Predation-mediated recruitment in the Georges Bank fish community

Save this PDF as:
 WORD  PNG  TXT  JPG

Size: px
Start display at page:

Download "Predation-mediated recruitment in the Georges Bank fish community"

Transcription

1 ICES Journal of Marine Science, 58: doi:1.16/jmsc , available online at on Predation-mediated recruitment in the Georges Bank fish community T. S. Tsou and J. S. Collie Tsou, T. S., and Collie, J. S. 21. Predation-mediated recruitment in the Georges Bank fish community. ICES Journal of Marine Science, 58: Understanding the mechanisms that control recruitment in marine fish populations is important for conservation and also for more efficient management through improved predictions of abundance. We examined the role of predation in regulating recruitment by applying multispecies virtual population analysis (MSVPA) to eight fish species on Georges Bank. Cod and silver hake were the most important predators and herring and silver hake were the most important prey species. Predation on cod and silver hake was compensatory, because cannibalism was the dominant source of predation for these species. Predation on herring and mackerel appeared to be slightly depensatory or density-independent. Predation on young fish also affected the relative cohort size at age 2. Therefore, predation should be considered when making medium- to long-term recruitment forecasts. 21 International Council for the Exploration of the Sea Keywords: Georges Bank, MSVPA, predation, recruitment. Published electronically 1 August 21. T.-S. Tsou and J. S. Collie: Graduate School of Oceanography, University of Rhode Island, Narragansett, RI 2882, USA. Present address T.-S. Tsou: Florida Marine Research Institute, 1 8th Avenue SE, St. Petersburg, FL 3371, USA; Tel: ; fax: ; Introduction Predation among exploited fish species contributes significantly to their natural mortality and varies from year to year (Pope, 1991). The magnitude and the variability of predation on young fish play an important role in regulating species abundance in many marine ecosystems (Bax, 1991). Therefore, understanding how species interact is essential for making long-term predictions about multispecies fisheries. In the North Sea, for example, decreasing the fishing mortality on cod, haddock, whiting, and saithe would have the contraintuitive result of reducing overall yield because of increased predation (Pope, 1991). The combined effects of fishing and predation can be estimated with multispecies virtual population analysis (MSVPA), which provides useful insight into the role of predator prey interactions by quantifying the food consumption of major predators (Sparre, 198). MSVPA simultaneously estimates fishing and predation mortality by year and age group for any number of predator and prey species on the basis of catch-at-age and stomach-content data. MSVPA also estimates (constant) predator prey suitability coefficients, which can be used to forecast predation under varying abundance of predators and prey. In MSVPA, natural mortality is a function of predation. This is a significant departure from the singlespecies philosophy, which generally assumes constant natural mortality over all age groups and years (or a constant natural mortality age-array over all years). To this end, natural mortality is partitioned into predation mortality and residual natural mortality caused by other factors. is a dynamic function of predator consumption, predator preference, predator abundance, and prey abundance, and is considered to be the dominant source of mortality on young fish (Sparre, 1991). The residual mortality is usually treated as a constant and is considered to be much lower than predation mortality. Predation was identified as the largest source of mortality of pre-recruit fish on Georges Bank (Sissenwine et al., 1984), which is considered to be a predator-controlled ecosystem (Sissenwine, 1986; Overholtz et al., 1991). As part of the Coastal Ocean Program, Georges Bank predation study, we applied MSVPA to estimate predation mortality and to reconstruct the abundance-at-age of six important fish species (for details, see Tsou and Collie, 21). Here, our focus is the role of predation on young fish with respect to recruitment /1/ $35./ 21 International Council for the Exploration of the Sea

2 Predation-mediated recruitment in the Georges Bank fish community 995 Male dogfish Female dogfish Winter skate Haddock Materials and methods Silver hake MSVPA integrates the suitability concept of Andersen and Ursin (1977) for describing predator prey relationships with the traditional form of virtual population analysis (VPA). Sparre (1991) outlines the five basic equations of MSVPA. The first two equations represent the well-established basis for single-species VPA (Gulland, 1965), except that the natural mortality by age group is partitioned into predation mortality and residual natural mortality. can be estimated from the other three equations, which involve biomass-at-age of all species, ingestion ratesat-age for all predators, prey and predator-specific suitability indices (based on stomach content data for individual years or averages over a period), and some estimate of other food (not represented by the modelled species). Following the approach of Sparre (198), we assumed a constant biomass of the ecosystem, such that the expected changes in other food compensate for changes in the abundance of exploited prey. The total biomass of fish and their prey (15 million tonnes) was estimated with an energy budget of the Georges Bank ecosystem (Sissenwine et al., 1984). Results of the MSVPA were insensitive to the assumed level of other food (Tsou and Collie, 21). We performed an eight-species MSVPA of the Georges Bank fish community (Figure 1) for As predators, we included cod (Gadus morhua), silver hake (Merluccius bilinearis), haddock (Melanogrammus aeglefinus), spiny dogfish (Squalus acanthias), and winter skate (Raja ocellata); as prey, cod, silver hake, herring (Clupea harengus), mackerel (Scomber scombrus), and yellowtail flounder (Limanda ferruginea). Spiny dogfish and winter skate were incorporated as other predators, whose abundances were estimated external to the MSVPA from survey data (Tsou and Collie, 21). Inputs are quarterly catch-at-age, terminal Cod Yellowtail flounder Herring Mackerel Figure 1. Partial food web for the Georges Bank fish community as included in the MSVPA (the arrows point from prey to predator species). fishing mortality rates, residual natural mortality rates (.2 for age 1+;.3 for -group), mean body weights, predator-stomach contents, predator consumption rates, and an estimate of the total biomass of the ecosystem. We adopted catch-at-age data and terminal fishing mortalities from single-species stock-assessment documents. As silver hake, herring, and mackerel are assessed over a wider geographic area than Georges Bank, their catches were pro-rated by their average biomass proportions on Georges Bank (.19,.121, and.126, respectively; Collie and DeLong, 1999). Consumption rates by predator age were from previous studies: dogfish from Jones and Geen (1977); cod, silver hake, and haddock from Grosslein et al. (198); and winter skate from Nelson (1993). Diet composition by predator and prey size/age categories is the foundation of MSVPA. Since beginning in 1968, thousands of stomach samples from marine fish have been gathered along the northeast coast of North America and analysed. We extracted stomach-content data from the food-habit database. The stomach-content data for were tuned to the simulated populations in those years to obtain the suitability indices. A prey item was included in the analysis only if it was identified to species level and constituted more than.1% of a predator s diet composition over the sampling period and study area. Predator size was recorded for each stomach; however, only a small fraction of the stomach contents has been classified by size/age class of prey. To circumvent this problem, predator size-preference functions of cod, silver hake, haddock, dogfish, and winter skate were fitted with gamma distributions to distribute the ingested prey into size categories (Tsou and Collie, 21). Distributions by size classes were further converted into age classes based on growth curves. Results on young fish was high compared with fishing mortality and the constant natural mortality rate of.2 that is generally assumed for these stocks (Figure 2). Averaged across years, age and age 1 silver hake experienced the highest predation mortality among the six prey species: approximately 1.1 for age and 1.6 for age 1 silver hake. Cannibalism was the dominant source of predation mortality and accounted, on average, for 7% and 6% of predation mortality on age and age 1 silver hake, respectively. Haddock was another important predator of age silver hake, accounting for 15% of the predation, while cod was responsible for 3% of the predation mortality on age 1. Based on the estimated predator prey size preference function (Tsou and Collie, 21), cod prey heavily on age 1 silver hake, resulting in the higher predation

