication of the species names are always given, and the convention of placing author names in parentheses when the original generic name has been changed is maintained. The names used in the checklist follow the most recent revisions. The one exception to this involves the family Heptageniidae. Independent cladistic research (McCafferty & Wang, in preparation) has indicated that the radical reduction of the generic nomenclature proposed by Kluge (1988) is inappropriate and would preclude considerable phylogenetic and biological information now imparted by the generic names used in the current binomials. Also, Kluge (1988) potentially created a secondary homonym of Macdunnoa brunnea Flowers, 1982 by recombining the species name with Heptagenia. However, since that revision is not recognized, no homonym exists and no action is necessary. Edmunds & Allen (1957) regarded Callibaetis hageni Eaton, 1885 as a nomen dubium, and Edmunds et al. (1976) later indicated that different specimens assigned to this name belonged to C. americanus Banks, 1900, C. carolus Traver, 1935, and C. coloradensis Banks, McCafferty & Waltz (1990) showed the latter two names to be junior subjective synonyms of C. americanus. Callibaetis ferrugineus hageni, new status, is regarded here as the senior subjective synonym of C. americanus: Callibaetis ferrugineus hageni Eaton [=C. americanus Banks, new synonym]; C. ferrugineus ferrugineus (Walsh), new status, is the eastern subspecies

3 W. P. MCCAFFERTY 3 equivalent to the historical concept of C. ferrugineus, and C. ferrugineus hageni is the western subspecies equivalent to the historical concept of C. americanus and all names that have been synonymized under it (see McCafferty & Waltz 1990). Camelobaetidius arriaga (Traver & Edmunds), 1968, Camelobaetidius chiapas (Edmunds & Traver), 1968, and Camelobaetidius jenseni (Edmunds & Traver), 1968 were based on slight color variants and questionable genitalic differences (Lugo-Ortiz & McCafferty 1995) in adults that now appear to be from the same population of mayflies, all taken from the same locality in Mexico on the same date. Therefore, based on page priority, the latter two names are placed as junior subjective synonyms of Camelobaetidius arriaga (Edmunds & Jensen) [=C. chiapas (Edmunds & Traver), new synonym and =C. jenseni (Traver & Edmunds), new synonym]. Isonychia edmundsi Kondratieff & Voshell, 1984 has no consistent differences with I. sicca (Walsh), Adult coloration differences previously identified for northern (sicca) and Neotropical representatives (edmundsi) grade through Mexico, and no larval differences are known. Isonychia sicca (Walsh) [=I. edmundsi Kondratieff & Voshell, new synonym] is considered a far ranging and variable species. Eaton (1883) incorrectly considered Tortopus puella (Pictet), 1843 to be a junior synonym of Ephoron album (Say), 1824, and Burks (1953) considered the former species to be of dubius status. From Pictet's (1843) short description, key, and the type locality of New Orleans, it can be discerned that the species belongs to Tortopus. Its locality is consistent with that of T. incertus (Traver), 1935, not T. primus (McDunnough), 1924 (see McCafferty 1994). There is now sufficient evidence to consider Tortopus incertus as a subjective junior synonym of Tortopus puella (Pictet) [=T. incertus (Traver), new synonym]. Koss & Edmunds (1970) in Utah, and McCafferty et al. (1993) in Colorado noted the similarity of Lachlania saskatchewanensis Ide, 1941 and L. powelli Edmunds, Because no differences can be found between the two, and because of the now apparent homogeneous distribution, the latter name is here placed as a subjective junior synonym of Lachlania saskatchewanensis Ide [=L. powelli Edmunds, new synonym]. Except for C. hageni and T. puella, the other four species listed as nomina dubia by Edmunds and Allen (1957) remain as such. In addition, Baetis posticatus (Say), 1823, Baetisca callosa Traver, 1931, Callibaetis undatus (Pictet), 1843, Heptagenia manifesta (Eaton), 1885,Ironodes geminatus (Eaton), 1885, and Stenonema flaveolum (Pictet), 1843 (see McCafferty & Bae 1992) are here listed as nomina dubia because the species to which they actually refer cannot be determined. Cloeon cognatum Stephens, 1835 is an adventive species from Europe that until very recently has always been, in North America, incorrectly referred to as C. dipterum (Linneaus), 1761, another European species (see Flowers 1978). For tracking purposes, this historical misidentification is noted by placing the latter name in the index in reference to the former. In no other cases have misidentifications been noted.

4 4 NORTH AMERICA EPHEMEROPTERA SPECIES CHECKLIST Suborder Rectracheata Infraorder Lanceolata LEPTOPHLEBIIDAE 001. Choroterpes albiannulata McDunnough, Choroterpes basalis (Banks), Choroterpes ferruginea Traver, Choroterpes fusca Spieth, Choroterpes hubbelli Berner, Choroterpes inornata Eaton, Choroterpes terratoma Seemann, Farrodes texanus Davis, Habrophlebia vibrans Needham, Habrophlebiodes americana (Banks), Habrophlebiodes annulata Traver, Habrophlebiodes brunneipennis Berner, Habrophlebiodes celeteria Berner, Hydrosmilodon primanus (Eaton), Leptophlebia austrina (Traver), Leptophlebia bradleyi (Needham), Leptophlebia collina (Traver), Leptophlebia cupida (Say), Leptophlebia grandis (Traver), Leptophlebia gravastella (Eaton), Leptophlebia intermedia (Traver), Leptophlebia johnsoni McDunnough, Leptophlebia nebulosa (Walker), Leptophlebia pacifica (McDunnough), Neochoroterpes kossi (Allen), Neochoroterpes oklahoma (Traver), Neochoroterpes nanita (Traver), Neochoroterpes orientalis Henry, Paraleptophlebia adoptiva (McDunnough), Paraleptophlebia altana Kilgore & Allen, Paraleptophlebia aquilina Harper & Harper, Paraleptophlebia assimilis (Banks), Paraleptophlebia associata (McDunnough), Paraleptophlebia bicornuta (McDunnough), Paraleptophlebia bnmneipennis (McDunnough), Paraleptophlebia cachea Day, 1954

