A key for identification with description of new species of the genus Pyrrhalta Joannis, 1865 (Coleoptera: Chrysomelidae: Galerucinae) from Indochina

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1 Кавказский энтомол. бюллетень 9(2): CAUCASIAN ENTOMOLOGICAL BULL A key for identification with description of new species of the genus Pyrrhalta Joannis, 1865 (Coleoptera: Chrysomelidae: Galerucinae) from Indochina Определительная таблица с описанием новых видов рода Pyrrhalta Joannis, 1865 (Coleoptera: Chrysomelidae: Galerucinae) из Индокитая Л.Н. Медведев A.N. Severtsov Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospect, 33, Moscow Russia. lev.n.medvedev@mail.ru Институт проблем экологии и эволюции им. А.Н. Северцова РАН, Ленинский проспект, 33, Москва Россия Key words: Coleoptera, Chrysomelidae, Galerucinae, Pyrrhalta, Indochina, key, new taxa, new synonymy. Ключевые слова: Coleoptera, Chrysomelidae, Galerucinae, Pyrrhalta, Индокитай, определитель, новые таксоны, новые синонимы. Abstract. A key of 26 species of the genus Pyrrhalta from Indochina is given. Five species are described: P. tuberculata sp. n., P. laosica sp. n. (Laos), P. gorbunovi sp. n. (Laos, Thailand), P. fedorenkoi sp. n., P. ornata sp. n. (South Vietnam). New synonymy is established: P. multicostata (Pic, 1928) = Atysa octocostata L. Medvedev, 1999, syn. n. Poneridia costata (Allard, 1889) is possibly identical with Pyrrhalta unicostata (Pic, 1937). Резюме. Приводится определитель 26видовPyrrhalta из Индокитая. Описаны 5 новых видов: P. tuberculata sp. n., P. laosica sp. n. (Лаос), P. gorbunovi sp. n. (Лаос, Таиланд), P. fedorenkoi sp. n., P. ornata sp. n. (Южный Вьетнам). Atysa octocostata L. Medvedev, 1999, syn. n. является новым синонимом P. multicostata (Pic, 1928). Poneridia costata (Allard, 1889), возможно, идентичен Pyrrhalta unicostata (Pic, 1937). Introduction The genus Pyrrhalta Joannis, 1865 from Indochina is rather poorly studied till now. A faunistic list of the tribe Galerucini from Vietnam includes 9 species of Pyrrhalta [Samoderzhenkov, 1988]. In the monograph of Galerucinae from Indochina [Kimoto, 1989] only 7 species of Pyrrhalta were recorded, because new species described by Samoderzhenkov were still unknown to S. Kimoto (both mentioned publications appeared almost in the same time). Three more species of Pyrrhalta were described from Vietnam in the last years [Medvedev, 2012a, b; Skomorokhov, 2011]. In the key given below I include 25 species, among them 5 species (3 from Laos and 2 from South Vietnam) are new for science. But it seems that additional new species or records from this area are possible. For example, Chinese fauna of Pyrrhalta includes about 50 species [Catalogue..., 2010]. The genus Pyrrhalta was split in a few genera, because almost all its Palearctic subgenera were raised to generic status; however this splitting needs further investigation and confirmation. In this work I accept Pyrrhalta in the understanding of Ogloblin [936], Gressitt and Kimoto [1963] and Kimoto [1989]. It s very possibly, that in future Oriental Pyrrhalta will be divided in a few subgenera or even genera, because differences among many species are very sharp. Next abbreviations are used for the depository places: NHMB Naturhistorisches Museum Basel, Switzerland; JB collection of Jan Bezdĕk, Brno, Czech Republic; LM author s collection, Moscow, Russia. A key to species of the genus Pyrrhalta 1(8). Elytra with longitudinal or oblique ridges or tuberculate. 2(3). Elytra with tubercles, partly united in oblique ridges. Antennae with segments 6 11 strongly widened, segment 10 transverse. Upperside rusty fulvous with tubercles a little darker, antennae with 6 basal segments fulvous with black apices, segments 7 11 black. Underside black, legs fulvous. Length mm. Laos... P. tuberculata sp. n. 3(2). Elytra with longitudinal ridges. Antennae slender or slightly widened. 