Description of Panulirus brunneiflagellum new species with notes on its biology, evolution, and fisheries

Save this PDF as:
 WORD  PNG  TXT  JPG

Size: px
Start display at page:

Download "Description of Panulirus brunneiflagellum new species with notes on its biology, evolution, and fisheries"

Transcription

1 New Sekiguchi Zealand & George Description Journal of Marine and of Freshwater Panulirus brunneiflagellum Research, 2005, new Vol. species 39: /05/ The Royal Society of New Zealand Description of Panulirus brunneiflagellum new species with notes on its biology, evolution, and fisheries HIDEO SEKIGUCHI Faculty of Bioresources Mie University 1515 Kamihama-cho, Tsu Mie , Japan RAY W. GEORGE Centre of Excellence in Natural Resource Management University of Western Australia Albany WA 6330, Australia Abstract The new species is distinguished from its nearest relatives by the colour and patterning of the anntennual rami. Panulirus brunneiflagellum is morphologically close to P. japonicus and is restricted to the subtropical Ogasawara Islands in the west Pacific. It probably became isolated from the P. japonicus stock with the development of the unique Ogasawara oceanic circulatory system as a result of the deflection of the Kuroshio subgyre as the Iwo-Jima Ridge uplifted. Over 90% of the annual production of c. 5 t is frozen and transported to the Tokyo market, Japan. Keywords Panulirus brunneiflagellum; evolution; new species Panulirus; fishery M04038; Online publication date 8 June 2005 Received 10 February 2004; accepted 2 June 2004 INTRODUCTION A review of the Panulirus japonicus species complex was published by George & Holthuis (1965) in which keys, descriptions, and figures distinguished the geographically restricted species P. japonicus, P. marginatus, P. pascuensis, P. cygnus, and the widespread, and subsequently proven to be confusing, P. longipes (see George 1997 and Chan & Ng 2001, for details of taxa related to P. longipes, and debate over their nomenclature). George & Holthuis (1965) recognised two subspecies of P. longipes, the nominate P. longipes longipes with spotted legs of the Indian Ocean and a striped-legged form (called by them P. longipes femoristriga) inhabiting the western and central Pacific Ocean. George (1972), Sekiguchi (1991), and Chan & Chu (1996) recognised that P. l. femoristriga comprised more than one taxon and, as part of a study of evolution of Panulirus, George (1997) concluded that three taxa, all possessing striped legs, have been identified commonly as P. l. femoristriga. The key distinguishing features allowing separation of these three taxa with striped legs is the coloration and patterning of the antennular flagella. Two of the striped-legged taxa were described and figured by Sekiguchi (1991) who used the local Japanese names at Ogasawara Islands of Aka-ebi (brown whisker) and Shirahige-ebi (white-banded whisker) to distinguish them. Panulirus femoristriga sensus strictu is recognised by the distinctive one white and one darl ramus of each antennule and is known as Shironuke (plain white whisker) by Japanese fishers. According to Chan & Ng (2001) who examined the nomenclature of P. l. femoristriga, P. bispinosus, and P. albiflagellum, Shirahige-ebi is identical to P. l. bispinosus and Shironuke to P. femoristriga sensus strictu (synonym P. albiflagellum). Thus, we believe a new Panulirus name should be given to Aka-ebi. In December 1998, the two authors visited Chichi-Jima in the Ogasawara Islands to gather first hand data. This paper formally describes Aka-ebi as the new taxon and provides notes on its speciation,