5 W. P. MCCAFFERTY Paraleptophlebia calcarita Robotham & Allen, Paraleptophlebia californica Traver, Paraleptophlebia clara (McDunnough), Paraleptophlebia debilis (Walker), Paraleptophlebia falcula Traver, Paraleptophlebia georgiana Traver, Paraleptophlebia gregalis (Eaton), Paraleptophlebia guttata (McDunnough), Paraleptophlebia helena Day, Paraleptophlebia heteronea (McDunnough), Paraleptophlebia jeanae Berner, Paraleptophlebia kirchneri Kondratieff & Durfee, Paraleptophlebia memorialis (Eaton), Paraleptophlebia moerens (McDunnough), Paraleptophlebia mollis (Eaton), Paraleptophlebia ontario (McDunnough), Paraleptophlebia packi (Needham), Paraleptophlebia placeri Mayo, Paraleptophlebia praepedita (Eaton), Paraleptophlebia quisquilia Day, Paraleptophlebia rufivenosa (Eaton), Paraleptophlebia sculleni Traver, Paraleptophlebia sticta Burks, Paraleptophlebia strigula (McDunnough), Paraleptophlebia swannanoa (Traver), Paraleptophlebia temporalis (McDunnough), Paraleptophlebia vaciva (Eaton), Paraleptophlebia volitans (McDunnough), Paraleptophlebia zayante Day, Thraulodes brunneus Koss, Thraulodes ephippiatus Traver & Edmunds, Thraulodes gonzalesi Traver & Edmunds, Thraulodes hilaris (Eaton), Thraulodes humeralis Navas, Thraulodes lunatus Traver & Edmunds, Thraulodes mexicanus (Eaton), Thraulodes packeri Traver & Edmunds, Thraulodes spangleri Traver & Edmunds, Thraulodes speciosus Traver, Thraulodes zonalis Traver & Edmunds, Traverella albertana (McDunnough), Traverella lewisi Allen, Traverella presidiana (Traver), 1934

6 6 NORTH AMERICA EPHEMEROPTERA SPECIES BEHNINGIIDAE 080. Dolania americana Edmunds & Traver, 1959 POT AMANTHIDAE 081. Anthopotamus distinctus (Traver), Anthopotamus myops (Walsh), Anthopotamus neglectus disjunctus Bae & McCafferty, Anthopotamus neglectus neglectus (Traver), Anthopotamus verticis (Say), 1839 POLYMITARCYIDAE (=Euthyplociidae + Polymitarcyidae) 086. Campsurus cuspidatus Eaton, Campsurus decoloratus (Hagen), Ephoron album (Say), Ephoron leukon Williamson, Euthyplocia hecuba (Hagen), Tortopus circumfluus Ulmer, Tortopus primus (McDunnough), Tortopus puella (Pictet), 1843 EPHEMERIDAE (=Ephemeridae + Palingeniidae) 094. Ephemera blanda Traver, Ephemera compar Hagen, Ephemera guttulata Pictet, Ephemera simulans Walker, Ephemera traverae Spieth, Ephemera triplex Traver, Ephemera varia Eaton, Hexagenia albivitta (Walker), Hexagenia atrocaudata McDunnough, Hexagenia bilineata (Say), Hexagenia limbata (Serville), Hexagenia mexicana Eaton, Hexagenia orlando Traver, Hexagenia rigida McDunnough, Litobrancha recurvata (Morgan), Pentagenia robusta McDunnough, Pentagenia vittigera (Walsh), 1862

7 W. P. MCCAFFERTY 7 Infraorder Pannota BAETISCIDAE 111. Baetisca becki Schneider & Berner, Baetisca berneri Tarter & Kirchner, Baetisca callosa Traver, 1931 nomen dubium 114. Baetisca carolina Traver, Baetisca columbiana Edmunds, Baetisca escambiensis Berner, Baetisca gibbera Berner, Baetisca lacustris McDunnough, Baetisca laurentina McDunnough, Baetisca obesa (Say), Baetisca rogersi Berner, Baetisca rubescens (Provancher), 1878 NEOEPHEMERIDAE 123. Neoephemera bicolor McDunnough, Neoephemera compressa Berner, Neoephemera purpurea (Traver), Neoephemera youngi Berner, 1953 CAENIDAE 127. Amercaenis ridens (McDunnough), Brachycercus arcticus Soldan, Brachycercus berneri Soldan, Brachycercus edmundsi Soldan, Brachycercus flavus Traver, Brachycercus fioridicola Soldan, Brachycercus lacustris (Needham), Brachycercus maculatus Berner, Brachycercus nasutus Soldan, Brachycercus nitidus (Traver), Brachycercus pini Soldan, Brachycercus prudens (McDunnough), Brachycercus tuberculatus Soldan, Caenis amica Hagen, Caenis anceps Traver, Caenis bajaensis Allen & Murvosh, Caenis candida Harper & Harper, Caenis diminuta diminuta Walker, Caenis hilaris (Say), Caenis latipennis Banks, Caenis macafferti Provonsha, Caenis punctata McDunnough, Caenis tardata McDunnough, 1931