4(7). Elytra with 3 4 longitudinal ridges. Antennal segment 10 not more than 1.5 times as long as wide. 5(6). Each elytron with 4 ridges. Upperside reddish with spot on vertex, longitudinal stripe on prothorax, scutellum and sutural margin of elytra black; antennal underside and legs black. Length mm. Vietnam, Laos, Thailand, Myanmar... P. multicostata (Pic, 1928) = Atysa octocostata L. Medvedev, 1999, syn. n. 6(5). Each elytron with 3 ridges, but outermost one feeble, sometimes almost indistinct. Upperside fulvous to dark fulvous with black spots on head, prothorax and elytra (Fig. 1), antennae black, but segments 1 4 with fulvous bases, scutellum with black base. Aedeagus Fig. 10. Length mm. Laos, Thailand... P. gorbunovi sp. n. 7(4). Elytron with one ridge beginning from humerus and going parallel to side margin. Body entirely fulvous. Antennal segment 10 about 3 times as long as wide. Aedeagus Fig. 11. Length mm. Vietnam, Laos, Thailand, China (Hainan)... P. unicostata (Pic, 1937)

2 268 8(1). Elytra without ridges or tubercles (sometimes with feeble elevation along anterior margin). 9(14). Elytra largely greenish. 10(13). Antennae robust, in preapical segments nearly as long as wide. 11(12). Elytra greenish with lateral margin including epipleurae brownish. Body fulvous. Length 9 mm. North Vietnam P. laboissieri Kimoto, (11). Generally greenish, elytra with lateral margins broadly blue, abdomen brownish. Anterior and posterior margins of prothorax not bordered. Length mm. North Vietnam. (Previously described as Decoomanius limbatus Laboissière, 1927)... P. limbata (Laboissière, 1927) 13(10). Antennae slender, in preapical segments much wider than long. Fulvous, antennae blackish, elytra greenish with marginal area fulvous. Length 4.5 mm. North Vietnam. (Previously described as Chapalia jeanvoinei Laboissiere, 1929)... P. jeanvoinei (Laboissière, 1929) 14(9). Elytra without metallic green color. 15(16). Elytra black. Head, prothorax and scutellum red. Antennae black with fulvous basal segments, underside and femora fulvous, tibiae and tarsi black. Antennal segment 10 about 3 times as long as wide. Prothorax with deep transverse impression, more shallow in middle. Aedeagus Fig. 12. Length 4.1 mm. South Vietnam P. impressicollis Samoderzhenkov, (15). Upperside fulvous or reddish, sometimes with black pattern. 17(18). Elytra reddish brown with interior 1/3 black. Body black, clypeus brownish. Length 5 mm. North Vietnam P. rubromarginata (Laboissière, 1929) 18(17). Elytra otherwise marked or entirely fulvous, prothorax not entirely black. 19(26). Prothorax fulvous with black pattern. 20(25). Prothorax with 3 4 black spots. 21(24). Prothorax with 3 black spots. 22(23). Elytra without spots. A spot on vertex, apices of antennal segments, scutellum, apices of tibiae and tarsi black. Antennal segment 10 about twice as long as wide. Aedeagus Fig. 13. Length mm. South Vietnam... P. fedorenkoi sp. n. 23(22). Elytra with 5 black spots (Fig. 2), middle on vertex, scutellum and metasternum black. Antennal segment 10 a little longer than wide. Length mm. South Vietnam, Thailand, South-Eastern China, Taiwan P. maculata Gressitt et Kimoto, (21). Prothorax with 4 black spots: 2 before base and 2 elongate near anterior angles. 2 elongate stripes on vertex and scutellum black. Length 7.25 mm. South Vietnam: island Con Dao... P. prokofievi Skomorokhov, (20). Prothorax with large transverse black spot in middle. Fulvous, labrum except sides, large spot on vertex, 2 elongate spots on base of elytra and apical half of elytra except margins (fig. 3) black. Antennal segment 10 about 3 times as long as wide. Aedeagus Fig. 14. Length 8.2 mm. South Vietnam P. ornata sp. n. 26(19). Prothorax entirely fulvous or reddish. 27(36). Elytra fulvous or pale flavous with black pattern. 28(29). Elytra pale flavous (much lighter than red fulvous head and prothorax) with all margins narrowly black (Fig. 4). Antennae black, underside fulvous with black metasternum, legs fulvous with black upperside of tibiae. Antennal segment 10 about twice as long as wide. Aedeagus Fig. 15. Length mm. South Vietnam P. curticollis Samoderzhenkov, (28). Elytra not paler than prothorax. 