3 996 T. S. Tsou and J. S. Collie Cod Silver hake Haddock Mackerel Prey species Herring Yellowtail flounder Figure 2. Predation mortalities of age (closed) and age 1 (hatched) prey as estimated from MSVPA, averaged over years. mortality than on age. Predation on age and age 1 cod from the five predators was not high (average.35 and.2, respectively). The most important predator of age cod was silver hake, which accounted for more than 4% of the predation. Cannibalism and dogfish were each responsible for another 3%. Age 1 cod experienced lower predation than age, the dominant source being cannibalism. Haddock experienced high predation at age (1.1) and cod was its main predator. Average predation on young mackerel was low because its abundance increased sharply during the period, predators preferred older mackerel based on our predator size preference functions, and mackerel was infrequently identified to species level in the stomach-content database. Herring was another important prey species that experienced predation mortality of.2 at age and of.7 at age 1. Silver hake was the dominant predator (8% of predation mortality). Predation on age 1 was three times higher than on age because of the difference in preference of silver hake. Yellowtail flounder experienced very low age predation mortality and a predation mortality rate of.2 at age 1. This is because its major predator, dogfish, preferred the size of age 1 yellowtail flounder and age was not selected according to the size-preference function. Although yellowtail flounder constituted a small proportion of the spiny dogfish diet, predation mortality was substantial because dogfish far outnumbered the flounder. When summed over the age groups, the proportion of biomass consumed by predators increased with stock Cod Silver hake Proportion consumed Haddock Mackerel Herring.2 Yellowtail flounder Stock biomass (kt) Figure 3. Relationships between the proportion of biomass consumed and prey biomass as estimated from MSVPA (solid lines: smoothed with super smoother in S-Plus). 25

4 Predation-mediated recruitment in the Georges Bank fish community Age abundance (millions) Age abundance (millions) Age 2 abundance (millions) Figure 4. Silver hake: age 1 vs. age and age 2 vs. age 1 abundance (left panels; broken line: average survival rate) and predation mortality vs. age and age 1 abundance (right panels; solid line: smoothed values) Age abundance (millions) Age abundance (millions) 8 Age 2 abundance (millions) Figure 5. Herring: as in Figure 4. biomass for cod, silver hake, and yellowtail flounder, whereas the proportion declined with increasing stock biomass for haddock, mackerel, and herring (Figure 3). The increasing trends for cod and silver hake might be caused by cannibalism. The decreasing trends for mackerel and herring may indicate predator saturation

5 998 T. S. Tsou and J. S. Collie Age abundance (millions) Age abundance (millions) 12 Age 2 abundance (millions) Figure 6. Cod: as in Figure Age abundance (millions) Age abundance (millions) 5 Age 2 abundance (millions) Figure 7. Haddock: as in Figure 4 (the two largest year classes were excluded from the plot). because the abundance of pelagics has increased sharply since the mid-198s. The MSVPA equations predict that if predator abundance is constant predation rates should decline as a function of prey stock biomass. In the case of cannibalism, predation rates increase with stock biomass because predator abundance is a multiplier.

6 Predation-mediated recruitment in the Georges Bank fish community 999 Predation from cannibalism Predation from cannibalism (a) (b) Stock biomass Figure 8. on age (open circle) and age 1 (closed triangle) resulting from cannibalism for (a) cod and (b) silver hake. Year-class sizes estimated at different ages were substantially modified by age and age 1 predation. Because predation was high for both age and age 1 silver hake, the relationships of age 1 vs. age and age 2 vs. age 1 abundance were scattered along the mean mortality line (Figure 4). The abundant 1979 cohort of age silver hake experienced above average predation and thus did not result in a large cohort at age 1. Conversely, the 1986 year class experienced low predation mortality as 1-group, resulting in the largest cohort at age 2. For herring, the relationship between age 1 and age abundance was almost linear (Figure 5). was somewhat variable, but had only minor effects on relative year-class size. The relationship between age 2 and age 1 herring was more scattered because age 1 experienced predation mortality three times that of age. For cod (Figure 6), the relationships were fairly linear, because predation mortality was about the same as residual mortality. This was also the case for age 1 haddock (Figure 7), but the age 1 vs. age relationship was more scattered because of the high predation mortality on -group. on age and age 1 silver hake appeared to increase with prey abundance (Figure 4). This compensatory mortality would be expected when cannibalism is a significant component of the total predation. Age 1 cod also experienced a high cannibalism rate and exhibited compensatory mortality as well (Figure 6). In contrast, predation mortality on juvenile herring (Figure 5) and juvenile haddock (Figure 7) showed a weak negative relationship with their abundance, suggesting density independent or weakly depensatory predation. Cannibalism was important for cod and silver hake, but to a different extent among years (Figure 8). Cannibalism of cod was high before 1983 and low afterward, except in Cannibalism increased slightly with increasing stock biomass. Rates of cannibalism in silver hake were more variable for age than for age 1, but both showed a clear increasing trend with stock biomass. Discussion The MSVPA results agree with previous conclusions that predation is a dominant source of mortality in the Georges Bank fish community (Sissenwine et al., 1984; Fogarty and Murawski, 1998; Collie and DeLong, 1999). Strong interactions exist between cod and silver hake as predators and silver hake and herring as prey, while weaker trophic interactions exist for haddock, yellowtail flounder, dogfish, and winter skate. A comparison of age 1 recruits estimated from single species VPA and MSVPA (Figure 9) indicates a difference up to one order of magnitude for some prey species. For herring and yellowtail flounder, which experienced most predation at age 1, the relationship was more scattered than for other species. The results obtained are conditioned on the assumptions and input data for MSVPA. Silver hake abundance is uncertain because an analytical stock assessment is lacking. estimates were sensitive to the input values of predator consumption rates and the size-preference functions influenced the distribution of predation among prey age groups (Tsou and Collie, 21). There is also unobserved mortality of fish larvae and juveniles that is not included in the MSVPA owing to their absence from food-habits data. Nevertheless, our results suggest that predation mortality is large and variable enough to affect year-class size of herring and silver hake significantly. Therefore, predation mortality should be accounted for when making medium- and long-term forecasts of recruitment. Acknowledgements We thank Michael Fogarty for his advice on multispecies analysis and Rodney Rountree, Jason Link, and Lance Garrison for their help in accessing the NEFSC food-habits database; Jack Finn provided the original MSVPA FORTRAN program. The study was funded by the NOAA Coastal Ocean Program, Georges Bank Predation Study, NA36RG474.

