Releases and Recoveries of U.S. and NPRB Salmonid Data Storage Tags, and Recoveries of High Seas Tags in North America and Russia, 2005

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1 NPAFC Doc. 94 Rev. Releases and Recoveries of U.S. and NPRB Salmonid Data Storage Tags, and Recoveries of High Seas Tags in North America and Russia, 25 Robert V. Walker, Nancy D. Davis, and Katherine W. Myers SCHOOL OF AQUATIC AND FISHERY SCIENCES, University of Washington, Box 3552, Seattle, WA , U.S.A. John H. Helle NOAA, NMFS, Alaska Fisheries Science Center, AUKE BAY LABORATORY, 1135 Glacier Hwy., Juneau, Alaska , U.S.A. Masa-aki Fukuwaka HOKKAIDO NATIONAL FISHERIES RESEARCH INSTITUTE, Fisheries Research Agency 116 Katsurakoi, Kushiro 85-82, Shigehiko Urawa NATIONAL SALMON RESOURCE CENTER 2-2 Nakanoshima, Toyohira-ku, Sapporo , Vladimir I. Karpenko and Anatoly B. Dekshtein KamchatNIRO 18, Naberezhnaya St., Petropavlovsk-Kamchatsky 68362, Russia Sergei Zolotukhin Khabarovsk Branch, TINRO-Centre 13-A, Amursky al., Khabarovsk 6828, Russia Sergey Kovalenko SakhNIRO 196, Komsomol skaya St., Yuzhno-Sakhalinsk 69323, Russia submitted to the NORTH PACIFIC ANADROMOUS FISH COMMISSION by THE UNITED STATES OF AMERICA October 25 THIS PAPER MAY BE CITED IN THE FOLLOWING MANNER: Walker, R.V., N.D. Davis, K.W. Myers, J.H. Helle, M. Fukuwaka, S. Urawa, V.I. Karpenko, A.B. Dekshtein, and S. Zolotukhin. 25. Releases and recoveries of U.S. and NPRB salmonid data storage tags, and recoveries of high seas tags in North America and Russia, 25. (NPAFC Doc. 94.) 2 p. SAFS-UW-54. School of Aquatic and Fishery Sciences, University of Washington, Seattle, WA.

2 1 RELEASES AND RECOVERIES OF U.S. AND NPRB SALMONID DATA STORAGE TAGS, AND RECOVERIES OF HIGH SEAS TAGS IN NORTH AMERICA AND RUSSIA, 25 ABSTRACT Information is reported on all high-seas salmon tags recovered in North America from 1 October 24 through 3 September 25, and all releases and recoveries of U.S. and NPRB-funded data storage tags (DSTs). One hundred thirty-eight DSTs, which record temperature, depth, and geoposition based on light data, salinity, temperature, and depth, temperature and depth, or temperature-only data, were placed on Pacific salmon in the central North Pacific Ocean and Bering Sea during a research cruise aboard a ese vessel in June and July 25. During the cruise 63 sockeye, 24 chum, 33 pink, 13 coho, and 5 chinook salmon were tagged with DSTs. Twenty-two recoveries of fish tagged with DSTs are reported, two from North America, 18 from, and two from Russia. Notable distribution data includes a slight eastward range extension for maturing northern Okhotsk Sea coast chum salmon in the Bering Sea. Notable data tag recoveries include the first recovery for Russian salmon from the northern Okhotsk Sea coast (a chum salmon) and the first recovery for a Russian coho salmon (however, this tag failed). Data from northern Okhotsk and Hokkaido chum salmon showed typical maximum vertical movements to 6- m with deeper (25-4 m) vertical excursions in fish entering coastal waters with higher surface temperatures near Hokkaido. Graphs of ambient temperature and pressure data from the DSTs are presented. The most notable feature of DST recoveries from 24 tagging was the high incidence of tag failure. Overall, seven of the 2 tags received failed completely, and two had faulty depth data. In addition, two data tags were not recovered from fish where disk tags were found. We also include information on 13 previously unreported disk tag recoveries from previous years (12 from Russia and 1 Bering Sea recovery). These include three chum, eight pink, and two coho salmon, tagged in 1991 and 2-23 and recovered in those years. Recovery of a tag from an Okhotsk chum salmon provided a western range extension for immature fish of that region in the Bering Sea. The remaining Russian fish were recovered in eastern and western Kamchatka, Sakhalin Island, and unknown areas of the Russian Far East. INTRODUCTION Information is reported on all high-seas salmon (Oncorhynchus spp.) tags recovered in North America from 1 October 24 through 3 September 25, and all releases and recoveries of U.S. data storage tags (DSTs) and DSTs funded by the North Pacific Research Board (NPRB). The School of Aquatic and Fishery Sciences (SAFS), University of Washington, serves as a processing center for all North American recoveries of Canadian, ese, Russian, and U.S. high-seas salmon tags, and recoveries of U.S. and NPRB high-seas salmon tags and DSTs by all nations. Releases and recoveries of all U.S. and NPRB DSTs are reported, in order to have a complete record in one document. (Releases of U.S. DSTs from ese vessels and recoveries in Asia are also reported in Fukuwaka et al. 25.) Graphs of ambient temperature and pressure data from recovered DSTs are presented where data are available.

