Education. Positions held

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1 Education 2004 Ph.D. Ecology, University of Georgia, with Dr. J. Bruce Wallace 1999 M.S. Biology, University of Alabama, with Dr. Arthur C. Benke 1994 B.A. Biology, University of Colorado at Boulder Positions held present Postdoctoral Associate, Department of Zoology and Physiology, University of Wyoming (with Dr. Robert O. Hall, Jr.) 2005 Research Experience for Undergraduates program (REU-NSF) mentor, El Verde Field Station, Puerto Rico Postdoctoral researcher, Institute of Ecology, University of Georgia, El Verde Field Station, Puerto Rico (with Dr. Alan P. Covich) 2003 Ecology lecturer, Institute of Ecology, University of Georgia Teaching assistant, University of Georgia Research assistant, University of Georgia Teaching assistant, University of Alabama Research Technician, Center for Limnology, University of Colorado (with Dr. William M. Lewis Jr.) Experiential outdoor educator, University of Colorado Research interests Land-use effects on aquatic food webs and ecosystems Interactions and feedbacks among nutrients, food webs, and ecosystems Cross-ecosystem material subsidies Ecological Stoichiometry Publications Wyatt F. Cross Department of Zoology and Physiology University of Wyoming, Laramie, WY, Phone: , Fax: , wcross@uwyo.edu Mailing address: USGS, Grand Canyon Monitoring and Research Center 2255 N. Gemini Dr., Flagstaff, AZ, Cross, W. F., J. B. Wallace, A. D. Rosemond Nutrient enrichment reduces constraints on material flows in a detritus-based ecosystem. Ecology in press. Cross, W. F., A. Ramírez, A. Santana*, and L. Silvestrini Santiago*. Egestion rates of tropical stream consumers: towards separating the importance of consumption and bioturbation. Biotropica (in revision). *undergraduate authors. Greenwood, J. L., A. D. Rosemond, J. B. Wallace, W. F. Cross, and H. S. Weyers Nutrients stimulate leaf breakdown rates and detritivore biomass: bottom-up effects via heterotrophic pathways. Oecologia, in press.

2 Cross, W. F., J. B. Wallace, A. D. Rosemond, and S. L. Eggert Whole-system nutrient enrichment increases secondary production in a detritus-based ecosystem. Ecology 87: *Frost, P. C., J. P. Benstead, W. F. Cross, H. Hillebrand, J. H. Larson, M. A. Xenopoulus, and T. Yoshida Threshold elemental ratios of carbon and phosphorus in aquatic consumers. Ecology Letters 9: *Recommended by faculty of 1000 ( Johnson, B. J., J. B. Wallace, A. D. Rosemond, and W. F. Cross Larval salamander growth responds to enrichment of a nutrient poor headwater stream. Hydrobiologia 573: Cross, W. F., A. D. Rosemond, J. P. Benstead, S. L. Eggert, and J. B. Wallace Differential effects of consumers on C, N, and P dynamics: insights from long-term research. In: Dynamic Food Webs: Multispecies Assemblages, Ecosystem Development, and Environmental Change. deruiter, P. C., V. Wolters, and J. C. Moore (eds.). Elsevier. Scharler, U. M., F. D. Hulot, D. J. Baird, W. F. Cross, J. Culp, C. A. Layman, D. Raffaelli, M. Vos, and K. O. Winemiller Central issues for aquatic food webs: from chemical cues to whole system responses. In: Dynamic Food Webs: Multispecies Assemblages, Ecosystem Development, and Environmental Change. deruiter, P. C., V. Wolters, and J. C. Moore (eds.). Elsevier. Cross, W. F., J. P. Benstead, P. C. Frost, and S. A. Thomas Ecological stoichiometry in freshwater benthic systems: recent progress and perspectives. Freshwater Biology 50: Cross, W. F., B. R. Johnson, J. B. Wallace, and A. D. Rosemond Contrasting response of stream detritivores to long-term nutrient enrichment. Limnology and Oceanography 50: Frost, P. C., W. F. Cross, and J. P. Benstead Ecological stoichiometry in freshwater benthic ecosystems: an introduction. Freshwater Biology 50: Cross, W. F., J. P. Benstead, A. D. Rosemond, and J. B. Wallace Consumerresource stoichiometry in detritus-based streams. Ecology Letters 6: Johnson, B., W. F. Cross, and J. B. Wallace Long-term resource limitation reduces insect detritivore growth in a headwater stream. Journal of the North American Benthological Society 22: Cross, W. F., and A. C. Benke Intra- and interspecific competition among coexisting lotic snails. Oikos 96:

3 In preparation Benstead, J. P., A. D. Rosemond, J. B. Wallace, S. E. Eggert, K. Suberkropp, V. Gulis, P. J. Mulholland, J. L. Greenwood, W. F. Cross, and H. S. Weyers. Nitrogen and phosphorus loading alters the carbon budget of a headwater stream ecosystem. Target journal: Ecology. Rosemond, A. D., K. Suberkropp, W. F. Cross, J. Greenwood, V. Gulis, J. P. Benstead, H. Weyers, S. L. Eggert, R. Hilten, P. J. Mulholland, and J. B. Wallace. Nutrient enrichment of detritus-based stream ecosystems. Target journal: BioScience. Cross, W. F., A. P. Covich, A. Ramirez, T. A. Crowl. Production, longevity, and trophic support of freshwater shrimps in two tropical headwater streams. Target journal: Limnology and Oceanography. Benstead, J. P., W. F. Cross, J. G. March, A. Ramirez, A. P. Covich, W. H. McDowell. Consumer-driven nutrient recycling and its effects on whole-ecosystem flux in tropical streams. Target journal: Ecology Letters Teaching Experience Lecture 2003 General ecology, ECOL 3500, University of Georgia Guest lecturer for Trophic Interactions course, University of Georgia. Laboratory Teaching assistant: The Marine Environment I (physical and chemical), The Marine Environment II (biological), University of Georgia Teaching assistant: Ocean Science, General Biology, Human Anatomy and Physiology, University of Alabama. Student mentoring 2005 Research Experience for Undergraduates, Puerto Rico: Alexandra Santana, Role of freshwater shrimp in cycling of fine particulate organic matter award winner at the SACNAS meeting in Denver, Co Supervised 10 undergraduate research assistants, University of Georgia Mentored 2 undergraduate independent research projects, University of Georgia: Anthony Cohen, Leaf litter breakdown and invertebrate community structure in a nutrient-enriched stream ecosystem - presented at university-wide undergraduate research symposium; Catherine Evans, Organic matter dynamics in a nutrient-enriched stream ecosystem Experiential education Experiential outdoor educator. University of Colorado Outdoor Program. Courses taught: Fly-fishing, Fly-tying, Basic Entomology, Freshman Wilderness Program, Mountain Oriented First Aid (assistant), Backpacking, and Wilderness Ethics. 3

4 Awards and grants 2005 National Science Foundation cross-site LTER working group The importance of consumer-driven nutrient recycling in streams across a climatic gradient, $8, Best graduate student publication, Institute of Ecology, University of Georgia, (Ecology Letters 6: ) Institute of Ecology nominee for university-wide Graduate School Excellence in Research award (not received) University-Wide Dissertation Completion Award, University of Georgia Dean Lindholm Memorial Award, Institute of Ecology, University of Georgia Sigma Xi Scientific Research Society, Grant-in-Aid of Research, $ Best Student Presentation, 50 th annual meeting of North American Benthological Society, Pittsburg, Pennsylvania Best Student Presentation, Institute of Ecology Research Symposium, University of Georgia Chermock Award, Outstanding Graduate Student, University of Alabama Travel Award, University of Alabama Graduate School. Invited Seminars 2006 Cross, W. F. Linking consumers and nutrients in temperate and tropical stream ecosystems. University of Wyoming, Department of Zoology and Physiology Cross, W. F. Reciprocal dynamics of nutrients and consumers in stream ecosystems. University of California, Santa Cruz, Department of Ecology and Evolutionary Biology Cross, W. F. Reciprocal dynamics of nutrients and stream communities in Temperate and tropical ecosystems. Grand Canyon Monitoring and Research Center, USGS, Flagstaff, Arizona Cross, W. F. Reciprocal dynamics of nutrients and stream communities in temperate and tropical ecosystems. University of Hawaii at Hilo. Department of Biology Cross, W. F. Examining the role of tropical freshwater shrimps in stream ecosystem dynamics. International Institute for Tropical Forestry, U.S. Forest Service, Puerto Rico Cross, W. F. Effects of nutrient enrichment on a detritus-based stream ecosystem. Université du Québec à Trois Rivières. Department of Chemistry and Biology Cross, W. F., A. D. Rosemond, J. P. Benstead, S. L. Eggert, and J. B. Wallace. Differential effects of consumers on C, N, and P dynamics: insights from long-term research. Food Web Symposium 2003, Giessen Germany. 4