7 1 T. S. Tsou and J. S. Collie Cod Haddock Herring 5 Yellowtail flounder SSVPA recruits Mackerel SSVPA recruits Figure 9. Comparison between age 1 recruits (millions) estimated from single-species VPA (SSVPA) and MSVPA (extremely high recruitment of mackerel in 1979 and herring in 1983 were excluded from the plot). References Andersen, K. P., and Ursin, E A multispecies extension to the Beverton and Holt theory of fishing; with accounts of phosphorus, circulation and primary production. Meddelelser fra Danmarks Fiskeri- og Havundersgelser. N.S., 7: Bax, N. J A comparison of the fish biomass flow to fish, fisheries, and mammals in six marine ecosystems. ICES Marine Science Symposia, 193:

8 Predation-mediated recruitment in the Georges Bank fish community 11 Collie, J. S., and DeLong, A. K Multispecies interactions in the Georges Bank fish community. Ecosystem approaches for fisheries management, Alaska Sea Grant College Program, AK SG 99 1: Fogarty, M. J., and Murawski, S. A Large-scale disturbance and the structure of marine systems: fisheries impacts on Georges Bank. Ecological Applications, 8: S6 S22. Grosslein, M. D., Langton, R. W., and Sissenwine, M. P Recent fluctuations in pelagic fish stocks of the Northwest Atlantic, Georges Bank region, in relation to species interactions. Rapports et Procès-Verbaux des Réunions du Conseil International pour l Exploration de la Mer, 177: Gulland, J. A Estimation of mortality rates. Annex to Arctic Fisheries Working Group Report. ICES CM 1965: Doc. 3, 9 pp. Jones, B. C., and Geen, G. H Food and feeding of spiny dogfish (Squalus acanthias) in British Columbia waters. Journal of the Fisheries Research Board of Canada, 34: Nelson, G. A The potential impacts of skate abundances upon the invertebrate resources and growth of yellowtail flounder (Pleuronectes ferrugineus) on Georges Bank. Doctoral dissertation, University of Massachusetts. Overholtz, W. J., Murawski, S. A., and Foster, K. L Impact of predatory fish, marine mammals, and seabirds on the pelagic fish ecosystem of the northeastern USA. ICES Marine Science Symposia, 193: Pope, J. G The ICES Multispecies Assessment Working Group: Evolution, Insights, and Future Problems. ICES Marine Science Symposia, 193: Sissenwine, M. P Perturbation of a predator-controlled continental shelf ecosystem. In Variability and Management of Large Marine Ecosystems, pp Ed. by K. Sherman, and L. M. Alexander. American Association for the Advancement of Science, Selected Symposium Series, no. 99. Sissenwine, M. P., Cohen, E. B., and Grosslein, M. D Structure of the Georges Bank ecosystem. Rapports et Procès-Verbaux des Réunions du Conseil International pour l Exploration de la Mer, 183: Sparre, P A goal function of fisheries (Legion Analysis). ICES CM 198/G: 4. Sparre, P Introduction to multispecies virtual population analysis. ICES Marine Science Symposia, 193: Tsou, T. S., and Collie, J. S. 21. Estimating predation in the Georges Bank fish community. Canadian Journal of Fisheries and Aquatic Sciences, 58:

9.4.5 Advice September Widely distributed and migratory stocks Herring in the Northeast Atlantic (Norwegian spring-spawning herring)

9.4.5 Advice September Widely distributed and migratory stocks Herring in the Northeast Atlantic (Norwegian spring-spawning herring) 9.4.5 Advice September 212 ECOREGION STOCK Widely distributed and migratory stocks Herring in the Northeast Atlantic (Norwegian spring-spawning herring) Advice for 213 ICES advises on the basis of the

More information

Sprat (Sprattus sprattus) in subdivisions (Baltic Sea)

Sprat (Sprattus sprattus) in subdivisions (Baltic Sea) ICES Advice on fishing opportunities, catch, and effort Baltic Sea Ecoregion Published 31 May 2016 8.3.18 Sprat (Sprattus sprattus) in subdivisions 22 32 (Baltic Sea) ICES stock advice ICES advises that

More information

Interactions in a multispecies age-structured assessment model for the Gulf of Alaska

Interactions in a multispecies age-structured assessment model for the Gulf of Alaska Interactions in a multispecies age-structured assessment model for the Gulf of Alaska Kray F. Van Kirk SFOS UAF Juneau, Alaska Terrance J. Quinn II SFOS UAF Juneau, Alaska Jeremy S. Collie GSO URI Narragansett,

More information

Abundance and trophic interactions in North Sea fishes

Abundance and trophic interactions in North Sea fishes Abundance and trophic interactions in North Sea fishes Georgi M. Daskalov and Steven Mackinson CEFAS, Lowestoft Laboratory, Lowestoft, Suffolk, NR33 OHT, UK Tel/fax: +44 () 152 524584, email: georgi.daskalov@cefas.co.uk

More information

Haddock (Melanogrammus aeglefinus) in divisions 7.b k (southern Celtic Seas and English Channel)

Haddock (Melanogrammus aeglefinus) in divisions 7.b k (southern Celtic Seas and English Channel) ICES Advice on fishing opportunities, catch, and effort Celtic Seas, Greater North Sea, and Oceanic Northeast Atlantic ecoregions Published 29 June 2018 https://doi.org/10.17895/ices.pub.4453 Haddock (Melanogrammus

More information

North East Atlantic Fisheries Baltic Sprat Whitepaper March 2011

North East Atlantic Fisheries Baltic Sprat Whitepaper March 2011 North East Atlantic Fisheries Baltic Sprat Whitepaper March 2011 1. Introduction Sprat is a clupeid fish found widely in the North East Atlantic area. Sprat is a relatively short-lived species. The stock

More information

A Combined Recruitment Index for Demersal Juvenile Cod in NAFO Divisions 3K and 3L

A Combined Recruitment Index for Demersal Juvenile Cod in NAFO Divisions 3K and 3L NAFO Sci. Coun. Studies, 29: 23 29 A Combined Recruitment Index for Demersal Juvenile Cod in NAFO Divisions 3K and 3L David C. Schneider Ocean Sciences Centre, Memorial University St. John's, Newfoundland,

More information

Assessment Summary Report Gulf of Mexico Red Snapper SEDAR 7

Assessment Summary Report Gulf of Mexico Red Snapper SEDAR 7 Assessment Summary Report Gulf of Mexico Red Snapper SEDAR 7 Stock Distribution: Red snapper are found throughout the Gulf of Mexico, the Caribbean Sea, and from the U.S. Atlantic Coast to northern South

More information

3.4.3 Advice June Barents Sea and Norwegian Sea Cod in Subareas I and II (Norwegian coastal waters cod)

3.4.3 Advice June Barents Sea and Norwegian Sea Cod in Subareas I and II (Norwegian coastal waters cod) 3.4.3 Advice June 2013 ECOREGION STOCK Barents Sea and Norwegian Sea Cod in Subareas I and II (Norwegian coastal waters cod) Advice for 2014 ICES advises on the basis of the Norwegian rebuilding plan,

More information

Report of the Working Group on Widely Distributed

Report of the Working Group on Widely Distributed ICES WGWIDE REPORT 2017 ICES ADVISORY COMMITTEE ICES CM 2017/ACOM:23 Report of the Working Group on Widely Distributed Stocks (WGWIDE) 30 August -5 September 2017 ICES Headquarters, Copenhagen, Denmark