30(33). Elytra with black basal band. Antennal segment 10 about 3 times as long a wide. Underside fulvous. 31(32). Elytral basal band doesn t reach side margin (Fig. 5), suture fulvous, apical slope without black spot. Legs fulvous with more or less blackish upperside of tibiae. Aedeagus Fig. 16). Length mm. South Vietnam. Feeding was registered mostly on Trema (Ulmaceae), but also on Callicarpa (Verbenaceae) P. basifasciata Samoderzhenkov, (31). Elytral basal band reaches side margin, suture narrowly black, apical slope with black spot (Fig. 6), legs fulvous with black tibiae and darkened tarsi. Length 7.4 mm. North Vietnam P. basifasciata Samoderzhenkov, 1988 (? color form) 33(30). Elytra otherwise marked. 34(35). Elytra with longitudinal stripe from humeral tubercle along side margin and transverse spot at base, sometimes connected with stripe black (Figs 7, 8), very rarely stripe is reduced. Fulvous, metasternum and abdomen sometimes more or less darkened. Antennal segment 10 about 4 times as long as wide. Aedeagus fig. 17. Length mm. South Vietnam. Elytral pattern sometimes reduced (see item 46) P. ornatipennis Samoderzhenkov, (34). Elytral suture narrowly black (Fig. 9). Fulvous; vertex, antennae except bases, scutellum, meso- and metasternum black, upperside of tibiae and tarsi more or less darkened. Antennal segment 10 about twice as long as wide. Aedeagus Fig. 18. Length mm. North Vietnam P. kabakovi Samoderzhenkov, (27). Upperside, including head, entirely fulvous. 37(40). Legs bicolor. Underside fulvous. 38(39). Legs fulvous with black tibiae and tarsi. Antennae black with 2 basal segments fulvous (Fig. 26). Segment 10 about twice as long a wide. Aedeagus fig. 19. Length mm. South Vietnam... P. nigrotibialis L. Medvedev, (38). Legs fulvous with tarsi and mostly apices of tibiae black. Antennae with segments 1 3 and 8 11 fulvous, 4 7 black with fulvous bases (Fig. 27). Antennal segment 10 about 2.5 times as long as wide. Aedeagus Fig. 20. Length mm. North Vietnam P. apicicornis L. Medvedev, (37). Legs entirely fulvous. 41(44). Body small, not more than 5 mm. Antennal segment 10 about 1.5 times as long as wide. 42(43). Prothorax narrow, maximal width of elytra times wider than prothorax. Basal margin of prothorax distinctly emarginated before scutellum. Antennal segment 10 about 1/5 times as long as wide. Antennae black with 4 basal segments fulvous, underside and legs dark fulvous. Length mm. South Vietnam. Male unknown P. microphthalma Samoderzhenkov, (42). Prothorax more broad, maximal width of elytra only 1.6 times as wide as prothorax. Basal margin of prothorax not emarginated before scutellum. Antennae black with 3 basal segments fulvous, underside and legs fulvous. Length 3.8 mm. North Vietnam (Tam Dao, 1 female)... P. sp. 44(41). Body large, not less than 5.8 mm. 45(48). Antennae entirely fulvous. 46(47). Prothorax with longitudinal impression in middle, widened before scutellum and deep round groove on each side. Antennal segment 10 about 4 times as long as wide. Fulvous, metasternum and abdomen sometimes more or less darkened. Aedeagus Fig. 17. Length mm. South Vietnam. See also item P. ornatipennis Samoderzhenkov, (46). Prothorax without longitudinal impression in middle, with shallow round impression on each side. Antennal segment 10 about 3 times as long as wide. All body entirely fulvous. Aedeagus Fig. 21. Length mm. North Vietnam (Tam Dao), China... P. corpulenta Gressitt et Kimoto, (45). Antennae partly black. 49(52). Elytra with longitudinal concavity along side margin from