3 2 MATERIALS AND METHODS Fish were captured for tagging by research longline on a ese vessel in 25. U.S. highseas tags are 2 mm diameter plastic red-and-white Petersen disk tags. Four types of DSTs were used. One type is a small circuit board potted in a clear urethane, manufactured by Lotek Marine Technologies. Model _1-5 is 27 x 16 x 8 mm and weighs 5 g. These tags record temperature and depth data. Another Lotek tag, model _24, records temperature, depth, and estimated geoposition based on light data analyzed by the tag. ibutton-type tags are Thermochron ibutton data storage devices (DS-1921Z-F5) manufactured by Dallas Semiconductor, Inc., and repackaged in urethane for fish tagging by AlphaMach, Inc.; they record temperature data only. Model ibkrill tags are hexagonal, 25 x 13 x 8 mm and weigh 3.2 g. Model DST CTD tags are housed in a 27 x 16 x 8 mm ceramic shell and weigh 8 g. These tags, manufactured by Star-Oddi, record salinity, temperature, and depth data. _1 and ibutton tags were attached to fish just anterior to the dorsal fin using two 76 or 64 mm nickel pins, with labeled U.S. and ese disk tags placed on the pins on the other side of the fish. DST CTD and _24 tags were attached in the same location and method, but were affixed with stainless steel or surgical wire, with a small oval plastic plate on the opposite side of the fish. SAFS s high-seas tag processing center activities include: (1) advertising for tag recoveries, (2) returning tags and original recovery information to the appropriate release agencies, (3) mailing information on tag recoveries and a tag reward to fishermen and processors, (4) maintaining a file of original correspondence, data, and tags of all recoveries of U.S., U.S.-Russia, and -U.S. tags (1956-present), (5) maintaining and updating an all-agency tag release and computer database, and (6) reporting all recoveries of U.S., U.S.-Russia, and -U.S. high-seas tags to the North Pacific Anadromous Fish Commission (NPAFC). In addition, FRI scientists periodically prepare reports and maps based on historical recoveries of high-seas tags that describe the known ocean ranges of major regional stocks of Asian and North American salmonids (for example, Myers et al. 1996). The complete all-agency (Canada,, Russia, and United States) high-seas tag release and recovery computer database (1954-present) is available from FRI upon request from the parties of NPAFC so that all member nations can have access to a common database. RESULTS AND DISCUSSION One hundred thirty-eight DSTs, were placed on Pacific salmon in the central North Pacific Ocean and Bering Sea during a research cruise aboard a ese vessel in June and July 25 (Table 1). Ten DST CTDs (recording salinity, temperature, and depth data), 2 _24 (temperature, depth, and geoposition based on light), 1 _1-5 (temperature and depth), and seven ibkrill (temperature-only) tags were used. Sixty-three sockeye, 24 chum, 33 pink, 13 coho, and 5 chinook salmon were tagged with DSTs. From 1 October 24 through 3 September 25, two high-seas salmon tags from recoveries in North America have been reported (Table 2): one chinook and one sockeye salmon, both tagged with DSTs. The DST was not recovered with the sockeye salmon, and the DST on the chinook salmon failed. Both fish were tagged in the previous year (24) as immature fish, so the tag loss and failure are very unfortunate as lost opportunities to receive overwintering data. The chinook salmon was recovered at Kaltag in the Yukon River, and the sockeye salmon near Ilnik on the north coast of the Alaska Peninsula. Twenty salmon tagged with DSTs were recovered in waters of (18 chum) and Russia (1 chum and 1 coho) from tagging operations in 24 in the Bering Sea and North Pacific (Table 2). Notable data tag recoveries from Asia include the first recovery for salmon from the northern Okhotsk Sea coast (a chum salmon) and the first recovery for a Russian coho salmon (however, this tag failed). The northern Okhotsk chum salmon recovery is only the second from that region of a fish