2 564 New Zealand Journal of Marine and Freshwater Research, 2005, Vol. 39 habitat, and fishing importance. Measurements are of median carapace length from between the rostral horns and the posterior margin of the carapace. SYSTEMATICS Panulirus brunneiflagellum sp. nov. (Fig. 1, 2A C, 3A) Aka-ebi: Sekiguchi 1991; Chan & Chu 1996 Panulirus aka : George 1997 Material examined Holotype Ogasawara, Mukojima Chain, in trap, November 1997, male, mm (Western Australian Museum, WAM C23504). Paratypes Ogasawa, Mukojima Chain, in trap, November 1997, 1 male, 90.5 mm (WAM C23507), 2 females, 98.4 and mm, with spermatophores (WAM C23505 and C23506). Chichi-Jima, in commercial holding tanks 12 December 1998, 3 females each with spermatophore and 4 males, mm. Description of male holotype Antennular plate with two principal anterior spines and two posteriorly diverging rows of three spinules behind the principal pair of spines; ventral surfaces of distal two segments of antennal peduncle without spinules, distal segment with central pair of major spines, second segment with central pair and basal pair of major spines (Fig. 3A); exopod of third maxilliped with multi-articulate flagellum reaching half length of merus of endopod (Fig. 2C); exopod of second maxilliped with multi-articulate flagellum reaching well beyond endopod (Fig. 2B); posterior margin of thoracic sternum with distinct pair of posteriorly directed teeth; male reproductive opening with curved, raised chitinous ridge without setal tuft; outer margin of triangular plate (plate D) on first abdominal somite with very slight indentation; anterior margin of second abdominal pleura nearly straight with one very minute spinule (Fig. 2A); transverse grooves of second to fourth abdominal somites uninterrupted, very slightly medianly indented; lateral end of second transverse groove does not join pleural groove, lateral end of transverse groove of third somite on right side only, does not join pleural groove, remainder join corresponding grooves; all grooves covered by fringe of setae; apart from setae along posterior margin of the somites and those along the transverse grooves, abdominal terga entirely naked; male pleopods without endopods. Colour Antennule peduncle blackish-purple with white spot at distal articulations and lateral white irregular line; antennular flagella uniformly brownish without crossbands; antennular plate blackish-purple with a large central triangular white spot; antennal peduncle blackish-purple with pale spots and orange tips to dark spines; antennal flagella uniformly dark gray; supraorbital horns blackish-purple with orange tips and large baso-ventral white spot; dorsal carapace blackish-purple anteriorly, brownish centrally and posteriorly, laterally more orange with pale spots, many spines white and orange tipped; continuous mid lateral stripe on carapace, ventral stripe broken and spotted; posterior carapace margin blackishpurple with 13 distinct white spots; cervical groove with fine white spots and stripes; dorsal abdomen blackish-purple with many white and orange dots and laterally, small white spots and larger spots especially on plate D, behind each somite articulation and on abdominal pleura; pereiopods purplish, merus longitudinally streaked with five irregular, narrow white lines; carpus and propodus with dorsal orange stripe, articulations orange; pleopods bordered white with basal spot; telson bordered yellowish. Variations in paratypes In the series examined, the rows of diverging spinules behind the principal pair on the antennular plate ranged from 1 to 5; the exopod of the third maxilliped of the smallest male and in two females reaches only one third of endopod and in another female the exopod reaches only the base of the merus of the endopod; four of the five males have the distinct pair of spines on the thoracic sternum like the holotype, but in one male there is only one very small spinule on the right side; in two females (one with and one without spermatophore) it is reduced to two tiny spinules, and in the remaining mature three females (with spermatophores) the teeth are absent. For two of the four males and for three of the five females in the paratypical series, the transverse groove of the second abdominal somite does not join the pleural groove, for the other two males and one female this transverse groove does join the pleural groove (as is the situation for a specimen described by Sekiguchi (1991) from Hachijo Island), the joining of the transverse groove and pleural groove in the remaining female is very

3 Sekiguchi & George Description of Panulirus brunneiflagellum new species 565 Fig. 1 Panulirus brunneiflagellum. Female with spermatophore from Ogasawara Islands, 120 mm carapace length. A, dorsal view of abdomen; B, lateral view of abdomen; C, lateral view of carapace; D, dorsal view of carapace; E, antennular flagella (after Sekiguchi 1991).

4 566 New Zealand Journal of Marine and Freshwater Research, 2005, Vol. 39 Fig. 2 A C, Panulirus brunneiflagellum. D F, Panulirus longipes bispinosus. A and D, lateral view of abdomen; B and E, second maxilliped; C and F, third maxilliped. Short hairs along transverse and pleural grooves of abdomen and spines of second and third maxillipeds are omitted. Scale = 10 mm for A and D, and 5 mm for B, C, E, and F (after Sekiguchi 1991). indistinct. Thus six of the 10 have the condition as described for the holotype. For the females the small white spots and orange dots on the dorsal and lateral surfaces of the abdomen are larger and provide a more even cover than for the males. Fig. 3 Ventral view of distal two segments of antennal peduncle. A, Panulirus brunneiflagellum holotype male, WAM23504; B, Shirahige-ebi (Panulirus longipes bispinosus). Scale = 10 mm. Distribution Panulirus brunneiflagellum is the dominant species in the southern islands of the Izu Ridge at the Ogasawara Islands (Bonin Is.). These islands are c km south of Tokyo, Japan and 1500 km north of Saipan, Mariana Islands. It has also been recorded from Hachijo Island at the northern end of the Izu Ridge, c. 300 km south of Tokyo (Sekiguchi 1991). Panulirus l. bispinosus (referred to in Sekiguchi 1991 as P. l. femoristriga), the other less abundant taxon from Ogasawara Is., has been recorded from