3 A key for identification with description of new species of the genus Pyrrhalta Joannis, 1865 from Indochina 269 or with black apices (Figs 29, 30). Aedeagus asymmetrical (Fig. 23) and has other form. Side margin of elytra more narrow and curved upwards. Length of body mm. South Vietnam... P. nuibata L. Medvedev, (49). Elytra with longitudinal concavity only in anterior half of side margin and without fold incite of concavity. Underside of aedeagus with sharp ridge in basal half. Aedeagus feebly asymmetrical (Fig. 24). Antennae mostly with segments 1 2 fulvous, 3 rd bicolor, 4 8 black, 9 11 fulvous (Figs 31, 32). Length mm... P. laosica sp. n. Pyrrhalta tuberculata sp. n. Material. Holotype Laos, Khammouang Prov., Ban Khounkhan (Nahin), 18 1 / , 200 m, , leg. O. Gorbunov (LM). Paratype: same locality and date, 1 (LM). Description. Rusty fulvous, 6 basal antennal segments fulvous with black apices, 5 apical segments black, underside black, legs fulvous, pubescence light fulvous. Body elongate, widened to behind. Head and all upperside covered with adpressed pubescence. Head rugosely punctate, clypeus with central ridge, angularly emarginated on anterior margin, frontal tubercles obliquely placed and poorly delimited. Antennae reach anterior third of elytra, with 6 apical segments widened, proportions of segments are as 15 : 7 : 10 : 10 : 10 : 10 : 9 : 8 : 7 : 7 : 10, preapical segments feebly transverse. Prothorax twice as wide as long, broadest and produced in middle, hind margin excavated before scutellum, anterior angles obtuse, hind angles broadly rounded, surface uneven, grooved on each side and more feeble in middle before scutellum, densely punctured, but punctures not very distinct under pubescence. Scutellum trapeziform with rounded apex, its sculpture not seen under dense pubescence. Elytra 1.4 times as long as wide, moderately widened to behind, surface dull, uneven, each elytron with 4 tubercles: one in anterior quarter near suture, obliquely prolonged to humerus, other near middle, the third just before apical slope and the last before apex, punctures fine and moderately dense, but not very distinct. Apical abdominal sternite with feebly rounded hind margin. Length of body mm. Differential diagnosis. The new species is near P. digambara group from Northern India (P. digambara Maulik, 1936, P. aludela Maulik, 1936, P. aurata Maulik, 1936), but differs from all known species in having 4 large tubercles on each elytron. Pyrrhalta gorbunovi sp. n. (Figs 1, 10) Figs 1 9. Elytral patterns of species of the genus Pyrrhalta Joannis, Рис Рисунок надкрылий видов рода Pyrrhalta Joannis, P. gorbunovi sp. n.; 2 P. maculata Gressitt et Kimoto, 1963; 3 P. ornata sp. n.; 4 P. curticollis Samoderzhenkov, 1988; 5, 6 P. basifasciata Samoderzhenkov, 1988; 7, 8 P. ornatipennis Samoderzhenkov, 1988; 9 P. kabakovi Samoderzhenkov, base to apical slope, divided from dorsal surface with feeble fold going from humerus. Underside of aedeagus without ridge in basal half. 50(51). Antennal segment 4 9 black, next segments mostly bicolor or apical segment entirely fulvous (Fig. 28). Aedeagus practically symmetrical (Fig. 22). Length mm. South Vietnam... P. hanunga Samoderzhenkov, (50). Practically all intermediate antennal segments bicolor, black with fulvous bases, 3 or 4 apical segments entirely fulvous Material. Holotype ( ): Laos, Khammouang Prov., Ban Khounkhan (Nahin), N / E, 200 m, , leg. O. Gorbunov (LM). Paratypes: same locality and date, 3 (LM); Thailand, Mae Hong Son Prov., Kiwlom pass near Soppong, , 1400 m, 2 leg. R. and H. Fouque (JB). Description. Upperside fulvous to dark fulvous, head with spot on vertex and another between eyes black, antennal segments 1 6 mostly with darkened apices, next segments black, prothorax with 3 black spots in basal half, but lateral ones small and sometimes not very distinct, scutellum with black basal half, elytra with numerous elongate black marks between ridges and mostly on impressed areas (Fig. 1), underside black with fulvous apex of abdomen and pygidium, legs with spot on middle of femora apices of tibiae and tarsi blackish. Pubescence of upperside of upperside golden fulvous or partly white. Body elongate, slightly widened to behind. Head and all upperside covered with dense adpressed pubescent, radiated on elytra outwards from the middle of depressed areas. Head finely punctate, clypeus with central ridge and almost straight anterior margin, frontal tubercles oblique, very poorly delimited, microsculptured. Antennae reach anterior quarter of elytra, proportions of segments are as 11 : 7 : 7 : 7 : 7 : 7 : 7 : 7 : 7 : 7 : 10,