8 8 NORTH AMERICA EPHEMEROPTERA SPECIES 150. Caenis youngi Roemhild, Cercobrachys etowah Soldan, Cercobrachys serpentis Soldan, 1986 EPHEMERELLIDAE 153. Attenella attenuata (McDunnough), Attenella delantala (Mayo), Attenella margarita (Needham), Attenella soquele (Day), Caudatella cascadia (Allen & Edmunds), Caudatella edmundsi (Allen), Caudatella heterocaudata californica (Allen & Edmunds), Caudatella heterocaudata circia (Allen & Edmunds), Caudatella heterocaudata heterocaudata (McDunnough), Caudatella hystrix (Traver), Caudatella jacobi (McDunnough), Caurinella idahoensis Allen, Drunella allegheniensis (Traver), Drunella coloradensis (Dodds), Drunella cornuta (Morgan), Drunella cornutella (McDunnough), Drunella doddsi (Needham), Drunella Jlavilinea (McDunnough), Drunella grandis flavitincta (McDunnough), Drunella grandis grandis (Eaton), Drunella grandis ingens (McDunnough), Drunella lata (Morgan), Drunella longicornis (Traver), Dnmella pelosa (Mayo), Dnmella spinifera (Needham), Drunella tuberculata (Morgan), Drunella walkeri (Eaton), Drunella wayah (Traver), Ephemerella alleni Jensen & Edmunds, Ephemerella altana Allen, Ephemerella apopsis McCafferty, Ephemerella argo Burks, Ephemerella aurivillii (Bengtsson), Ephemerella berneri Allen & Edmunds, Ephemerella catawba Traver, Ephemerella choctawhatchee Berner, Ephemerella consimilis Walsh, 1862 nomen dubium 190. Ephemerella crenula Allen & Edmunds, Ephemerella dorothea Needham, 1908

9 W. P. MCCAFFERTY Ephemerella excrucians Walsh, Ephemerella floripara McCafferty, Ephemerella fratercula McDunnough, Ephemerella hispida Allen & Edmunds, Ephemerella inconstans Traver, Ephemerella inermis Eaton, Ephemerella infrequens McDunnough, Ephemerella invaria (Walker), Ephemerella lacustris Allen & Edmunds, Ephemerella maculata Traver, Ephemerella moffatae Allen, Ephemerella mollitia Seemann, Ephemerella mucronata (Bengtsson), Ephemerella needhami McDunnough, Ephemerella ora Burks, Ephemerella quebecensis (Provancher), 1876 nomen dubium 208. Ephemerella rama Allen, Ephemerella rossi Allen & Edmunds, Ephemerella rotunda Morgan, Ephemerella septentrionalis McDunnough, Ephemerella simila Allen & Edmunds, Ephemerella subvaria McDunnough, Ephemerella unicornis Needham, 1905 nomen dubium 215. Ephemerella verruca Allen & Edmunds, Eurylophella aestiva (McDunnough), Eurylophella bartoni (Allen), Eurylophella bicolor (Clemens), Eurylophella bicoloroides (McDunnough), Eurylophella coxalis (McDunnough), Eurylophella doris (Traver), Eurylophella enoensis Funk, Eurylophella funeralis (McDunnough), Eurylophella lodi (Mayo), Eurylophella lutulenta (Clemens), Eurylophella macdunnoughi Funk, Eurylophella minimella (McDunnough), Eurylophella poconoensis Funk, Eurylophella prudentalis (McDunnough), Eurylophella temporalis (McDunnough), Eurylophella versimilis (McDunnough), Serratella carolina (Berner & Allen), Serratella deficiens (Morgan), Serratella frisoni (McDunnough), Serratella levis (Day), Serratella micheneri (Traver), Serratella molita (McDunnough), Serratella sequoia (Allen & Collins), Serratella serrata (Morgan), 1911

10 10 NORTH AMERICA EPHEMEROPTERA SPECIES 240. Serratella serratoides (McDunnough), Serratella sordida (McDunnough), Serratella spiculosa (Bemer & Allen), Serratella teresa (Traver), Serratella tibialis (McDunnough), Serratella velmae (Allen & Edmunds), Timpanoga hecuba hecuba (Eaton), Timpanoga hecuba pacifica (Allen & Edmunds), Timpanoga Zita (Burks), Timpanoga provonshai (McCafferty), Timpanoga simplex (McDunnough), 1925 LEPTOHYPHIDAE (=Tricorythidae, subfamily Leptohyphinae) 251. Leptohyphes alleni Brusca, Leptohyphes apache Allen, Leptohyphes berneri Traver, Leptohyphes brunneus Allen & Brusca, Leptohyphes canstaneus Allen, Leptohyphes dicinctus Allen & Brusca, Leptohyphes dolani Allen, Leptohyphes ferruginus Allen & Brusca, Leptohyphes hispidus Allen & Brusca, Leptohyphes lestes Allen & Brusca, Leptohyphes michaeli Allen, Leptohyphes mints Allen, Leptohyphes nigripunctus Traver, Leptohyphes packeri Allen, Leptohyphes paraguttatus Allen, Leptohyphes pilosus Allen & Brusca, Leptohyphes quercus Kilgore & Allen, Leptohyphes robacki Allen, Leptohyphes sabinus Traver, Leptohyphes spiculatus Allen & Brusca, Leptohyphes succinus Allen, Leptohyphes tarsos Allen & Murvosh, Leptohyphes vescus Allen, Leptohyphes zalope Traver, Tricorythodes albilineatus Bemer, Tricorythodes allectus (Needham), Tricorythodes angulatus Traver, Tricorythodes atratus (McDunnough), Tricorythodes comus Traver, Tricorythodes condylus Allen, Tricorythodes corpulentus Kilgore & Allen, Tricorythodes curvatus Allen, Tricorythodes dimorphus Allen, 1967