More information

A. SOUTHERN NEW ENGLAND / MID-ATLANTIC (SNE/MA) WINTER FLOUNDER ASSESSMENT SUMMARY FOR 2011

A. SOUTHERN NEW ENGLAND / MID-ATLANTIC (SNE/MA) WINTER FLOUNDER ASSESSMENT SUMMARY FOR 2011 A. SOUTHERN NEW ENGLAND / MID-ATLANTIC (SNE/MA) WINTER FLOUNDER ASSESSMENT SUMMARY FOR 2011 State of Stock: In 2010 the SNE/MA winter flounder stock was overfished but overfishing was not occurring. The

More information

Trends in Scottish Fish Stocks 2018

Trends in Scottish Fish Stocks 2018 Port Arthur, Scalloway, Shetland, ZE1 UN, Scotland, UK Tel: +44 ()1595 772 Fax: +44 ()1595 7721 Email: info@nafc.uhi.ac.uk Web: www.nafc.ac.uk Trends in Scottish Fish Stocks 218 Ian R. Napier 14 th September

More information

Ecological interactions between forage fish, rorquals, and fisheries in Haida Gwaii

Ecological interactions between forage fish, rorquals, and fisheries in Haida Gwaii Ecological interactions between forage fish, rorquals, and fisheries in Haida Gwaii Szymon Surma MSc candidate UBC Fisheries Centre The problem rorquals: lunge-feeding baleen whales eat large quantities

More information

Spurdog (Squalus acanthias) in the Northeast Atlantic

Spurdog (Squalus acanthias) in the Northeast Atlantic ICES Advice on fishing opportunities, catch, and effort Northeast Atlantic Published 11 October 2016 9.3.17 Spurdog (Squalus acanthias) in the Northeast Atlantic ICES stock advice ICES advises that when

More information

Trends in Scottish Fish Stocks 2017

Trends in Scottish Fish Stocks 2017 Port Arthur, Scalloway, Shetland, ZE1 0UN, Scotland, UK Tel: +44 (0)1595 772000 Fax: +44 (0)1595 772001 Email: info@nafc.uhi.ac.uk Web: www.nafc.ac.uk Trends in Scottish Fish Stocks 2017 Ian R. Napier

More information

Advice May Herring in Subdivisions and 32 (excluding Gulf of Riga herring)

Advice May Herring in Subdivisions and 32 (excluding Gulf of Riga herring) 8.3.10 Advice May 2014 ECOREGION STOCK Baltic Sea Herring in Subdivisions 25 29 and 32 (excluding Gulf of Riga herring) Advice for 2015 ICES advises on the basis of the MSY approach that catches in 2015

More information

Ecological Reference Points for Atlantic Menhaden

Ecological Reference Points for Atlantic Menhaden Ecological Reference Points for Atlantic Menhaden Ken Hinman National Coalition for Marine Conservation June 2009 Among the ASMFC s tasks this year for conserving and managing Atlantic menhaden, according

More information

Advice October 2012

Advice October 2012 9.4.20 Advice October 2012 ECOREGION STOCK Widely distributed and migratory stocks Portuguese dogfish (Centroscymnus coelolepis) in the Northeast Atlantic Advice for 2013 and 2014 ICES advises on the basis

More information

Length and species-dependent diurnal variation of catch rates in the Norwegian Barents Sea bottom-trawl surveys

Length and species-dependent diurnal variation of catch rates in the Norwegian Barents Sea bottom-trawl surveys ICES Journal of Marine Science, 56: 284 291. 1999 Article No. jmsc.1999.0440, available online at http://www.idealibrary.com on Length and species-dependent diurnal variation of catch rates in the Norwegian

More information

Map Showing NAFO Management Units

Map Showing NAFO Management Units Map Showing NAFO Management Units Biology Are 6 species of seals in Atlantic Canadian waters, all of which occur in Newfoundland Two Arctic Species (Ringed, Bearded) Two temperate (Grey, Harbour) Two migratory

More information

Fishery Stock Status Fishing Mortality

Fishery Stock Status Fishing Mortality Northeast Groundfish Interim Findings: Ecological Indicators Version: February 2014 Fishery Stock Status Fishing Mortality This indicator measures the rate at which fishing removed fish each stock. This

More information

Herring (Clupea harengus) in subdivisions and 32 (central Baltic Sea, excluding Gulf of Riga)

Herring (Clupea harengus) in subdivisions and 32 (central Baltic Sea, excluding Gulf of Riga) ICES Advice on fishing opportunities, catch, and effort Baltic Sea Ecoregion Published 31 May 2016 8.3.14 Herring (Clupea harengus) in subdivisions 25 29 and 32 (central Baltic Sea, excluding Gulf of Riga)

More information

ASMFC Stock Assessment Overview: Red Drum

ASMFC Stock Assessment Overview: Red Drum Purpose The purpose of this document is to improve the understanding and transparency of the Commission s stock assessment process and results. It is the first of several that will be developed throughout

More information

Cod distribution and temperature in the North Sea

Cod distribution and temperature in the North Sea ICES mar. Sei. Symp., 198: 244-. 1994 Cod distribution and temperature in the North Sea Henk Heessen and Niels Daan Heessen, H. J. L., and Daan, N. 1994. Cod distribution and temperature in the North Sea.

More information

Mackerel (Scomber scombrus) in subareas 1 8 and 14, and in Division 9.a (the Northeast Atlantic and adjacent waters)

Mackerel (Scomber scombrus) in subareas 1 8 and 14, and in Division 9.a (the Northeast Atlantic and adjacent waters) ICES Advice on fishing opportunities, catch, and effort Ecoregions in the Northeast Atlantic and Arctic Ocean Published 29 September 2017 DOI: 10.17895/ices.pub.3023 Mackerel (Scomber scombrus) in subareas

More information

Comparison of EU and US Fishery management Systems Ernesto Penas Principal Adviser DG Mare

Comparison of EU and US Fishery management Systems Ernesto Penas Principal Adviser DG Mare Comparison of EU and US Fishery management Systems Ernesto Penas Principal Adviser DG Mare Stock and Fisheries Status Seminar Brussels, 26 September 2017 Why comparing the EU and the US? Need to put the

More information

HADDOCK ON THE SOUTHERN SCOTIAN SHELF AND IN THE BAY OF FUNDY (DIV. 4X/5Y)

HADDOCK ON THE SOUTHERN SCOTIAN SHELF AND IN THE BAY OF FUNDY (DIV. 4X/5Y) Canadian Science Advisory Secretariat Science Advisory Report 26/47 HADDOCK ON THE SOUTHERN SCOTIAN SHELF AND IN THE BAY OF FUNDY (DIV. 4X/5Y) Context Haddock (Melanogrammus aeglefinus) are found on both

More information

9.4.5 Advice October Widely Distributed and Migratory Stocks Herring in the Northeast Atlantic (Norwegian spring-spawning herring)

9.4.5 Advice October Widely Distributed and Migratory Stocks Herring in the Northeast Atlantic (Norwegian spring-spawning herring) 9.4.5 Advice October 21 ECOREGION STOCK Widely Distributed and Migratory Stocks Herring in the Northeast Atlantic (Norwegian spring-spawning herring) Advice for 211 Management Objective (s) Landings in