4 3 tagged in the Bering Sea, and is a slight eastward range extension in the Bering Sea (55 3 N, W; previous record 55 3 N, E). Data from the northern Okhotsk chum salmon showed typical maximum vertical movements to 6-8 m (Fig. 1). The fish passed through an area of colder surface water (6 C), similar to temperatures seen in data from tags on Hokkaido and Sakhalin chum salmon as they migrate through the Kuril islands. Data from four chum salmon returning to Hokkaido showed typical maximum vertical movements to 6- m with deeper (25-4 m) vertical excursions as they encountered higher surface temperatures near Hokkaido (Figs. 2-3). Temperature data patterns from six other Hokkaido chum salmon carrying temperature-only tags are similar to data from Hokkaido chum salmon with temperature and depth tags (Figs. 4-6). Of the remaining eight DSTs on Hokkaido chum salmon, five failed completely, two had faulty depth data, and one was not recovered when the disk tags on the fish were found. The most notable feature of DST recoveries from 24 tagging was the high incidence of tag failure. Overall, seven of the 2 tags received failed completely, and two had faulty depth data. Tags of all three types (from three different manufacturers) failed. In addition, two data tags were not recovered from fish where disk tags were found. We also include information on 13 previously unreported disk tag recoveries from previous years (12 from Russia and 1 high seas recovery; Table 3). These include three chum, eight pink, and two coho salmon, tagged in 1991 and One pink salmon was recovered by the vessel that tagged it the morning after it was released in the Bering Sea. The remaining fish were recovered in eastern and western Kamchatka (8), the western coast of the Okhotsk Sea (1), Sakhalin Island (2), and unknown areas of the Russian Far East (1). The Sakhalin recoveries are the fifth and sixth recoveries of maturing Sakhalin chum tagged in the Bering Sea; four of these six recoveries have been from tagging since 21. The Okhotsk chum recovery is the seventh of an immature northern Okhotsk Sea chum salmon tagged in the Bering Sea (previous recoveries from tagging in and 1982), and is a western range extension in the Bering Sea (56 3 N, 178 W; previous record at N, 18 ). Release data for DSTs from 24 tagging operations and other recoveries in that year were reported by Walker et al. (24). Release data for disk tags from 1991 and 2-23 tagging operations were reported by Ogura (1991) and Fukuwaka et al. (2, 21, 22, 23). Recoveries of coded-wire tags by high seas research vessels and groundfish fisheries are reported in Myers et al. (25). ACKNOWLEDGMENTS ese, Russian, and U.S. fishermen, processors, biologists, and fisheries agencies who participated in the high-seas tagging program are gratefully acknowledged for returning tags and providing recovery information. Special acknowledgments are given to Captain Y. Murata and the officers, crew, and students aboard the Wakatake maru, and Captain J. Doi and the officers, crew, and students aboard the Kaiyo maru, who cooperated in high-seas salmonid tagging experiments in Funding for SAFS s high-seas tagging operations and tag processing center is from the Ocean Carrying Capacity program of the Auke Bay Laboratory of the Alaska Fisheries Science Center, NMFS (NOAA contract 5ABNF-1-2).