4 270 five apical segments slightly thickened, about 1.5 times as long as wide. Prothorax 1.75 times as wide as long, broadest and angulate in anterior third, anterior angles obtuse, posterior angles angulate and removed on posterior third of lateral margin, hind margin not excavated before scutellum, surface dull, uneven, with deep groove on each side, punctures fine, but practically invisible under pubescence. Scutellum trapeziform with truncate apex, strongly convex, hill-like. Elytra 1.7 times as long as wide, dull, uneven, with 2 ridges in middle and one along side margin, punctures dense but not seen under pubescence. Apical abdominal sternite of female broadly rounded on hind margin. Aedeagus (Fig. 10) parallel-sided with short triangular apex, not curved in lateral view. Length of male 3.6 mm, of female mm. Differential diagnosis. The new species is near P. digambara (Maulik, 1936) and P. medvedevi Sprecher et Zoia, 2002 from India and Nepal, differs from both in sculpture and color of upperside and form of aedeagus, which has rounded apex in P. digambara and cuneiform in P. medvedevi (see Sprecher-Uebersax and Zoia [2002]). Pyrrhalta fedorenkoi sp. n. (Figs 13, 25) Material. Holotype ( ): Vietnam, Binh Phuoc Prov., Bu Gia Map National Park, / , m, at light, , leg. D. Fedorenko (LM). Paratypes: same locality and date, 3 ex. (LM). Description. Fulvous, spot on vertex, 3 spots on prosternum, apices of many antennal segments (Fig. 25), scutellum, apices of tibiae and tarsi black. Pubescence white. Body elongate, slightly widened to behind. Head and upperside with more or less erect hairs. Clypeus impunctate, frons and vertex densely punctate, frontal tubercles obliquely placed, convex, finely punctate and microsculptured, interantennal space convex, vertex without impressions. Antennae reach anterior third of elytra, proportions of segments are as 10 : 5 : 6 : 8 : 8 : 7 : 7 : 9 : 7 : 7 : 6 : 7, preapical segments about twice as long as wide. Prothorax 2.1 times as wide as long, broadest behind middle, side margins rounded, all angles obtuse, basal margin without emargination in middle, surface strongly punctate, especially on sides, with very feeble, scarcely visible lateral depression and practically indistinct central depression. Scutellum with broadly rounded apex, distinctly punctate. Elytra times as long as wide, very densely punctate, with interspaces much smaller than diameter of punctures. Epipleurae distinct to apical quarter of elytra. Segment 1 of anterior tarsus feebly widened in male. Aedeagus (Fig. 13) almost symmetrical, with acute triangular apex. Length of body mm. Differential diagnosis. The new species is near P. nuibata L. Medvedev, 2012 from the same locality, but differs in having spotted prothorax and vertex, black scutellum, bicolor legs and other form of aedeagus. Pyrrhalta ornata sp. n. (Figs 3, 14) Material. Holotype ( ): Vietnam, Tuyen Quang Prov., 3 km SE of Pac Ban village, Na Hang Nature Reserve, N / E, at light, , leg. G. Csorba (LM). Description. Fulvous, labrum except sides, large spot in middle of vertex and in middle of prothorax, 2 spots on base of elytra and apical half of elytra except margins (Fig. 3) black. Body elongate, slightly widened to behind. Head and all upperside covered with white adpressed pubescence. Clypeus triangular, impunctate, convex between antennal insertions, frontal tubercles subquadrangular, convex, shining and impunctate, poorly delimited posteriorly, vertex strongly and densely punctate. Antennae reach middle of elytra, proportions of segments are as 13 : 7 : 10 : 12 : 12 : 11 : 11 : 10 : 10 : 9 : 7, preapical segments about 3 4 