11 W. P. MCCAFFERTY Tricorythodes edmundsi Allen, Tricorythodes explicatus (Eaton), Tricorythodes fictus Traver, Tricorythodes minutus Traver, Tricorythodes mulaiki Traver, Tricorythodes notatus Allen & Brusca, Tricorythodes peridius Burks, Tricorythodes stygiatus McDunnough, Tricorythodes texanus Traver, Tricorythodes ulmeri Allen & Brusca, 1973 Suborder Pisciforma ACANTHAMETROPODIDAE (=Acanthametropodidae + Analetrididae) (=Siphlonuridae, subfamily Acanthametropodinae) 294. Acanthametropus pecatonica (Burks), Analetris eximia Edmunds, 1972 AMELETIDAE (=Siphlonuridae, subfamily Siphlonurinae in part) 296. Ameletus aequivocus McDunnough, Ameletus alticolus McDunnough, Ameletus amador Mayo, Ameletus browni McDunnough, Ameletus celer McDunnough, Ameletus celeroides McDunnough, Ameletus connectinus McDunnough, Ameletus connectus McDunnough, Ameletus cooki McDunnough, Ameletus cryptostimulus Carle, Ameletus dissitus Eaton, Ameletus exquisitus Eaton, Ameletus facilis Day, Ameletus falsus McDunnough, Ameletus imbellis Day, Ameletus lineatus Traver, Ameletus ludens Needham, Ameletus mantis Mayo, Ameletus oregonensis McDunnough, Ameletus querulus McDunnough, Ameletus shepherdi Traver, Ameletus similior McDunnough, Ameletus sparsatus McDunnough, Ameletus subnotatus Eaton, Ameletus suffusus McDunnough, Ameletus tarteri Burrows, Ameletus tertius McDunnough, Ameletus tuberculatus McDunnough, 1939

12 12 NORTH AMERICA EPHEMEROPTERA SPECIES 324. Ameletus validus McDunnough, Ameletus vancouverensis McDunnough, Ameletus velox Dodds, Ameletus vernalis McDunnough, Ameletus walleyi Harper, 1970 AMETROPODIDAE 329. Ametropus albrighti Traver, Ametropus ammophilus Allen & Edmunds, Ametropus neavei McDunnough, 1928 BAETIDAE 332. Acentrella ampla Traver, Acentrella insignificans (McDunnough), Acentrella lapponica Bengtsson, Acentrella turbida (McDunnough), Acerpenna akataleptos (McDunnough), Acerpenna harti (McDunnough), Acerpenna macdunnoughi (Ide), Acerpenna pletura Lugo-Ortiz & McCafferty, Acerpenna pygmaea (Hagen), Apobaetis etowah (Traver), Apobaetis indeprensus Day, Baetis adonis Traver, Baetis alachua (Berner), Baetis alius Day, Baetis armillatus McCafferty & Waltz, Baetis bicaudatus Dodds, Baetis bimaculatus (Berner), Baetis brunneicolor McDunnough, Baetis bundyae Lehmkuhl, Baetis caelestis Allen & Murvosh, Baetis caurinus Edmunds & Allen, Baetis cinctutus McCafferty & Waltz, Baetis diablus Day, Baetis dubius (Walsh), Baetis elliotti (Daggy), Baetis flavistriga McDunnough, Baetis Joemina McDunnough, Baetis futile (McDunnough), Baetis hudsonicus Ide, Baetis intercalaris McDunnough, Baetis magnus McCafferty & Waltz, Baetis moffati Dodds, Baetis notos Allen & Murvosh, 1987

13 W. P. MCCAFFERTY Baetis palisadi Mayo, Baetis parallelus Banks, Baetis persecutus McDunnough, Baetis piscatoris Traver, Baetis pluto McDunnough, Baetis posticatus (Say), 1823 nomen dubium 371. Baetis punctiventris (McDunnough), Baetis rubrolaterale (McDunnough), Baetis rusticans McDunnough, Baetis sulfurosus Day, Baetis thermophilos McDunnough, Baetis tricaudatus Dodds, Baetis veteris (McDunnough), Baetis virile (McDunnough), Baetodes adustus Cohen & Allen, Baetodes alleni McCafferty & Provonsha, Baetodes arizonensis Koss, Baetodes bibranchius McCafferty & Provonsha, Baetodes caritus Cohen & Allen, Baetodes deficiens Cohen & Allen, Baetodes edmundsi Koss, Baetodes fortinensis Mayo, Baetodes fuscipes Cohen & Allen, Baetodes inermis Cohen & Allen, Baetodes longus Mayo, Baetodes obesus Mayo, Baetodes pallidus Cohen & Allen, Baetodes pictus Cohen & Allen, Baetodes tritus Cohen & Allen, Barbaetis benfieldi Kennedy, Barbaetis cestus (Provonsha & McCafferty), Callibaetis californicus Banks, Callibaetis centralis Peters, Callibaetis ferrugineus ferrugineus (Walsh), 1862 new status 399. Callibaetis ferrugineus hageni Eaton, 1885 new status 400. Callibaetis floridanus Banks, Callibaetis fluctuans (Walsh), Callibaetis montanus Eaton, Callibaetis pallidus Banks, Callibaetis pictus (Eaton), Callibaetis pretiosus Banks, Callibaetis punctilusus McCafferty & Provonsha, Callibaetis skokianus Needham, Callibaetis undatus (Pictet), 1843 nom~n dubium

14 14 NORTH AMERICA EPHEMEROPTERA SPECIES 409. Camelobaetidius arriaga (Traver & Edmunds), Camelobaetidius mexicanus (Traver & Edmunds), Camelobaetidius musseri (Traver & Edmunds), Camelobaetidius similis Lugo-Ortiz & McCafferty, Camelobaetidius sinaloa (Allen & Murvosh), Camelobaetidius waltzi Mccafferty, Camelobaetidius warreni (Traver & Edmunds), Centroptilum alamance (Traver), Centroptilum bifurcatum McDunnough, Centroptilum selanderorum Edmunds, Centroptilum semirufum McDunnough, Centroptilum triangulifer (McDunnough), Centroptilum victoriae McDunnough, Cloeodes excogitatus Waltz & McCafferty, Cloeodes macrolamellus Waltz & McCafferty, Cloeodes peninsulus Waltz & McCafferty, Cloeon cognatum Stephens, Diphetor devinctus (Traver), Diphetor hageni (Eaton), Fallceon eatoni (Kimmins), Fallceon longifolius (Kluge), Fallceon quilleri (Dodds), Heterocloeon berneri (Miiller-Liebenau), Heterocloeon curiosum (McDunnough), Heterocloeon frivolum (McDunnough), Heterocloeon petersi (Miiller-Liebenau), Labiobaetis apache McCafferty & Waltz, Labiobaetis dardanus (McDunnough), Labiobaetis ephippiatus (Traver), Labiobaetis frondalis (McDunnough), Labiobaetis longipalpus (Morihara & McCafferty), Labiobaetis propinquus (Walsh), Moribaetis macaferti Waltz, Paracloeodes minutus (Daggy), Procloeon album (McDunnough), Procloeon asperatum (Traver), Procloeon bellum (McDunnough), Procloeon caliginosum (McDunnough), Procloeon conturbatum (McDunnough), 1929