5 4 REFERENCES Fukuwaka, M., S. Urawa, I. Ono, H. Tanaka, N. Davis, and R.V. Walker. 2. Recoveries of highseas tags in in 1999, and tag releases and recoveries of fin-clipped salmon from ese research vessel surveys in the North Pacific Ocean in 2. (NPAFC Doc. 482.) Hokkaido National Fisheries Research Institute, Fisheries Research Agency, Kushiro. 12 p. Fukuwaka, M., S. Urawa, I. Ono, N. Davis, and R.V. Walker. 21. Recoveries of high-seas tags in in 2, and tag releases and recoveries of fin-clipped salmon from ese research vessel surveys in the North Pacific Ocean in 21. (NPAFC Doc. 545.) Hokkaido National Fisheries Research Institute, Fisheries Research Agency, Kushiro. 12 p. Fukuwaka, M., S. Urawa, K. Hirasawa, H. Tanaka, N. Davis, and R.V. Walker. 22. Recoveries of high-seas tags in in 21, and tag releases and recoveries of fin-clipped salmon from ese research vessel surveys in the North Pacific Ocean in summer of 22. (NPAFC Doc. 621.) Hokkaido National Fisheries Research Institute, Fisheries Research Agency, Kushiro. 11 p. Fukuwaka, M., S. Urawa, K. Hirasawa, N. Davis, and R.V. Walker. 23. Recoveries of high-seas tags in in 22, and tag releases and recoveries of fin-clipped salmon from ese research vessel surveys in the North Pacific Ocean in fall of 22 and summer of 23. (NPAFC Doc. 715.) Hokkaido National Fisheries Research Institute, Fisheries Research Agency, Kushiro. 11 p. Fukuwaka, M., S. Urawa, S. Yoshimitsu, N.D. Davis, and R.V. Walker. 25. Recoveries of highseas tags in in 24, and tag releases and recoveries of fin-clipped salmon from ese research vessel surveys in the North Pacific Ocean in summer of 25. (NPAFC Doc. 889.) Hokkaido National Fisheries Research Institute, Fisheries Research Agency of, Kushiro. 12 p. Myers, K.W., K.Y. Aydin, R.V. Walker, S. Fowler, and M.L. Dahlberg Known ocean ranges of stocks of Pacific salmon and steelhead as shown by tagging experiments, (NPAFC Doc. 192.) FRI-UW Fisheries Research Institute, University of Washington, Seattle, WA. 4 p. + figs. and appends. Myers, K.W., N.D. Davis, A.G. Celewycz, E.V. Farley, Jr., J.F.T. Morris, M. Trudel, M. Fukuwaka, S. Kovalenko, and A.O. Shubin. 25. High seas salmonid coded-wire tag recovery data, 25. (NPAFC Doc. 95.) SAFS-UW-55. School of Aquatic and Fishery Sciences, University of Washington, Seattle, WA. Nagasawa, T., M. Fukuwaka, S. Urawa, K. Hirasawa,N. Davis, and R.V. Walker. 24. Recoveries of high-seas tags in in 23, and tag releases and recoveries of fin-clipped salmon from ese research vessel surveys in the North Pacific Ocean in fall of 23 and summer of 24. (NPAFC Doc. 786.) Hokkaido National Fisheries Research Institute, Fisheries Research Agency, 116 Katsurakoi, Kushiro p. Ogura, M Release data for ese salmon tagging experiments in 1991 and recovery data up to August, (INPFC Doc ) National Research Institute of Far Seas Fisheries, Fisheries Agency of, Shimizu. 24 p.

6 5 Walker, R.V., N.D. Davis, K.W. Myers, and M. Fukuwaka. 22. Releases and recoveries of U.S. salmonid data storage tags, and recoveries of high seas tags in North America, 22. (NPAFC Doc. 68.) SAFS-UW-24. School of Aquatic and Fishery Sciences, University of Washington, Seattle, WA. 8 p. Walker, R.V., N.D. Davis, K.W. Myers, and M. Fukuwaka. 23. Releases and recoveries of U.S. salmonid data storage tags, and recoveries of high seas tags in North America, 23. (NPAFC Doc. 722.) SAFS-UW-38. School of Aquatic and Fishery Sciences, University of Washington, Seattle, WA. 14 p. Walker, R.V., N.D. Davis, K.W. Myers, S. Urawa, and K. Hirasawa. 24. Releases and recoveries of U.S. and NPRB salmonid data storage tags, and recoveries of high seas tags in North America, 24. (NPAFC Doc. 86.) SAFS-UW-46. School of Aquatic and Fishery Sciences, University of Washington, Seattle, WA. 23 p.

7 6 Table 1. U.S. and NPRB archival data storage tags placed on salmonids in the North Pacific Ocean and Bering Sea aboard the Wakatake maru in 25. Geo tags record geographical position, temperature and depth data (Lotek _24); tags record temperature and depth data (_-5); CTD tags record salinity, temperature, and depth (StarOddi DST CTD); and ibk tags record temperature only (AlphaMach ibkrill). JFRA = Fisheries Research Agency of. Data Tag Release Location Fork Other tags Tag # Model Species Date Latitude Longitude Length Age US JFRA 8114 coho 6/16/5 42 N LL85 MM coho 6/16/5 42 N X.1 LL86 MM coho 6/16/5 42 N LL87 MM coho 6/16/5 42 N LL88 MM coho 6/16/5 42 N LL89 MM coho 6/17/5 43 N LL8149 MM coho 6/17/5 43 N LL815 MM coho 6/17/5 43 N X.1 LL8154 MM coho 6/18/5 44 N LL8158 MM358 K112 ibk pink 6/21/5 47 N LL8166 MM coho 6/22/ N LL8175 MM chum 6/23/ N LL8177 MM chum 6/23/ N LL8178 MM chum 6/23/ N LL8179 MM chum 6/23/ N LL818 MM coho 6/23/ N LL824 MM sockeye 6/23/ N LL825 MM CTD chinook 6/24/ N X.2 LL8211 MM sockeye 6/24/ N LL8212 MM sockeye 6/24/ N LL8213 MM sockeye 6/24/ N LL8219 MM3119 K113 ibk coho 6/24/ N LL8227 MM chum 6/24/ N LL8228 MM3128 K114 ibk pink 6/25/5 5 3 N LL8229 MM chum 6/25/5 5 3 N LL823 MM CTD sockeye 6/25/5 5 3 N LL8238 MM CTD sockeye 6/25/5 5 3 N LL8239 MM3139 A13245 Geo sockeye 6/25/5 5 3 N LL824 MM314 K117 ibk coho 6/25/5 5 3 N LL8241 MM sockeye 6/25/5 5 3 N LL8242 MM sockeye 6/25/5 5 3 N LL8243 MM3143 A1325 Geo sockeye 6/25/5 5 3 N LL8246 MM sockeye 6/26/ N LL8249 MM chum 6/26/ N X.5 LL825 MM315 K116 ibk pink 6/27/ N LL8251 MM pink 6/28/ N LL8255 MM pink 6/28/ N LL8256 MM3156 continued