times as long as wide. Prothorax twice as wide as long, broadest and angulate in middle, side margin straight before and concave after angulation, anterior angles obtuse, posterior angles rounded obtuse, surface uneven, with large round impression on each side, strongly and densely punctate with very narrow interspaces. Scutellum almost quadrate, densely punctate. Elytra 1.55 times as long as wide, dull, even, punctures moderately fine and very dense. Last abdominal sternite incised on hind margin. Aedeagus Fig. 14. Length of body 8.2 mm. Differential diagnosis. Pyrrhalta ornata sp. n. differs well from all Oriental species with very specific elytral pattern. Pyrrhalta laosica sp. n. (Figs 24, 31, 32) Material. Holotype ( ):N Laos, 17 km NNE Oudum Xai, N / E, about 1100 m, , leg. Vit Kuban (MNHB). Paratypes: same locality and date, 7 ex. (4 ex. MNHB, 3 ex. LM). Description. Entirely fulvous, except color of antennae which is variable: dark antennae have basal segments fulvous, 3 rd bicolor, 4 8 black, 9 11 bicolor; light antennae have segments 1 3 fulvous, 4 8 bicolor, 9 11 fulvous; there are also intermediate types of color (Figs 31, 32). Pubescence white. Body elongate. Head and upperside with adpressed hairs. Clypeus impunctate, frons and vertex densely punctate, frontal tubercles feebly convex, not sharply delimited, punctate, without microsculpture, interantennal space convex, vertex thin impressed line. Antennae reach middle of elytra, proportions of segments are as 9 : 5 : 7 : 10 : 9 : 9 : 9 : 8 : 8 : 8 : 9, preapical segments about 3 times as long as wide. Prothorax twice as wide as long, broadest just before middle, side margins rounded, anterior angles acute, posterior angles obtuse, surface densely and rather strongly punctate, with feeble longitudinal impression widened at base and shallow impression on each side. Scutellum with broadly rounded, almost truncate apex, strongly punctate. Elytra times as long as wide, very densely punctate, with interspaces smaller than diameter of punctures, with longitudinal concavity in anterior half of side margin and without fold inside of concavity. Epipleurae reach almost apex of elytra. Segment 1 of anterior tarsus of male very feebly widened. Aedeagus (Fig. 24) asymmetrical, with sharp ridge in basal part of underside. Length of body mm. Differential diagnosis. The new species is near P. hanunga Samoderzhenkov, 1988 and P. nuibata L. Medvedev, 2012, differs from both in other sculpture of elytra, form of aedeagus and a few differences in color of antennae. Pyrrhalta limbata (Laboissière, 1927) Remark. Kimoto [1989] included the genus Decoomanius Laboissière, 1927 in the genus Pyrrhalta, but this species differs from all other Pyrrhalta in having anterior and posterior borders of prothorax unimagined, therefore this question needs further study. Pyrrhalta hanunga Samoderzenkov, 1988 Remark. Description of this species was based on a single male and according to description it has labrum, spot on vertex and 4 spots on prothorax. In reality in the holotype labrum is fulvous, only apices of mandibles are black, middle of vertex is a little darkened, but without a real spot, on prothorax there are 4 dark fulvous elevated areas which scarcely differs from main color, very poorly

5 A key for identification with description of new species of the genus Pyrrhalta Joannis, 1865 from Indochina 271 Figs Aedeagi of species of the genus Pyrrhalta Joannis, Рис Эдеагусы видов рода Pyrrhalta Joannis, P. gorbunovi sp. n.; 11 P. unicostata Pic, 1937; 12 P. impressicollis Samoderzhenkov, 1988; 13 P. fedorenkoi sp. n.; 14 P. ornata sp. n.; 15 P. curticollis Samoderzhenkov, 1988; 16 P. basifasciata Samoderzhenkov, 1988; 17 P. ornatipennis Samoderzhenkov, 1988; 18 P. kabakovi Samoderzhenkov, 1988; 19 P. nigrotibialis L. Medvedev, 2010; 20 P. apicicornis L. Medvedev, 2012; 21 P. corpulenta Gressitt et Kimoto, 1963; 22 P. hanunga Samoderzhenkov, 1988; 23 P. nuibata L. Medvedev, 2012; 24 P. laosica sp. n.