More information

BLACK SEA WHITING, MERLANGIUS MERLANGUS EUXINUS NORDMANN

BLACK SEA WHITING, MERLANGIUS MERLANGUS EUXINUS NORDMANN 82 BLACK SEA WHITING, MERLANGIUS MERLANGUS EUXINUS NORDMANN The taxonomic position of the Black Sea whiting like that of a great number of Black Sea fish species is rather complicated, and is still a controversial

More information

NOAA FISHERIES SERVICE. Habitat Assessment Prioritization

NOAA FISHERIES SERVICE. Habitat Assessment Prioritization NOAA FISHERIES SERVICE Habitat Assessment Prioritization What is a Habitat Assessment? A habitat assessment is both the process and products associated with consolidating, analyzing, and reporting the

More information

2016 ANNUAL FISH TRAWL SURVEY REPORT

2016 ANNUAL FISH TRAWL SURVEY REPORT 216 ANNUAL FISH TRAWL SURVEY REPORT The University of Rhode Island Graduate School of Oceanography The Graduate School of Oceanography (GSO) Fish trawl survey began weekly sampling two stations in Narragansett

More information

ASSESSMENT OF THE WEST COAST OF NEWFOUNDLAND (DIVISION 4R) HERRING STOCKS IN 2011

ASSESSMENT OF THE WEST COAST OF NEWFOUNDLAND (DIVISION 4R) HERRING STOCKS IN 2011 Canadian Science Advisory Secretariat Science Advisory Report 212/24 ASSESSMENT OF THE WEST COAST OF NEWFOUNDLAND (DIVISION 4R) HERRING STOCKS IN 211 Context Figure 1. Map of unit areas of NAFO Division

More information

10.3 Advice May 2014

10.3 Advice May 2014 1.3 Advice May 214 ECOREGION STOCK North Atlantic Atlantic salmon from North America Advice for 214 Because the NASCO Framework of Indicators of North American stocks for 213 (run in January 214) did not

More information

ICES Advice on fishing opportunities, catch, and effort Celtic Seas and Greater North Sea ecoregions Published 30 June 2016

ICES Advice on fishing opportunities, catch, and effort Celtic Seas and Greater North Sea ecoregions Published 30 June 2016 ICES Advice on fishing opportunities, catch, and effort Celtic Seas and Greater North Sea ecoregions Published 30 June 2016 5.3.57 Sea bass (Dicentrarchus labrax) in divisions 4.b c, 7.a, and 7.d h (central

More information

Fishing mortality in relation to highest yield. Fishing mortality in relation to agreed target

Fishing mortality in relation to highest yield. Fishing mortality in relation to agreed target 3.4 Stock summaries 3.4. Northeast Arctic cod State of the stock Spawning biomass in relation to precautionary limits Full reproductive capacity Fishing mortality in relation to precautionary limits/management

More information

Harvest Control Rules in a multispecies world: The Barents Sea and beyond. Daniel Howell IMR Bergen

Harvest Control Rules in a multispecies world: The Barents Sea and beyond. Daniel Howell IMR Bergen Harvest Control Rules in a multispecies world: The Barents Sea and beyond Daniel Howell IMR Bergen Single species HCR 1997 2012 Multispecies HCRs Will give examples of Existing multispecies HCRs (explicitly

More information

3.3.2 Cod (Gadus morhua) in subareas 1 and 2 (Northeast Arctic)

3.3.2 Cod (Gadus morhua) in subareas 1 and 2 (Northeast Arctic) ICES Advice on fishing opportunities, catch, and effort Barents Sea and Norwegian Sea Ecoregions Published 10 June 2016 3.3.2 Cod (Gadus morhua) in subareas 1 and 2 (Northeast Arctic) ICES stock advice

More information

EU request to ICES on in-year advice on haddock (Melanogrammus aeglefinus) in Division 7.a (Irish Sea)

EU request to ICES on in-year advice on haddock (Melanogrammus aeglefinus) in Division 7.a (Irish Sea) ICES Special Request Advice Celtic Seas Ecoregion Published 1 March 2017 EU request to ICES on in-year advice on haddock (Melanogrammus aeglefinus) in Division 7.a (Irish Sea) Advice summary Please note:

More information

Recruitment of cod and haddock in the North Atlantic: a comparative analysis

Recruitment of cod and haddock in the North Atlantic: a comparative analysis ICES Journal of Marine Science, 5: 952 961. 21 doi:1.16/jmsc.21.11, available online at http://www.idealibrary.com on Recruitment of cod and haddock in the North Atlantic: a comparative analysis Michael

More information

Northwest Atlantic Harp Seals

Northwest Atlantic Harp Seals Fisheries Pêches and Oceans et Océans DFO Science Stock Status Report E1-01(2000) Background The Harp seal is an abundant, medium sized seal which migrates annually between Arctic and sub-arctic regions

More information

Year Avg. TAC Can Others Totals

Year Avg. TAC Can Others Totals SKATE IN DIVISIONS 3L, 3N, 3O AND SUBDIVISION 3Ps Background There are 8 to 1 species of skate in the waters around Newfoundland. Of these, thorny skate (Raja radiata) is by far the most common, comprising

More information

ICES advises that when the MSY approach is applied, catches in 2019 should be no more than tonnes.

ICES advises that when the MSY approach is applied, catches in 2019 should be no more than tonnes. Advice on fishing opportunities, catch, and effort Celtic Seas, Greater North Sea, and Oceanic Northeast Atlantic ecoregions Published 29 June 2018 Version 2: 4 September 2018 https://doi.org/10.17895/ices.pub.4487

More information

ASSESSMENT OF THE WEST COAST OF NEWFOUNDLAND (DIVISION 4R) HERRING STOCKS IN 2013

ASSESSMENT OF THE WEST COAST OF NEWFOUNDLAND (DIVISION 4R) HERRING STOCKS IN 2013 Canadian Science Advisory Secretariat Quebec Region Science Advisory Report 214/56 ASSESSMENT OF THE WEST COAST OF NEWFOUNDLAND (DIVISION 4R) HERRING STOCKS IN 213 Quebec 4Ra 4S 4R 4Rc 4Rb U.S.A. (Maine)

More information

Effects of juvenile nursery on geographic spawning distribution in Norwegian spring-spawning herring (Clupea harengus L.)