8 7 Table 1. continued. Vessel and Tag Release Location Fork Other tags Lengt Data Tag # Model Species Date Latitude Longitude h Age FRI JFRA A13243 Geo sockeye 6/28/ N LL8257 MM pink 6/28/ N LL8268 MM pink 6/28/ N LL8269 MM chum 6/28/ N LL827 MM CTD chum 6/28/ N LL8271 MM chum 6/28/ N LL8272 MM chum 6/28/ N LL8273 MM pink 6/29/ N LL8286 MM pink 6/29/ N LL8287 MM pink 6/29/ N LL837 MM327 A13257 Geo chum 6/29/ N LL838 MM328 A13244 Geo sockeye 6/3/ N LL839 MM chum 6/3/ N LL8314 MM3214 A13249 Geo sockeye 6/3/ N LL8315 MM pink 6/3/ N LL8316 MM pink 6/3/ N LL8317 MM sockeye 6/3/ N LL8318 MM pink 6/3/ N LL8319 MM pink 6/3/ N LL832 MM pink 6/3/ N LL8321 MM CTD sockeye 6/3/ N LL8322 MM CTD sockeye 6/3/ N LL8323 MM pink 6/3/ N LL8324 MM pink 7/1/ N LL8334 MM pink 7/1/ N LL8335 MM chum 7/1/ N LL8337 MM3237 A13251 Geo sockeye 7/1/ N LL8343 MM3243 A13252 Geo sockeye 7/1/ N LL8344 MM chum 7/1/ N LL835 MM325 A13253 Geo sockeye 7/1/ N LL8351 MM chum 7/1/ N LL8354 MM3254 A13259 Geo chum 7/1/ N LL8355 MM CTD chum 7/1/ N LL836 MM pink 7/2/ N LL8362 MM pink 7/2/ N LL8363 MM3263 A13254 Geo sockeye 7/2/ N LL8366 MM3266 A13255 Geo sockeye 7/2/ N X.X LL8367 MM sockeye 7/2/ N X.3 LL8369 MM pink 7/7/ N LL837 MM sockeye 7/7/ N LL8371 MM sockeye 7/7/ N LL8372 MM sockeye 7/7/ N LL8373 MM3273 continued

9 8 Table 1. continued. Vessel and Tag Release Location Fork Other tags Data Tag # Model Species Date Latitude Longitude Length Age US JFRA 947 sockeye 7/7/ N LL8374 MM pink 7/7/ N LL8375 MM pink 7/7/ N LL8376 MM pink 7/7/ N LL8377 MM pink 7/7/ N LL8378 MM3278 K3 ibk chinook 7/7/ N LL8379 MM sockeye 7/7/ N LL838 MM sockeye 7/7/ N LL841 MM sockeye 7/7/ N LL842 MM sockeye 7/7/ N LL843 MM333 K111 ibk chinook 7/8/ N 179 W LL844 MM sockeye 7/8/ N 179 W LL845 MM chinook 7/8/ N 179 W 535 X.2 LL846 MM sockeye 7/9/ N 178 W LL848 MM pink 7/9/ N 178 W LL849 MM pink 7/9/ N 178 W LL84 MM pink 7/9/ N 178 W LL8411 MM sockeye 7/9/ N 178 W LL8412 MM sockeye 7/9/ N 178 W LL8413 MM sockeye 7/9/ N 178 W LL8414 MM sockeye 7/9/ N 178 W LL8415 MM sockeye 7/9/ N 178 W LL8416 MM pink 7/9/ N 178 W LL8417 MM pink 7/9/ N 178 W LL8418 MM pink 7/9/ N 178 W LL8419 MM sockeye 7/9/ N 178 W LL842 MM sockeye 7/9/ N 178 W LL8421 MM sockeye 7/9/ N 178 W LL8422 MM sockeye 7/9/ N 178 W LL8423 MM sockeye 7/9/ N 178 W LL8424 MM chinook 7/9/ N 178 W LL8425 MM sockeye 7/9/ N 178 W 534 X.2 LL8443 MM sockeye 7/9/ N 178 W LL8444 MM sockeye 7// N 178 W LL8445 MM3345 sockeye 7// N 178 W LL8446 MM sockeye 7// N 178 W LL8447 MM sockeye 7// N 178 W LL8448 MM sockeye 7// N 178 W LL8449 MM sockeye 7// N 178 W LL845 MM sockeye 7// N 178 W LL8451 MM sockeye 7// N 178 W LL8452 MM pink 7/7/ N LL8377 MM3277 continued