15 W. P. MCCAFFERTY Procloeon convexum (Ide), Procloeon elsa (Traver), Procloeonfragile (McDunnough), Procloeon hobbsi (Berner), Procloeon implicatum (McDunnough), Procloeon inanum (McDunnough), Procloeon ingens (McDunnough), Procloeon insignificans (McDunnough), Procloeon intermediale (McDunnough), Procloeon irrubrum Lowen & Flannagan, Procloeon mendax (Walsh), Procloeon minor (McDunnough), Procloeon oreophilum (Edmunds), Procloeon ozburni (McDunnough), Procloeon pennulatum (Eaton), Procloeon quaesitum (McDunnough), Procloeon rivulare (Traver), Procloeon rubropictum (McDunnough), Procloeon rufostrigatum (McDunnough), Procloeon simile (McDunnough), Procloeon simplex (McDunnough), Procloeon texanum McCafferty & Provonsha, Procloeon venosum (Traver), Procloeon vicinum (Hagen), Procloeon viridocularis (Berner), Procloeon walshi (McDunnough), Pseudocentroptiloides usa Waltz & McCafferty, 1989 METRETOPODIDAE 475. Metretopus alter Bengtsson, Metretopus borealis (Eaton), Siphloplecton basale (Walker), Siphloplecton brunneum Berner, Siphloplecton costalense Spieth, Siphloplecton fuscum Berner, Siphloplecton interlineatum (Walsh), Siphloplecton simile Berner, Siphloplecton speciosum Traver, 1932 SIPHLONURIDAE (=Siphlonuridae, subfamily Siphlonurinae in part) 484. Edmundsius agilis Day, Parameletus chelifer Bengtsson, Parameletus columbiae McDunnough, Parameletus croesus (McDunnough), 1923

16 16 NORlH AMERICA EPHEMEROPTERA SPECIES 488. Parameletus midas (McDunnough), Siphlonisca aerodromia Needham, Siphlonurus alternatus (Say), Siphlonurus autumnalis McDunnough, Siphlonurus barbaroides McDunnough, Siphlonurus barbarus McDunnough, Siphlonurus columbianus McDunnough, Siphlonurus decants Traver, Siphlonurus demaryi Kondratieff & Voshell, Siphlonurus luridipennis (Burmeister), Siphlonurus marginatus Traver, Siphlonurus marshalli Traver, Siphlonurus minnoi Provonsha & McCafferty, Siphlonurus mints (Eaton), Siphlonurus noveboracana (Lichtenstein), 1796 nomen dubium 503. Siphlonurus occidentalis (Eaton), Siphlonurus phyllis McDunnough, Siphlonurus quebecensis (Provancher), Siphlonurus rapidus McDunnough, Siphlonurus securifer McDunnough, Siphlonurus spectabilis Traver, Siphlonurus typicus (Eaton), 1885 Suborder Setisura ISONYCHIIDAE ( =Oligoneuriidae, subfamily Isonychiinae) (=Siphlonuridae, subfamily Isonychiinae) 510. Isonychia arida (Say), Isonychia berneri Kondratieff & Voshell, Isonychia bicolor (Walker), Isonychia campestris McDunnough, Isonychia diversa Traver, Isonychia georgiae McDunnough, Isonychia hoffmani Kondratieff & Voshell, Isonychia intermedia (Eaton), Isonychia notata Traver, Isonychia obscura Traver, Isonychia nifa McDunnough, Isonychia sayi Burks, Isonychia serrata Traver, Isonychia sicca (Walsh), Isonychia similis Traver, Isonychia tusculanensis Berner, Isonychia velma Needham, 1932

17 W. P. MCCAFFERTY 17 OLIGONEURIIDAE 527. Homoeoneuria alleni Pescador & Peters, Homoeoneuria ammophila (Spieth), Homoeoneuria cahabensis Pescador & Peters, Homoeoneuria dolani Edmunds, Berner & Traver, Homoeoneuria salviniae Eaton, Lachlania dencyanna Koss, Lachlania iops Allen & Chao, Lachlania saskatchewanensis Ide, 1941 PSEUDIRONIDAE ( =Heptageniidae, subfamily Pseudironinae) (=Siphlonuridae, subfamily Pseudironinae) 535. Pseudiron centralis McDunnough, 1931 ARTHROPLEIDAE (=Heptageniidae, subfamily Arthropleinae) 536. Arthroplea bipunctata (McDunnough), 1924 HEPTAGENIIDAE (=Heptageniidae, subfamily Heptageniinae) 537. Acanthomola pubescens Whiting & Lehmkuhl, Anepeorus rusticus McDunnough, Anepeorus simplex (Walsh), Cinygma dimicki McDunnough, Cinygma integrum Eaton, Cinygma lyriformis (McDunnough), Cinygmula confusa (McDunnough), Cinygmula gartrelli McDunnough, Cinygmula kootenai McDunnough, Cinygmula mimus (Eaton), Cinygmula par (Eaton), Cinygmula ramaleyi (Dodds), Cinygmula reticulata McDunnough, Cinygmula subaequalis (Banks), Cinygmula tarda (McDunnough), Cinygmula tioga Mayo, Cinygmula uniformis McDunnough, Epeorus albertae (McDunnough), Epeorus deceptivus (McDunnough), Epeorus dispar (Traver), Epeorus dulciana (McDunnough), 1935