6 272 Figs Color of antennae of species of the genus Pyrrhalta Joannis, Рис Окраска усиков видов рода Pyrrhalta Joannis, P. fedorenkoi sp. n.; 26 P. nigrotibialis L. Medvedev, 2010; 27 P. apicicornis L. Medvedev, 2012; 28 P. hanunga Samoderzhenkov, 1988; 29, 30 P. nuibata L. Medvedev, 2012; 31, 32 P. laosica sp. n. delimited and can not be named as spots. To the moment I have 6 more specimens of P. hanunga which have head and prothorax entirely fulvous. Poneridia costata (Allard, 1889) = Galerucella costata Allard, Remark. Galerucella costata Allard, 1889, till now known only with type, was removed in the Australian genus Poneridia Weise, 1908 [Laboissière, 1935] and this was accepted also by Kimoto [1989]. This genus includes 6 species from Australia and neighbouring regions, but it looks very strange that one species inhabits Vietnam. But according to original description it seems that this species is identical with Pyrrhalta unicostata (Pic, 1937). Additional investigation of type is necessary. Ackowledgements Author is very much obliged to Jan Bezděk (Mendel University in Brno, Czech Republic) for valuable information and review of the work. References Catalogue of Palaearctic Coleoptera (I. Löbl, A. Smetana eds.). Vol. 6. Chrysomeloidea Stenstrup: Apollo Books. 924 p. Gressitt J.L., Kimoto S The Chrysomelidae (Coleopt.) of China and Korea, Part 2. Pacific Insects Monograph. 1B: Kimoto S Chrysomelidae of Thailand, Cambodia, Laos and Vietnam. IV. Galerucinae. Esakia. 27: Laboissière V Rectification au mèmoire de E. Allard Contributions a la fauna Indo-Chinoise, Galerucides et Alticides. Annales de l Association des Naturalistes de Levallois-Perret. 21 [ ]: Medvedev L.N. 2012a. New species of Chrysomelidae (Coleoptera) from Indochina. Euroasian Entomological Journal. 11(1): Medvedev L.N. 2012b. New and interesting Chrysomelidae (Insecta, Coleoptera) from the collection of Naturkundemuseum Erfurt. Veröffentlichungen des Naturkundemuseums Erfurt. 31: Ogloblin D.A Fauna SSSR. Nasekomye zhestkokrylye. T. 26. Vyp. 1. Listoedy, Galerucinae [Fauna of the USSR. Beetles. Vol. 26. Iss. 1. Chrysomelidae, Galerucinae]. Moscow Leningrad: Academy of Sciences of the USSR Publ. 457 p. (in Russian). Samoderzhenkov E.V Leaf beetles of the tribe Galerucini (Chrysomelidae, Galerucinae) in fauna of Vietnam. In: Fauna i ekologiya nasekomykh Vietnama [Fauna and ecology of Vietnam insects]. Moscow: Nauka: (in Russian). Skomorokhov M.O To the knowledge of Chrysomelidae (Coleoptera) from the islands of southern Vietnam. Russian Entomological Journal. 20(2): Sprecher-Uebersax E., Zoia S Pyrrhalta medvedevi sp. nov., a new species from the Nepal Himalayas (Coleoptera, Chrysomelidae, Galerucinae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft. 75(3/4):