Effects of juvenile nursery on geographic spawning distribution in Norwegian spring-spawning herring (Clupea harengus L.) ICES Journal of Marine Science, 55: 987 996. 1998 Article No. jm980371 Effects of juvenile nursery on geographic spawning distribution in Norwegian spring-spawning herring (Clupea harengus L.) J. C. Holst

More information

Technical Briefing. Northern Cod (NAFO Div. 2J3KL) Newfoundland & Labrador March 23, 2018

Technical Briefing. Northern Cod (NAFO Div. 2J3KL) Newfoundland & Labrador March 23, 2018 Technical Briefing Northern Cod (NAFO Div. 2J3KL) Newfoundland & Labrador March 23, 2018 Outline Purpose of this briefing Science advice & the precautionary approach Summary of stock status How we estimate

More information

ICES advice on fishing opportunities

ICES advice on fishing opportunities ICES Advice on fishing opportunities, catch, and effort Northeast Atlantic and Arctic Ocean Published 22 October 2018 Version 2: 25 October 2018 https://doi.org/10.17895/ices.pub.4568 Herring (Clupea harengus)

More information

Effective Collaboration Between Scientists, Managers and Policy Makers

Effective Collaboration Between Scientists, Managers and Policy Makers 2009/FWG/WKSP/007 Effective Collaboration Between Scientists, Managers and Policy Makers Submitted by: World Bank Workshop on Implementing Ecosystem Approaches to Fisheries in the Context of the Broader

More information

Growth: Humans & Surf Clams

Growth: Humans & Surf Clams Growth: Humans & Surf Clams East Coast MARE Materials For the leader: Projector Whiteboard to project data graph onto White paper & markers For the activity: Copy of data table Copy of map Class graphing

More information

Advice June 2014

Advice June 2014 5.3.23 Advice June 2014 ECOREGION STOCK Celtic Sea and West of Scotland Plaice in Division VIIa (Irish Sea) Advice for 2015 Based on ICES approach to data-limited stocks, ICES advises that catches should

More information

Predator diet as an indicator of comb jellyfish (Ctenophora) abundance dynamics in the Barents Sea

Predator diet as an indicator of comb jellyfish (Ctenophora) abundance dynamics in the Barents Sea Predator diet as an indicator of comb jellyfish (Ctenophora) abundance dynamics in the Barents Sea Eriksen, E. 1, Dolgov, A. 2, Bogstad, B. 1 1 Institute of Marine Research (IMR), Norway 2 Polar Research

More information

Advice June Capelin in Subareas V and XIV and Division IIa west of 5 W (Iceland East Greenland Jan Mayen area).

Advice June Capelin in Subareas V and XIV and Division IIa west of 5 W (Iceland East Greenland Jan Mayen area). 2.4.12 Advice June 2012 ECOREGION Iceland and East Greenland STOCK Capelin in Subareas V and XIV and Division IIa west of 5 W (Iceland East Greenland Jan Mayen area) Advice for 2013 ICES advises on the

More information

Stock characteristics, fisheries and management of Greenland halibut (Reinhardtius hippoglossoides (Walbaum)) in the Northeast Arctic

Stock characteristics, fisheries and management of Greenland halibut (Reinhardtius hippoglossoides (Walbaum)) in the Northeast Arctic The 10 th Norwegian Russian Symposium Stock characteristics, fisheries and management of Greenland halibut (Reinhardtius hippoglossoides (Walbaum)) in the Northeast Arctic by Kjell H. Nedreaas (IMR) &

More information

Age-Structured Multispecies Model of the Georges Bank Fish Community

Age-Structured Multispecies Model of the Georges Bank Fish Community University of Rhode Island DigitalCommons@URI Open Access Dissertations 212 AgeStructured Multispecies Model of the Georges Bank Fish Community Kiersten Lynn Curti University of Rhode Island, kiersten.curti@gmail.com

More information

STOCK ASSESSMENT OF ALBACORE TUNA IN THE NORTH PACIFIC OCEAN IN 2011

STOCK ASSESSMENT OF ALBACORE TUNA IN THE NORTH PACIFIC OCEAN IN 2011 STOCK ASSESSMENT OF ALBACORE TUNA IN THE NORTH PACIFIC OCEAN IN 2011 Report of the ISC-Albacore Working Group Stock Assessment Workshop 3rd Science Advisory Committee Meeting Inter-American Tropical Tuna

More information

Science-based management of fish stocks and long-term sustainability

Science-based management of fish stocks and long-term sustainability Science-based management of fish stocks and long-term sustainability Jóhann Sigurjónsson, Director General Marine Research Institute, Iceland Seminar on the Role of Science and Technology in Responsible

More information

Pelagic fishery for Sebastes mentella in the Irminger Sea

Pelagic fishery for Sebastes mentella in the Irminger Sea 3.2.6.d Pelagic fishery for Sebastes mentella in the Irminger Sea The stock structure of deep-sea redfish S. mentella in Sub-area XII, Division Va and Sub-area XIV and NAFO Div. 1F remains generally uncertain.

More information

Feeding ecology of herring and mackerel in Icelandic waters

Feeding ecology of herring and mackerel in Icelandic waters Workshop on the Future Potential of Zooplankton and Mesopelagic Species October 1st - 2nd 2018, Reykjavík, Iceland Feeding ecology of herring and mackerel in Icelandic waters and other aspects of relevance

More information

Forage indicators and consumption profiles for Chesapeake Bay fishes

Forage indicators and consumption profiles for Chesapeake Bay fishes Forage indicators and consumption profiles for Chesapeake Bay fishes Andre Buchheister Ed Houde Carlos Lozano Presentation to Fisheries GIT Dec 14, 2015 Outline Background & Objectives Result Highlights

More information

Impact of climate variability and change on winter survival of Bristol Bay sockeye salmon

Impact of climate variability and change on winter survival of Bristol Bay sockeye salmon Impact of climate variability and change on winter survival of Bristol Bay sockeye salmon Alaska Fisheries Science Center Ed Farley (NOAA), Greg Ruggerone (NRC) Phil Mundy (NOAA), Ellen Yasumiishi (NOAA)

More information

Summary of discussion

Summary of discussion Tweedsmuir Caribou Modelling Project: Caribou Population Ecology Meeting Notes held March 5, 2008 Participants: Mark Williams, Debbie Cichowski, Don Morgan, Doug Steventon, Dave Daust Purpose: The purpose

More information

3.4 Stock Summaries Advice June 2011

3.4 Stock Summaries Advice June 2011 3.4 Stock Summaries 3.4.1 Advice June 211 ECOREGION STOCK Barents Sea and Norwegian Sea Cod in Subareas I and II (Northeast Arctic cod) Advice for 212 ICES advises on the basis of the Joint Russian Norwegian

More information

2.3.1 Advice May Capelin in Subareas V and XIV and Division IIa west of 5 W (Iceland East Greenland Jan Mayen area).

2.3.1 Advice May Capelin in Subareas V and XIV and Division IIa west of 5 W (Iceland East Greenland Jan Mayen area). 2.3.1 Advice May 2014 ECOREGION Iceland and East Greenland STOCK Capelin in Subareas V and XIV and Division IIa west of 5 W (Iceland East Greenland Jan Mayen area) Advice for 2014/2015 ICES advises on

More information

J. G. Pope and C. T. Macer. Introduction

J. G. Pope and C. T. Macer. Introduction ICES Journal of Marine Science, 53: 1157 1169. 1996 An evaluation of the stock structure of North Sea cod, haddock, and whiting since 192, together with a consideration of the impacts of fisheries and

More information

Why has the cod stock recovered in the North Sea?