10 9 Table 1. continued. Vessel and Tag Release Location Fork Other tags Data Tag # Model Species Date Latitude Longitude Length Age US JFRA 22 sockey 7// N 178 W LL8453 MM3353 e 17 pink 7// N 178 W LL8459 MM sockey 7// N 178 W LL846 MM336 e 19 sockey 7// N 178 W LL8461 MM3361 e A1326 Geo chum 7// N 178 W 538 X.X LL8462 MM3362 A13261 Geo chum 7// N 178 W LL8463 MM3363 A13263 Geo chum 7// N 178 W 55.3 LL8465 MM CTD chum 7/11/ N 179 W LL8466 MM pink 7/11/ N 179 W LL8468 MM sockey 7/11/ N 179 W LL8471 MM3371 e A13265 Geo chum 7/11/ N 179 W LL8472 MM CTD sockey 7/12/ N 179 E 554 X.3 LL8473 MM3373 e 26 sockey 7/12/ N 179 E LL8474 MM3374 e A13267 Geo sockey 7/13/ N 178 E LL8475 MM3375 e A13268 Geo sockey 7/13/ N 178 E LL8478 MM3378 e 221 CTD chum 7/13/ N 178 E LL8483 MM3383 A1327 Geo sockey 7/13/ N 178 E 596 X.X LL8484 MM3384 e A13271 Geo sockey e 7/13/ N 178 E LL8485 MM3385

11 Table 2. Preliminary release and recovery information for U.S. tags and cooperative -U.S. tags returned from 1 October 24 to 3 September 25. A blank indicates the information is not available. HL=hook and line, LL=longline, GN= gillnet, SGN=set gillnet. Age designation is the European method, first number is the number of freshwater annuli, second number is the number of ocean annuli. FL=fork length, and BW=body weight. Data storage tags: =Lotek _1-5 (records temperature and depth); ib=alphamach ibkrill (records temperature only); CTD=StarOddi DST CTD (records salinity, temperature, and depth). * = data tag not recovered U.S. Release Recovery Tag. Tag Lat. 2 X5 FL Lat. Area FL BW Gonad Nos. No. Date ( N) Long. Area Gear (mm) Age Date ( N) Long. Code Gear Sex (mm) (g) (g) Age Location A. Sockeye Salmon NN137, ibk W128* B. Chum Salmon LL7612, ibk K5 LL766, ibk K52 NN198, 5963* NN29, 598 NN237, 5991 NN245, 743 LL7811, 85 MM Jun W852 LL Jun W 5-2 GN M Ilnik, N. Alaska Peninsula, Alaska, USA BB Jun W W856 HL Oct E 2- SGN F Rausu, Nemuro BB Jun W W854 HL Oct E 2- SGN Rausu, Nemuro MM Jun W854 LL Sep E 2-2 SGN Ohmu, Okhotsk MM229 3-Jun W854 LL Oct E E 2-2 SGN F Hamatonbetsu, Okhotsk coast, Hokkaido, MM Jul W856 LL Sep E 2-1 SGN M Hiroo, Pacific MM Jul W856 LL Sep E M Urahoro, Pacific BB Jul W W854 HL Sep E 2-2 SGN M Abashiri, Okhotsk coast, Hokkaido,