18 18 NORTH AMERICA EPHEMEROPTERA SPECIES 558. Epeorus fragilis (Morgan), Epeorus frisoni (Burks), Epeorus grandis (McDunnough), Epeorus hesperus (Banks), Epeorus lagunitas (Traver), Epeorus longimanus (Eaton), Epeorus margarita Edmunds & Allen, Epeorus metlacensis Traver, Epeorus namatus (Burks), Epeorus permagnus (Traver), Epeorus pleuralis (Banks), Epeorus punctatus (McDunnough), Epeorus rubidus (Traver), Epeorus subpallidus (Traver), Epeorus suffusus (McDunnough), Epeorus vitreus (Walker), Heptagenia adequata McDunnough, Heptagenia culacantha Evans, Botts & Flowers, Heptagenia diabasia Burks, Heptagenia dolosa Traver, Heptagenia elegantula (Eaton), Heptagenia fiavescens (Walsh), Heptagenia julia Traver, Heptagenia manifesta (Eaton), 1885 nomen dubium 582. Heptagenia marginalis Banks, Heptagenia patoka Burks, Heptagenia pulla (Clemens), Heptagenia solitaria McDunnough, Heptagenia townesi Traver, Ironodes arctus Traver, Ironodes californicus (Banks), Ironodes fiavipennis Traver, Ironodes geminatus (Eaton), 1885 nomen dubium 591. Ironodes lepidus Traver, Ironodes nitidus (Eaton), Leucrocuta aphrodite (McDunnough), Leucrocuta hebe (McDunnough), Leucrocuta jewetti (Allen), Leucrocuta juno (McDunnough), Leucrocuta maculipennis (Walsh), Leucrocuta minerva (McDunnough), Leucrocuta petersi (Allen), Leucrocuta thetis (Traver), Leucrocuta umbratica (McDunnough), Leucrocuta walshi (McDunnough), 1926

19 W. P. MCCAFFERTY Macdunnoa brunnea Flowers, Macdunnoa nipawinia Lehmkuhl, Macdunnoa persimplex (McDunnough), Nixe bella (Allen & Cohen), Nixe criddlei (McDunnough), Nixe flowersi McCafferty, Nixe horrida (McDunnough), Nixe inconspicua (McDunnough), Nixe kennedyi (McDunnough), Nixe lucidipennis (Clemens), Nixe otiosa (McDunnough), Nixe perfida (McDunnough), Nixe rodocki (Traver), Nixe rosea (Traver), Nixe rusticalis (McDunnough), Nixe salvini (Kimmins), Nixe simplicioides (McDunnough), Nixe spinosa (Traver), Raptoheptagenia cruentata (Walsh), Rhithrogena amica Traver, Rhithrogena anomala McDunnough, Rhithrogena brunneotincta McDunnough, Rhithrogena decora Day, Rhithrogena exilis Traver, Rhithrogena fasciata Traver, Rhithrogena flavianula (McDunnough), Rhithrogena fuscifrons Traver, Rhithrogena futilis McDunnough, Rhithrogena gaspeensis McDunnough, Rhithrogena hageni Eaton, Rhithrogena impersonata (McDunnough), Rhithrogena jejuna Eaton, Rhithrogena morrisoni (Banks), Rhithrogena notialis Allen & Chao, Rhithrogena pellucida Daggy, Rhithrogena plana Allen & Chao, Rhithrogena robusta Dodds, Rhithrogena rubicunda Traver, Rhithrogena uhari Traver, Rhithrogena undulata (Banks), Rhithrogena virilis McDunnough, Rhithrogena vitta Allen & Chao, Stenacron candidum (Traver), Stenacron carolina (Banks), Stenacron floridense (Lewis), 1974

20 20 NORTH AMERICA EPHEMEROPTERA SPECIES 648. Stenacron gildersleevei (Traver), Stenacron interpunctatum (Say), Stenacron minnetonka (Daggy), Stenacron pallidum (Traver), Stenonema bednariki McCafferty, Stenonema carlsoni Lewis, Stenonema exiguum Traver, Stenonema femoratum (Say), Stenonema flaveolum (Pictet), 1843 nomen dubium 657. Stenonema ithaca (Clemens & Leonard), Stenonema lenati McCafferty, Stenonema luteum (Clemens), Stenonema mediopunctatum arwini Bednarik & McCafferty, Stenonema mediopunctatum mediopunctatum (McDunnough), Stenonema meririvulanum Carle & Lewis, Stenonema mexicanum integrum (McDunnough), Stenonema mexicanum mexicanum (Ulmer), Stenonema modestum (Banks), Stenonema pudicum (Hagen), Stenonema pulchellum (Walsh), Stenonema sinclairi Lewis, Stenonema smithae Traver, Stenonema terminatum placitum (Banks), Stenonema terminatum terminatum (Walsh), Stenonema vicarium (Walker), 1853 INDEX OF SPECIES NAME EQUIVALENTS (Only currently accepted names in italics) Acanthametropus pecatonica (Burks), Acanthomola pubescens Whiting & Lehmkuhl, Acentrella ampla Traver, Acentrella carolina (Banks), Acentrella insignificans (McDunnough), Acentrella lapponica Bengtsson, Acentrella turbida (McDunnough), Acerpenna akataleptos (McDunnough), Acerpenna harti (McDunnough), Acerpenna macdunnoughi (Ide), Acerpenna pletura Lugo-Ortiz & McCafferty, Acerpenna pygmaea (Hagen), Ameletus aequivocus McDunnough, Ameletus alticolus McDunnough,