Why has the cod stock recovered in the North Sea? Why has the cod stock recovered in the North Sea? Summary The expansion of European fisheries during the 1970s and 1980s resulted in high fishing pressure on stocks of cod, haddock, whiting and saithe

More information

Bycatch accounting and management in the Ross Sea toothfish fishery

Bycatch accounting and management in the Ross Sea toothfish fishery Bycatch accounting and management in the Ross Sea toothfish fishery Steve Parker Sophie Mormede Stuart Hanchet New Zealand Ross Sea Region 4.5 million km 2 of ocean High seas fishery Managed by CCAMLR

More information

Plaice (Pleuronectes platessa) in Division 7.e (western English Channel)

Plaice (Pleuronectes platessa) in Division 7.e (western English Channel) Celtic Seas and Greater North Sea ecoregions Published 30 June 2016 Version 2: 15 May 2017 5.3.51 Plaice (Pleuronectes platessa) in Division 7.e (western English Channel) ICES stock advice ICES advises

More information

Advice June, revised September Herring in Division IIIa and Subdivisions (Western Baltic spring spawners)

Advice June, revised September Herring in Division IIIa and Subdivisions (Western Baltic spring spawners) 6.4.15 Advice June, revised September 21 ECOREGION STOCK North Sea and Baltic Herring in Division IIIa and Subdivisions 22 24 (Western Baltic spring spawners) Advice for 211 Management Objective (s) Catches

More information

Groundfish Operational Assessments 2017 Overview

Groundfish Operational Assessments 2017 Overview Groundfish Operational Assessments 2017 Overview Northeast Fisheries Science Center http://www.nefsc.noaa.gov/groundfish/operational-assessments-2017/ Summary Presentation Operational Assessment Process

More information

Golden redfish (Sebastes norvegicus) in subareas 5, 6, 12, and 14 (Iceland and Faroes grounds, West of Scotland, North of Azores, East of Greenland)

Golden redfish (Sebastes norvegicus) in subareas 5, 6, 12, and 14 (Iceland and Faroes grounds, West of Scotland, North of Azores, East of Greenland) ICES Advice on fishing opportunities, catch, and effort Arctic Ocean, Celtic Seas, Faroes, Greenland Sea, Icelandic Waters, Published 13 June 2017 Norwegian Sea, and Oceanic Northeast Atlantic Ecoregions

More information

Determination of the von Bertalanffy Growth Equation for the Southern New England-Middle Atlantic~ Bank and Gulf of Maine stocks of Silver Hake by

Determination of the von Bertalanffy Growth Equation for the Southern New England-Middle Atlantic~ Bank and Gulf of Maine stocks of Silver Hake by Determination of the von Bertalanffy Growth Equation for the Southern New England-Middle Atlantic~ Bank and Gulf of Maine stocks of Silver Hake by F. P. Almeida Georges 3 National Marine Fisheries Service

More information

Introduction to population dynamics and stock assessments

Introduction to population dynamics and stock assessments South East Fisheries Science Center Introduction to population dynamics and stock assessments Adyan Rios MREP USVI Tuesday, May 23, 2017 Introduction to population dynamics and stock assessments Part 1

More information

CATCH-AT-SIZE AND AGE ANALYSIS FOR ATLANTIC SWORDFISH

CATCH-AT-SIZE AND AGE ANALYSIS FOR ATLANTIC SWORDFISH SCRS/2017/136 Collect. Vol. Sci. Pap. ICCAT, 74(3): 1258-1274 (2017) CATCH-AT-SIZE AND AGE ANALYSIS FOR ATLANTIC SWORDFISH Alex Hanke 1, Laurie Kell, Rui Coelho and Mauricio Ortiz SUMMARY Analyses of the

More information

2000 REVIEW OF THE ATLANTIC STATES MARINE FISHERIES COMMISSION FISHERY MANAGEMENT PLAN FOR BLUEFISH (Pomatomus saltatrix)

2000 REVIEW OF THE ATLANTIC STATES MARINE FISHERIES COMMISSION FISHERY MANAGEMENT PLAN FOR BLUEFISH (Pomatomus saltatrix) 2000 REVIEW OF THE ATLANTIC STATES MARINE FISHERIES COMMISSION FISHERY MANAGEMENT PLAN FOR BLUEFISH (Pomatomus saltatrix) Prepared by: Robert Beal (ASMFC) Bluefish Plan Review Team Elliot Atstupenas (USFWS)

More information

Multispecies Models Relevant to Management of Living Resources

Multispecies Models Relevant to Management of Living Resources ICES mar. Sei. Symp., 193: 1. 1991 Multispecies Models Relevant to Management of Living Resources Preface Scientific quality is undoubtedly the most important aspect of a contribution to a symposium, but

More information

I. What is a Fishery? II. What is Fisheries Management? III. What is Fisheries Science? I. Brief history of the evolution of fisheries science.

I. What is a Fishery? II. What is Fisheries Management? III. What is Fisheries Science? I. Brief history of the evolution of fisheries science. I. What is a Fishery? II. What is Fisheries Management? III. What is Fisheries Science? I. Brief history of the evolution of fisheries science. Archeological records Spears: 90,000 BP Nets: 40,000 BP

More information

3.3.2 Advice June 2014

3.3.2 Advice June 2014 3.3.2 Advice June 2014 ECOREGION STOCK Barents Sea and Norwegian Sea Cod in Subareas I and II (Northeast Arctic cod) Advice for 2015 ICES advises on the basis of the Joint Russian Norwegian Fisheries Commission

More information

Hatcheries: Role in Restoration and Enhancement of Salmon Populations

Hatcheries: Role in Restoration and Enhancement of Salmon Populations Hatcheries: Role in Restoration and Enhancement of Salmon Populations Hatcheries play a large role in the management, ecology, and evolution of Pacific salmon. Why were/are they built? What are the assumptions

More information

Long-term trends in the biomass of commercial fish in the North Sea fishing impacts, predator-prey interactions and temperature change

Long-term trends in the biomass of commercial fish in the North Sea fishing impacts, predator-prey interactions and temperature change Long-term trends in the biomass of commercial fish in the North Sea fishing impacts, predator-prey interactions and temperature change S10 Forecasting climate change impacts on fish populations and fisheries

More information

Developments in managing small pelagic fisheries

Developments in managing small pelagic fisheries Developments in managing small pelagic fisheries Prof Ray Hilborn Your essential event for networking and information Impacts of fishing forage fish on their predators: Progress report October 2017 Ray

More information

Essential Fish Habitat Description Atlantic cod (Gadus morhua)

Essential Fish Habitat Description Atlantic cod (Gadus morhua) Description Atlantic cod (Gadus morhua) In its Report to Congress: Status of the Fisheries of the United States (September 1997), NMFS determined the Gulf of Maine stock of cod is considered overfished,

More information

A REVIEW AND EVALUATION OF NATURAL MORTALITY FOR THE ASSESSMENT AND MANAGEMENT OF YELLOWFIN TUNA IN THE EASTERN PACIFIC OCEAN

A REVIEW AND EVALUATION OF NATURAL MORTALITY FOR THE ASSESSMENT AND MANAGEMENT OF YELLOWFIN TUNA IN THE EASTERN PACIFIC OCEAN A REVIEW AND EVALUATION OF NATURAL MORTALITY FOR THE ASSESSMENT AND MANAGEMENT OF YELLOWFIN TUNA IN THE EASTERN PACIFIC OCEAN Mark N. Maunder and Alex Aires-da-Silva Outline YFT history Methods to estimate

More information

Preliminary results of SEPODYM application to albacore. in the Pacific Ocean. Patrick Lehodey