12 11 Table 2. continued U.S. Release Recovery Tag. Tag Lat. 2 X5 FL Lat. Area FL BW Gonad Nos. No. Date ( N) Long. Area Gear (mm) Age Date ( N) Long. Code Gear Sex (mm) (g) (g) Age Location B. Chum Salmon - continued NN358, ibk W149 LL7836, 86 NN421, CTD 1379 NN541, 7467 NN562, 7469 NN581, 7473 NN67, 748 NN687, ibk W153 NN688, ibk W154 NN696, ibk W179 MM Jul W W856 LL Oct E 2- SGN M Nemuro, Nemuro N48 5-Jul W W854 HL Sep E F Kukhtui R., N. Okhotsk coast, Russia MM Jul W E7556 LL Sep E F Horoto River, Pacific coast, Hokkaido, MM Jul W W856 LL Oct E E MM Jul E E7556 LL Sep E E 2-1 SGN M Shibetsu, Nemuro coast, Hokkaido, M Taiki, Pacific MM Jul E E7556 LL Oct E 2-1 SGN F Erimo, Pacific MM267 -Jul E E7556 LL Sep E 2-1 SGN Kushiro, Pacific MM Jul E E7556 LL Sep E F Kushiro River, Pacific coast, Hokkaido, MM Jul E E7556 LL Sep E E 2- SGN M Rausu, Nemuro MM Jul E E7556 LL Sep E 2- SGN F Bekkai, Nemuro

13 12 Table 2. continued U.S. Release Recovery Tag. Tag Lat. 2 X5 FL Lat. Area FL BW Gonad Nos. No. Date ( N) Long. Area Gear (mm) Age Date ( N) Long. Code Gear Sex (mm) (g) (g) Age Location B. Chum Salmon - continued NN72, ibk W185 NN75, ibk W188 C. Coho Salmon LL769, 81 MM Jul E E7556 LL Oct E E 2- SGN M Shibetsu, Nemuro coast, Hokkaido, MM Jul E E7556 LL Sep E 2-1 SGN F Shiranuka, Pacific coast, Hokkaido, BB Jul W W85 HL Aug E 11-1 GN M Kamchatka R., E Kamchatka, Russia D. Chinook Salmon NN686, 7484 MM Jul E E7556 LL 7 X.3 3-Jun W 42-1 GN F Kaltag, Yukon R., western Alaska, USA

14 13 Table 3. Preliminary release and recovery information for previously unreported ese and cooperative -U.S. tags recovered in Russia or the Bering Sea and returned from 1991 to 3 September 25. A blank indicates the information is not available. LL=longline, GN= gillnet, SGN=set gillnet, TN=trapnet. Age designation is the European method, first number is the number of freshwater annuli, second number is the number of ocean annuli. FL=fork length and BW=body weight. U.S. Release Recovery Tag. Tag Lat. 2 X5 FL Lat. Area FL BW Gonad Nos. No. Date ( N) Long. Area Gear (mm) Age Date ( N) Long. Code Gear Sex (mm) (g) (g) Age Location A. Chum Salmon LL5853 Y Jul W W856 LL Oct E F Monetka Hatchery, Ostrovka R., Aniva Bay, SE Sakhalin, Russia - Y Jul E E7556 LL Aug E M Sokolnikovskiy Hatchery, Zavetinka R., SW Sakhalin, Russia LL453 LL353 7-Jul W W856 LL Jul E 7-1 SGN M Ayan City, W. Okhotsk coast, Russia B. Pink Salmon LL4353 LL Jul W854 LL E. Kamchatka, Russia LL4537 LL Jul W W856 LL Aug E Podsobnaya R., Korf Bay, E. Kamchatka, Russia LL685 Z Jun W848 LL June-July E 22 GN western Bering Sea, off E. Kamchatka LL6191 Z Jun W852 LL Jul E M Tymlat R., Karaginski Bay, E. Kamchatka, Russia

15 14 Table 3. continued U.S. Release Recovery Tag. Tag Lat. 2 X5 FL Lat. Area FL BW Gonad Nos. No. Date ( N) Long. Area Gear (mm) Age Date ( N) Long. Code Gear Sex (mm) (g) (g) Age Location B. Pink Salmon - continued LL6382 Z Jun W854 LL Aug E F Karaginski Bay near Ivashka R., E. Kamchatka, Russia LL657 Z747 3-Jul W W856 LL E 12-1 TN M Dranka R., Karaginski Bay, E. Kamchatka, Russia LL6716 Z Jul E E7556 LL E 12-1 TN F Dranka R., Karaginski Bay, E. Kamchatka, Russia LL6773 Z7673 -Jul E E7556 LL Jul E 24 GN Bering Sea, recovered by tagging vessel C. Coho Salmon - HH Jun E E7544 LL Far East, Russia LL3345 EE Jun W846 LL Oct Kotelnikova R., W. coast Kamchatka, Russia

16 15 TD 86 - N. Okhotsk Chum Recovered: 5 September 24 Kukhtui R., N. Okhotsk coast female, 56 mm, 1.95 kg Depth (m) Tagged: 5 July 24 Location: 55-3 N, W Length: 54 mm, Age: July August September Figure 1. Temperature and depth data recorded on a data storage tag placed on a 54 mm chum salmon in the Bering Sea on 5 July 24 and recovered in the Kukhtui River, north Okhotsk Sea coast, on 5 September 24.