21 W. P. MCCAFFERTY 21 Ameletus amador Mayo, Ameletus browni McDunnough, Ameletus celer McDunnough, Ameletus celeroides McDunnough, Ameletus connectina McDunnough, Ameletus connectinus McDunnough, Ameletus connectus McDunnough, Ameletus cooki McDunnough, Ameletus cryptostimulus Carle, Ameletus dissitus Eaton, Ameletus exquisitus Eaton, Ameletus facilis Day, Ameletus falsus McDunnough, Ameletus imbellis Day, Ameletus lineatus Traver, Ameletus ludens Needham, Ameletus monta Mayo, Ameletus mantis Mayo, Ameletus oregonensis McDunnough, Ameletus querulus McDunnough, Ameletus shepherdi Traver, Ameletus similior McDunnough, Ameletus sparsatus McDunnough, Ameletus subnotatus Eaton, Ameletus suffusus McDunnough, Ameletus tarteri Burrows, Ameletus tertius McDunnough, Ameletus tuberculatus McDunnough, Ameletus validus McDunnough, Ameletus vancouverensis McDunnough, Ameletus velox Dodds, Ameletus vernalis McDunnough, Ameletus walleyi Harper, Amercaenis ridens (McDunnough), Ametropus albrighti Traver, Ametropus ammophilus Allen & Edmunds, Ametropus neavei McDunnough, Analetris eximia Edmunds, Anepeorus rusticus McDunnough, Anepeorus simplex (Walsh), Anthopotamus diaphanus (Needham), Anthopotamus distinctus (Traver), Anthopotamus inequalis (Needham), Anthopotamus myops (Walsh),

22 22 NORTH AMERICA EPHEMEROPTERA SPECIES Anthopotamus neglectus (Traver), Anthopotamus neglectus disjunctus Bae & McCafferty, Anthopotamus neglectus neglectus (Traver), Anthopotamus rufous (Argo), Anthopotamus verticis (Say), Anthopotamus walkeri (Ide), Apobaetis etowah (Traver), Apobaetis indeprensus Day, Arthroplea bipunctata (McDunnough), Attenella attenuata (McDunnough), Attenella delantala (Mayo), Attenella margarita (Needham), Attenella soquele (Day), Baetis adonis Traver, Baetis akataleptos McDunnough, Baetis alachua (Berner), Baetis alba Say, Baetis albivitta Walker, Baetis albus Say, Baetis alius Day, Baetis alternata Say, Baetis alternatus Say, Baetis amplus (Traver), Baetis anachris Burks, Baetis annulata Walker, Baetis annulatus Walker, Baetis arida Say, Baetis armillatus McCafferty & Waltz, Baetis australis Traver, Baetis baeticatus Burks, Baetis basalis Walker, Baetis bicaudatus Dodds, Baetis bilineata Say, Baetis bimaculatus (Berner), Baetis bnmneicolor McDunnough, Baetis buenoi Allen, Baetis bundyae Lehmkuhl, Baetis bundyi Lehmkuhl, Baetis byblis Allen & Murvosh, Baetis caelestis Allen & Murvosh, Baetis canadensis Walker, Baetis caurinus Edmunds & Allen, Baetis cinctutus McCafferty & Waltz, Baetis cingulatus McDunnough, Baetis cingulatus (McDunnough),

23 W. P. MCCAFFERTY 23 Baetis cleptis Burks, Baetis dardanus McDunnough, Baetis debilis Walker, Baetis debilis Walsh, Baetis devinctus Traver, Baetis diablus Day, Baetis dubius (Walsh), Baetis eatoni Kimmins, Baetis elachistus Burks, Baetis elliotti (Daggy), Baetis endymion Traver, Baetis ephippiatus Traver, Baetis erebus Traver, Baetis femorata Provancher, Baetis femorata Say, Baetis femorata Walsh, Baetis flaveola Pictet, Baetis flavistriga McDunnough, Baetis fluctuans (Walsh), Baetis foemina McDunnough, Baetis frivolus McDunnough, Baetis frondalis McDunnough, Baetis fusca Walker, Baetis fuscata Walker, Baetis futile (McDunnough), Baetis hageni Eaton, Baetis harti McDunnough, Baetis herodes Burks, Baetis hiemalis Leonard, Baetis hudsonicus Ide, Baetis ignava Hagen, Baetis incertans McDunnough, Baetis insignificans McDunnough, Baetis intercalaris McDunnough, Baetis interlineata Walsh, Baetis intermedius Dodds, Baetis interpunctata Say, Baetis invaria Walker, Baetis jesmondensis McDunnough, Baetis lapponicus (Bengtsson), Baetis lasallei Banks, Baetis leechi Burks, Baetis levitans McDunnough, Baetis libos Allen & Murvosh, Baetis longifolius Kluge, Baetis longipalpus Morihara & McCafferty, Baetis luridipennis Burmeister, Baetis macani bundyae Lehmkuhl, Baetis macdunnoughi Ide,

24 24 NORTH AMERICA EPHEMEROPTERA SPECIES Baetis ma gnus McCafferty & Waltz, Baetis minimus Dodds, Baetis moffati Dodds, Baetis nanus McDunnough, Baetis notos Allen & Murvosh, Baetis noveboracana (Lichtenstein), Baetis obesa Say, Baetis ochris Burks, Baetis palisadi Mayo, Baetis pallidulus McDunnough, Baetis parallelus Banks, Baetis parvulus (McDunnough), Baetis parvus Dodds, Baetis persecutus McDunnough, Baetis phoebus McDunnough, Baetis phyllis Burks, Baetis pictus Eaton, Baetis piscatoris Traver, Baetis pluto McDunnough, Baetis posticatus (Say), Baetis propinquus (Walsh), Baetis punctiventris (McDunnough), Baetis pygmaeus (Hagen), Baetis quebecensis Hubbard, Baetis quilleri Dodds, Baetis rubescens (Provancher), Baetis rubrolaterale (McDunnough), Baetis rusticans McDunnough, Baetis sicca Walsh, Baetis sonora Allen & Murvosh, Baetis sp. A Morihara & McCafferty, Baetis sp. B Morihara & McCafferty, Baetis sp. C Morihara & McCafferty, Baetis sp. No. 1 Traver, Baetis spiethi Berner, Baetis spinosus McDunnough, Baetis sulfurosus Day, Baetis tessellata Hagen, Baetis tessellata Walker, Baetis thermophilos McDunnough, Baetis tricaudatus Dodds, Baetis unicolor (Hagen), Baetis vagans McDunnough, Baetis verticis Say, Baetis veteris (McDunnough), Baetis vicaria Walker, Baetis virile (McDunnough), Baetisca bajkovi Neave,