Preliminary results of SEPODYM application to albacore. in the Pacific Ocean. Patrick Lehodey SCTB15 Working Paper ALB-6 Preliminary results of SEPODYM application to albacore in the Pacific Ocean Patrick Lehodey Oceanic Fisheries Programme Secretariat of the Pacific Community Noumea, New Caledonia

More information

Fifty years ago, a single cod was large enough to feed a family of four or five. Today it is barely enough for one

Fifty years ago, a single cod was large enough to feed a family of four or five. Today it is barely enough for one Fifty years ago, a single cod was large enough to feed a family of four or five. Today it is barely enough for one Lord Perry of Walton, UK House of Lords (1997) (as cited in Stergiou 2002) Minimum size

More information

Atlantic croaker, Micropogonias undulatus (Linnaeus, 1766)

Atlantic croaker, Micropogonias undulatus (Linnaeus, 1766) Atlantic croaker, Micropogonias undulatus (Linnaeus, 1766) Although Atlantic croaker occur throughout much of Florida, they are seldom found south of Tampa Bay on the gulf coast or south of the Indian

More information

Sustainable Seas - Marine Fisheries Fisheries and Fishing

Sustainable Seas - Marine Fisheries Fisheries and Fishing Sustainable Seas - Marine Fisheries Fisheries and Fishing Maximum Sustainable Yield and Fishery Management Mariculture Reading: Ch 9: 9.12 Ch 14: 14.7-14.9 Ch 17: 17.1, 17.17-17.21,17.25 Graphic: Chub

More information

Top down modeling and bottom up dynamics: Linking fisheries-based multispecies models with climate hypotheses in the Northern California Current

Top down modeling and bottom up dynamics: Linking fisheries-based multispecies models with climate hypotheses in the Northern California Current Top down modeling and bottom up dynamics: Linking fisheries-based multispecies models with climate hypotheses in the Northern California Current John Field 1, Robert Francis 2, and Kerim Aydin 3 1 Santa

More information

Advice June 2012

Advice June 2012 2.4.1 Advice June 212 ECOREGION STOCK Iceland and East Greenland Beaked redfish (Sebastes mentella) in Subareas V, XII, and XIV and NAFO Subareas 1+2 (Deep pelagic stock > 5 m) Advice for 213 The advice

More information

Whiting (Merlangius merlangus) in divisions 7.b c and 7.e k (southern Celtic Seas and western English Channel)

Whiting (Merlangius merlangus) in divisions 7.b c and 7.e k (southern Celtic Seas and western English Channel) ICES Advice on fishing opportunities, catch, and effort Celtic Seas, Greater North Sea, and Oceanic Northeast Atlantic ecoregions Published 30 June 2017 Version 2: 07 July 2017 DOI: 10.17895/ices.pub.3269

More information

Atlantic Striped Bass Draft Addendum V. Atlantic Striped Bass Board May 9, 2017

Atlantic Striped Bass Draft Addendum V. Atlantic Striped Bass Board May 9, 2017 Atlantic Striped Bass Draft Addendum V Atlantic Striped Bass Board May 9, 2017 Timeline May 2017: Consider Approval of Draft Addendum V for Public Comment May July 2017: Public Comment period August 2017:

More information

Managing Lower Trophic Level Species in the Mid-Atlantic Region

Managing Lower Trophic Level Species in the Mid-Atlantic Region Managing Lower Trophic Level Species in the Mid-Atlantic Region Forage Fish Workshop Mid-Atlantic Fishery Management Council Raleigh, orth Carolina 11 April 2013 E. D. Houde orthwest Atlantic Coastal and

More information

SOCIETAL GOALS TO DETERMINE ECOSYSTEM HEALTH: A FISHERIES CASE STUDY IN GALVESTON BAY SYSTEM, TEXAS

SOCIETAL GOALS TO DETERMINE ECOSYSTEM HEALTH: A FISHERIES CASE STUDY IN GALVESTON BAY SYSTEM, TEXAS SOCIETAL GOALS TO DETERMINE ECOSYSTEM HEALTH: A FISHERIES CASE STUDY IN GALVESTON BAY SYSTEM, TEXAS Anthony S. Pait, NOAA, National Centers for Coastal Ocean Science Thomas P. O Connor, David R. Whitall,

More information

Recruitment processes of jack mackerel (Trachurus

Recruitment processes of jack mackerel (Trachurus Recruitment processes of jack mackerel (Trachurus japonicus) ) in the East China Sea (ECS) in relation to environmental conditions Chiyuki Sassa 1, Youichi Tsukamoto 1, Yoshinobu Konishi 1* Songguang Xie

More information

Surf Clams: Latitude & Growth

Surf Clams: Latitude & Growth Surf Clams: Latitude & Growth East Coast MARE Materials For the leader: Projector Whiteboard to project data graph onto For the activity: Copy of data table Copy of map Computer program to graph in or

More information

Fish larvae atlas of the NE Atlantic

Fish larvae atlas of the NE Atlantic Fish larvae atlas of the NE Atlantic Results from the Continuous Plankton Recorder survey 1948-2005 Sir Alister Hardy Foundation for Ocean Science Monitoring the health of the oceans since 1931 Citation:

More information

North Labrador Arctic Charr

North Labrador Arctic Charr Fisheries and Oceans Pêches et Océans Canada Canada DFO Science Newfoundland Region Stock Status Report D2-07(2001) substantive in some years. The north Labrador area is composed of various stock complexes

More information

Sandeel (Ammodytes spp.) in Divisions IIIa, IVa, and IVb, SA 3 (Skagerrak and Kattegat, North and Central North Sea)

Sandeel (Ammodytes spp.) in Divisions IIIa, IVa, and IVb, SA 3 (Skagerrak and Kattegat, North and Central North Sea) ICES Advice on fishing opportunities, catch and effort Greater North Sea Ecoregion Published 23 February 2015 6.3.39 Sandeel (Ammodytes spp.) in Divisions IIIa, IVa, and IVb, SA 3 (Skagerrak and Kattegat,

More information

Below you will find 4 different student projects that combine fisheries science and marine ecology in various ways

Below you will find 4 different student projects that combine fisheries science and marine ecology in various ways Below you will find 4 different student projects that combine fisheries science and marine ecology in various ways PROJECT #1: How much forage fish do fisheries need to leave in the sea to support ecosystem

More information

Common Resource Problem-Fishing

Common Resource Problem-Fishing Common Resource Problem-Fishing Many examples exist of how overfishing has brought about the collapse of fisheries such as Canada s northern cod fishery, innumerable salmon fisheries, many shell- fish

More information

Lect 19 - Populations - Chapter 23. Different Levels of Ecological Organization. Populations

Lect 19 - Populations - Chapter 23. Different Levels of Ecological Organization. Populations Lect 19 - - Chapter 23 Different Levels of Ecological Organization Individuals Assemblage Community Ecosystem - all individuals of a particular species in a given area - but need to recognize genetic structure

More information

ASMFC Stock Assessment Overview: Atlantic Menhaden

ASMFC Stock Assessment Overview: Atlantic Menhaden Introduction This document presents a summary of the 217 Stock Assessment Update for Atlantic menhaden. The assessment is an update to the 215 Benchmark Stock Assessment that was peer reviewed by an independent

More information