5 Presentations 2007 Cross, W. F., C. V. Baxter, R. O. Hall, T. A. Kennedy, S. Rogers, E. J. Rosi-Marshall, M. Yard. Using ecosystem production budgets to assess resource limitation of native and non-native fishes in the Colorado River, Grand Canyon. Upcoming North American Benthological Society meeting in Columbus, SC Kennedy, T. A., R. O. Hall, E. J. Rosi-Marshall, W. F. Cross, B. Ralston. Allochthonous organic matter from tributary floods dominates organic matter inputs to the Colorado River, Grand Canyon. Upcoming North American Benthological Society meeting in Columbus, SC Hall, R. O., T. Kennedy, E. J. Rosi-Marshall, W. F. Cross, M. D. Yard, C. V. Baxter. Open-channel estimates of primary production in the Colorado River, Grand Canyon. Annual meeting of the American Society for Limnology and Oceanography, Santa Fe, NM Davis, J. M., A. D. Rosemond, W. F. Cross, S. L. Eggert, and J. B. Wallace. Effects of long-term nutrient enrichment on macroinvertebrate biomass, abundance, and community composition of a detrital-based system. Annual meeting of Ecological Society of America, Memphis, TN Cross, W. F., A. P. Covich, A. Ramirez, M. A. Chadwick, and T. A. Crowl. Growth rates, longevity, and production of tropical freshwater shrimps in two headwater streams of Puerto Rico. 54 rd annual meeting of the North American Benthological Society, Anchorage, AK Covich, A. P., K. S. Fraiola, W. F. Cross, and T. A. Crowl. Beyond species richness: biodiversity and detrital processing by long-lived decapod crustaceans in tropical headwater streams. 54 rd annual meeting of the North American Benthological Society, Anchorage, AK Rosemond, A. D., V. I. Gulis, J. L. Greenwood, W. F. Cross, J. M. Davis, K. Suberkropp, and J. B. Wallace. Consequences of nutrient enrichment to biodiversity in headwater streams: an assessment of effects on multiple taxonomic groups. 54 rd annual meeting of the North American Benthological Society, Anchorage, AK Davis, J. M., A. D. Rosemond, W. F. Cross, S. L. Eggert, and J. B. Wallace. Effects of long-term nutrient enrichment on macroinvertebrate biomass, abundance, and community composition of a detrital-based system. 54 rd annual meeting of the North American Benthological Society, Anchorage, AK Santana, A., L. Silvestrini, W. F. Cross, and A. Ramirez. Consumer-mediated sediment removal in tropical streams: estimating the relative importance of consumption versus non-consumptive bioturbation. 54 rd annual meeting of the North American Benthological Society, Anchorage, AK. 5

6 2006 Silvestrini L., A. Santana, A. Ramirez, and W. F. Cross. Egestion rates of tropical stream consumers and implications for the cycling of fine particulate material. 54 rd annual meeting of the North American Benthological Society, Anchorage, AK Cross, W. F., J. P. Benstead, J. G. March, A. Ramirez, W. H. McDowell, and A. P. Covich. Consumer-driven nutrient recycling and its effect on whole-ecosystem flux in tropical streams. 53 rd annual meeting of the North American Benthological Society, New Orleans, LA Covich, A. P., T. A. Crowl, W. F. Cross, T. Heartsill-Scalley, K. M. Fraiola, and K. Hein. The importance of scale in tropical headwater stream food webs: loss of trophic diversity and detrital processing by decapod crustaceans. Annual meeting of Ecological Society of America, Montreal, Canada Frost, P. C., J. P. Benstead, W. F. Cross, H. Hillebrand, J. H. Larson, C. MacKenzie, M. A. Xenopoulos, and T. Yoshida. Threshold carbon:phosphorus ratios of aquatic organisms: relative effects of metabolism and elemental composition. Summer Meeting of the American Society of Limnology and Oceanography, Santiago de Compostela, Spain Frost, P. C., J. P. Benstead, W. F. Cross, H. Hillebrand, J. H. Larson, C. MacKenzie, M. A. Xenopoulos, and T. Yoshida. Threshold carbon:phosphorus ratios of aquatic consumers: effects of growth rate, metabolism, and body elemental composition. Annual meeting of Ecological Society of America, Montreal, Canada Covich, A. P., T. A. Crowl, T. Heartsill-Scalley, J. Bithorn, W. F. Cross, K. M. Fraiola, and K. Hein. The role of decapods in ecosystem function in Puerto Rico streams. Summer Meeting of the American Society of Limnology and Oceanography, Santiago de Compostela, Spain Wallace, J. B., W. F. Cross, S. L. Eggert, J. L. Meyer, and A. D. Rosemond. 28 years of secondary production in headwater streams at Coweeta Hydrologic Laboratory. 53 rd annual meeting of the North American Benthological Society, New Orleans, LA Rosemond, A. D., K. Suberkropp, W. F. Cross, J. L. Greenwood, S. L. Eggert, N. Taylor, and J. B. Wallace. Top-down feedback exacerbates effects of nutrient enrichment in detritus-based ecosystems via increased loss of basal carbon. 53 rd annual meeting of the North American Benthological Society, New Orleans, LA Cross, W. F., J. B. Wallace, A. D. Rosemond. Effects of nutrient enrichment on the trophic basis of production of detritus-based headwater streams. 52 nd annual meeting of the North American Benthological Society, Vancouver, British Colombia Rosemond, A. D., J. P. Benstead, J. W. Greenwood, K. Suberkropp, R. Hilten, S. L. Eggert, W. F. Cross, H. S. Weyers, and J. B. Wallace. Nutrient enrichment affects carbon dynamics in a detritus-based stream. 52 nd annual meeting of the North American Benthological Society, Vancouver, British Colombia. 6