17 16 18 TD Tag Hokkaido Chum Recovered: 22 October 24 Shibetsu, Nemuro coast, Hokkaido Male Length: 65 mm Weight: 2.8 kg Depth (m) Tagged: 8 July 24 Location: 56-3 N, 179- W Length: 583 mm Age: July August September October TD Hokkaido Chum Recovered: 27 September 24 Taiki, Pacific coast, Hokkaido Male Depth (m) Tagged: 9 July 24 Location: 56-3 N, 179- E Length: 641 mm Age: July August September 3 Figure 3. Temperature and depth data recorded on data storage tags placed on two chum salmon in the Bering Sea on 8 July 24 (top) and 9 July 24 (bottom) and recovered in Hokkaido,, on the Nemuro coast on 22 October 24 (top) and the Pacific coast on 27 September 24 (bottom).

18 17 25 TD Tag Hokkaido Chum Recovered: 11 October 24 Hamatonbetsu, Okhotsk coast, Hokkaido Female Length: 575 mm Weight: 2.2 kg Depth (m) 5 35 Tagged: 3 June 24 Location: 54-3 N, 18- Length: 521 mm Age: July August September October TD 85 - Hokkaido Chum Recovered: 28 September 24 Abashiri, Okhotsk coast, Hokkaido Male Length: 619 mm Weight: 2.45 kg Tagged: 5 July 24 Location: 55-3 N, W Length: 58 mm Age: Depth (m) July August September 45 Figure 4. Temperature and depth data recorded on data storage tags placed on two chum salmon in the Bering Sea on 3 June 24 (top) and 5 July 24 (bottom) and recovered in Hokkaido,, on the Okhotsk coast on 11 October 24 (top) and 28 September 24 (bottom).

19 18 ib Tag W153 - Hokkaido Chum Recovered: 21 September 24 Kushiro River, Pacific coast, Hokkaido Female Length: 745 mm Weight: 4.43 kg Tagged: 11 July 24 Location: 56-3 N, 177- E Length: 719 mm Age: July August September ib Tag W154 - Hokkaido Chum Recovered: 29 September 24 Rausu, Nemuro coast, Hokkaido Male Length: 698 mm Weight: 3.59 kg 2 15 Tagged: 11 July 24 Location: 56-3 N, 177- E Length: 564 mm Age: July August September Figure 5. Temperature data recorded on data storage tags placed on two chum salmon in the Bering Sea on 11 July 24 and recovered in Hokkaido,, on the Pacific coast on 21 September 24 (top) and on the Nemuro coast on 29 September 24 (bottom).

20 19 18 ib Tag W185 - Hokkaido Chum Recovered: 22 October 24 Shibetsu, Nemuro coast, Hokkaido Male Length: 65 mm Weight: 2.2 kg Tagged: 13 July 24 Location: 57-3 N, 176- E Length: 588 mm Age: July August September October 18 ib Tag W188 - Hokkaido Chum Recovered: 2 September 24 Shiranuka, Pacific coast, Hokkaido Female Length: 78 mm Weight: 5.2 kg Tagged: 13 July 24 Location: 57-3 N, 176- E Length: 741 mm Age: July August September Figure 5. Temperature data recorded on data storage tags placed on two chum salmon in the Bering Sea on 13 July 24 and recovered in Hokkaido,, on the Nemuro coast on 22 October 24 (top) and on the Pacific coast on 2 September 24 (bottom).

21 2 2 ib Tag W149 - Hokkaido Chum Recovered: 2 October 24 Nemuro, Hokkaido Male Length: 7 mm Weight: 3.5 kg Tagged: 5 July 24 Location: 57-3 N, 179- W Length: 637 mm Age: July August September 2 ib Tag W179 - Hokkaido Chum Recovered: 21 September 24 Bekkai, Nemuro coast, Hokkaido Female Length: 615 mm Weight: 2.65 kg Tagged: 12 July 24 Location: 57-3 N, 177- E Length: 68 mm Age: July August September Figure 5. Temperature data recorded on data storage tags placed on two chum salmon in the Bering Sea on 6 July 24 (top) and 12 July 24 (bottom) and recovered in Hokkaido,, on the Nemuro coast on 2 October 24 (top) and on 21 September 24 (bottom).