25 W. P. MCCAFFERTY 25 Baetisca becki Schneider & Berner, Baetisca berneri Tarter & Kirchner, Baetisca callosa Traver, Baetisca carolina Traver, Baetisca columbiana Edmunds, Baetisca escambiensis Berner, Baetisca gibbera Berner, Baetisca lacustris McDunnough, Baetisca laurentina McDunnough, Baetisca obesa (Say), Baetisca rogersi Berner, Baetisca rubescens (Provancher), Baetisca thomsenae Traver, Baetodes adustus Cohen & Allen, Baetodes alleni McCafferty & Provonsha, Baetodes arizonensis Koss, Baetodes bellus Mayo, Baetodes bibranchius McCafferty & Provonsha, Baetodes caritus Cohen & Allen, Baetodes deficiens Cohen & Allen, Baetodes edmundsi Koss, Baetodes fortinensis Mayo, Baetodes furvus Mayo, Baetodes fuscipes Cohen & Allen, Baetodes inermis Cohen & Allen, Baetodes longus Mayo, Baetodes obesus Mayo, Baetodes pallidus Cohen & Allen, Baetodes pictus Cohen & Allen, Baetodes sigillatus Allen & Chao, Baetodes tritus Cohen & Allen, Baetodes veracrusensis Mayo, Barbaetis benfieldi Kennedy, Barbaetis cestus (Provonsha & McCafferty), Blasturus austrinus Traver, Blasturus collinus Traver, Blasturus concinnus (Walker), Blasturus gracilis Traver, Blasturus gravastellus Eaton, Blasturus intermedius Traver, Blasturus nebulosus (Walker), Blasturus pacificus McDunnough, Blasturus vibex Traver, Brachycercus arcticus Soldan, Brachycercus berneri Soldan, Brachycercus edmundsi Soldan,

26 26 NORTH AMERICA EPHEMEROPTERA SPECIES Brachycercus fiavus Traver, Brachycercus fioridicola Soldan, Brachycercus idei (Lestage), Brachycercus lacustris (Needham), Brachycercus maculatus Berner, Brachycercus nasutus Soldan, Brachycercus nitidus (Traver), Brachycercus pini Soldan Brachycercus prudens (McDunnough), Brachycercus sp. A Berner, Brachycercus tuberculatus Soldan, Caenis amica Hagen, Caenis anceps Traver, Caenis bajaensis Allen & Murvosh, Caenis candida Harper & Harper, Caenis delicata Traver, Caenis diminuta Walker, Caenis diminuta diminuta Walker, Caenis forcipata McDunnough, Caenis gigas Burks, Caenis hilaris (Say), Caenis jocosa McDunnough, Caenis lacustris Needham, Caenis latipennis Banks, Caenis macafferti Provonsha, Caenis punctata McDunnough, Caenis ridens McDunnough, Caenis simulans McDunnough, Caenis tardata McDunnough, Caenis youngi Roemhild, Callibaetis americanus Banks, Callibaetis brevicostatus Daggy, Callibaetis californicus Banks, Callibaetis carolus Traver, Callibaetis centralis Peters, Callibaetis claudiae Edmunds, 1945 nomen nudum- 399 Callibaetis coloradensis Banks, Callibaetis doddsi Traver, Callibaetis evergreenensis Thew, Callibaetis ferrugineus ferrugineus (Walsh), Callibaetis ferntgineus (Walsh), Callibaetis ferrugineus hageni Eaton, Callibaetis floridanus Banks, Callibaetis fluctuans (Walsh), Callibaetis fuscus Dodds, Callibaetis hageni Eaton, Callibaetis hebes Upholt,

27 W. P. MCCAFFERTY 27 Callibaetis montanus Eaton, Callibaetis montanus montanus Eaton, Callibaetis montanus punctilusus McCafferty & Provonsha, Callibaetis nigritus Banks, Callibaetis pacificus Seemann, Callibaetis pallidus Banks, Callibaetis pictus (Eaton), Callibaetis pretiosus Banks, Callibaetis punctilusus McCafferty & Provonsha, Callibaetis semicostatus Banks, Callibaetis signatus Banks, Callibaetis skokianus Needham, Callibaetis traverae Upholt, Callibaetis undatus (Pict et), Callibaetis vitrea Dodds, Camelobaetidius arriaga (Traver & Edmunds), Camelobaetidius cepheus (Traver & Edmunds), Camelobaetidius chiapas (Traver & Edmunds), Camelobaetidius jenseni (Traver & Edmunds), Camelobaetidius mexicanus (Traver & Edmunds), Camelobaetidius musseri (Traver & Edmunds), Camelobaetidius navis (Allen & Chao), Camelobaetidius salinus (Allen & Chao), Camelobaetidius similis Lugo-Ortiz & McCafferty, Camelobaetidius sinaloa (Allen & Murvosh), Camelobaetidius trivialis (Allen & Chao), Camelobaetidius waltzi McCafferty, Camelobaetidius warreni (Traver & Edmunds), Camelobaetidius zenobia (Traver & Edmunds), Campsurus cuspidatus Eaton, Campsurus decoloratus (Hagen), Campsurus incertus Traver, Campsurus primus McDunnough, Campsurus puella (Pictet), Caudatella cascadia (Allen & Edmunds), Caudatella edmundsi (Allen), Caudatella heterocaudata californica (Allen & Edmunds), Caudatella heterocaudata circia (Allen & Edmunds), Caudatella heterocaudata heterocaudata (McDunnough), Caudatella hystrix (Traver), Caudatella jacobi (McDunnough), Caurinella idahoensis Allen, Centroptilum alamance (Traver), Centroptilum album McDunnough,