7 2003 Rosemond, A. D., K. Suberkropp, W. F. Cross, J. L. Greenwood, V. Gulis, P. J. Mulholland, J. B. Wallace, H. S. Weyers, J. P. Benstead, and R. Hilten. Relative response of consumers, resources, and ecosystem function to nutrient enrichment in a detritus-based system. 88 th annual meeting of the Ecological Society of America, Savannah, Georgia Cross, W. F., J. P. Benstead, A. D. Rosemond, and J. B. Wallace. Ecological stoichiometry of a detritus-based stream food web. 51 st annual meeting of the North American Benthological Society, Athens, Georgia Greenwood, J. L., A. D. Rosemond, J. B. Wallace, W. F. Cross, H. S. Weyers. Response of leaf breakdown to nutrient enrichment in an Appalachian headwater stream. 51 st annual meeting of the North American Benthological Society, Athens, Georgia Cross, W. F., A. D. Rosemond, J. P. Benstead, J. L. Greenwood, P. J. Mullholland, K. Suberkropp, J. B. Wallace, H. S. Weyers. Stream nutrient addition experiment. Coweeta Hydrologic Laboratory Annual Meeting, University of Georgia, Athens, Georgia Rosemond, A. D., J. P. Benstead, K. Suberkropp, W. F. Cross, P. J. Mullholland, J. L. Greenwood, J. B. Wallace. Does nutrient enrichment increase carbon limitation of a detritus-based stream food web? 87 th annual meeting of the Ecological Society of America Meeting, Tuscon, Arizona Benstead, J. P., A. D. Rosemond, J. B. Wallace, K. Suberkropp, J. L. Greenwood, and W. F. Cross. Long-term nutrient enrichment alters organic matter dynamics in a headwater stream. 50 th annual meeting of the North American Benthological Society, Pittsburgh, Pennsylvania Cross, W. F., J. B. Wallace, A. D. Rosemond, and B. Johnson. Effects of nutrient loading on invertebrate growth rates and secondary production. 50 th annual meeting of the North American Benthological Society, Pittsburgh, PA Cross, W. F., S. Eggert, E. R. Siler, and J. B. Wallace. Variation in secondary production of southeastern headwater streams: estimating the relative importance of biotic and abiotic factors. Coweeta Hydrologic Laboratory Annual Meeting, University of Georgia, Athens, Georgia Cross, W. F., S. Eggert, E. R. Siler, and J. B. Wallace. Variation in secondary production of southeastern headwater streams: estimating the relative importance of biotic and abiotic factors. 49 th annual meeting of the North American Benthological Society, La Crosse, Wisconsin Greenwood, J. L., W. F. Cross, H. S. Weyers, A. D. Rosemond, and J. B. Wallace. Initial response of leaf litter processing to nutrient enrichment in an Appalachian headwater stream. 49 th annual meeting of the North American Benthological Society, La Crosse, Wisconsin Wallace, J. B., W. F. Cross, S. Eggert, and E. R. Siler. Seventeen years of 7

8 abundance, biomass, and secondary production from three headwater streams: trophic structure and interannual variability. International Society for Theoretical and Applied Limnology (SIL), Melbourne, Australia Cross, W. F., and A. C. Benke. Competition among coexisting lotic snails. 47 th annual meeting of the North American Benthological Society, Duluth, Minnesota. Professional Activities 2006 present Participant in the development of National Ecological Observatory Network aquatic studies for the Pacific region 2000 present Ad-hoc reviewer: Archiv für Hydrobiologie, Aquatic Ecology, Ecological Applications, Ecology Letters, Environmental Conservation, Freshwater Biology, Journal of the North American Benthological Society, Limnology and Oceanography, Oecologia, Wetlands 2000 present Member of Ecological Society of America, North American Benthological Society 2005 Guest editor of special issue in Freshwater Biology: Ecological stoichiometry of freshwater benthic ecosystems Co-organized special session on Ecological Stoichiometry for the 52 nd annual meeting of the North American Benthological Society, Athens, Georgia Organized student-run ecology symposium series, Peanut Butter Club, University of Georgia. 8

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