5 Sekiguchi & George Description of Panulirus brunneiflagellum new species 567 two separate subtropical regions of the west Pacific. It occurs in the north-west Pacific (10 35 N) at the near-asian islands of Japan, Ryukyu Archipelago, Taiwan, and Philippines and the offshore islands of Ogasawara and Hachijo (Sekiguchi 1991). Another population occurs in the south-west Pacific (15 32 S) on the moderate sized islands of Cook, Fiji, Vanuatu, and New Caledonia as well as the mid east coast of Australia (George & Holthuis 1965; George 1972; Sekiguchi 1991; Chan & Chu 1996). Habitat Panulirus brunneiflagellum lives in open caves, down steep volcanic slopes from surface to 90 m in clear oceanic waters. It usually occupies the entrance of the caves, occasionally sharing the cave with P. l. bispinosus, however P. l. bispinosus usually shelters singly or in small groups in the deep recesses at the back of the cave (Nishikiori & Sekiguchi 2000). In the spawning season the two species form separate groups in separate caves. Tides at the Ogasawara are semidiurnal with a spring range of 1.5 m. Currents can run strongly between islands but swells are minimal. Surface water temperatures range from 19 to 29 C. In contrast, habitat records of P. l. bispinosus are as follows: Coral reef (Fiji and Heron Island), speared or netted in shallow water (Japan, Philippines, and New Caledonia) and taken in lobster traps to 130 m (Vanuatu and Australia) (George & Holthuis 1965; George 1972; Juinio et al. 1991). In the Ogasawara Islands, P. l. bispinosus usually shelters singly or in small groups in the deep recesses at the back of the cave (Nishikiori & Sekiguchi 2000). Fishery Nishikiori & Sekiguchi (2001) report that Japanese fishers have taken lobsters ever since the Ogasawara Islands were occupied more than 150 years ago. They are usually caught between 10 and 40 m depths in steel-framed rectangular traps covered with plastic mesh. Each trap measures c. 1 m 0.5 m 0.5 m and has a funnel entrance at each end. The buoyed traps are set singly each day on the volcanic rock slopes, baited with fresh fish. Twenty fishing boats work out from Chichi-Jima Island and two to four boats from Haha-Jima Island, catching an average of c. 5 t annually, 90% of which is frozen and transported to the Tokyo market. P. brunneiflagellum makes up 98% of the catch and the remainder is P. l. bispinosus, except for a few P. penicillatus. The catch was 6 t in 1997 and 9 t in The fishery is regulated by Tokyo Metropolitan Fisheries; a closed season operates from June to August to protect berried females. Species relationships and distinguishing features among the Panulirus japonicus species group Panulirus brunneiflagellum and P. l. bispinosus belong to the P. japonicus group (Group 1 of George & Main 1967) because they possess multi-articulate flagella on the exopod of both the second and third maxillipeds (Fig. 2B,C,E,F). Like all Indo-West Pacific Group 1 they also have uninterrupted dorsal transverse grooves across the abdominal segments and have one major pair of spines on the antennular plate thus separating them from the East Pacific P. interruptus and the West Atlantic P. argus which have medianly interrupted abdominal transverse grooves and two pairs of major spines on the antennular plate. Panulirus brunneiflagellum is distinguished from P. marginatus, P. cygnus, and P. femoristriga in having the transverse groove on the second abdominal segment not joining the corresponding pleural groove, whereas the grooves on all abdominal segments join the corresponding pleural grooves in these other species. In this character P. brunneiflagellum is closest to P. japonicus and P. pascuensis. However, P. japonicus also has the third and fourth abdominal transverse grooves not joining the pleural grooves and the males of P. japonicus are unique in usually having simple or bifid endopods of the pleopods of the second to fifth abdominal segments (George & Carlberg 1995). In P. brunneiflagellum, antennular flagella are plain and the abdomen spotted whereas in P. pascuensis antennular flagella are banded and pale bands are present on the abdomen. The two subspecies of P. longipes, P. l. longipes and P. l. bispinosus, together with P. femoristriga form a close group sharing the basic morphological characteristics of Group 1 multiarticulate flagella on the endopods of the second and third maxillipeds (Fig. 2E,F), medially uninterrupted absomenal transverse grooves that all join laterally with the pleural grooves (Fig. 2D) and one pair of major spines on the antennular plate. In the past, positive identification of preserved specimens was often very difficult when colour patterns had been obliterated or modified. The colours of fresh material make positive identification possible by combinations of patterns. P. l. longipes and P. l. bispinosus have banded antennal and antennular flagella (Fig. 4E), a distinct bow mark behind the cervical groove, and a single pale stripe along the lateral carapace (Fig.

6 568 New Zealand Journal of Marine and Freshwater Research, 2005, Vol. 39 Fig. 4 Shirahige-ebi (Panulirus longipes bispinosus). Male from Ogasawara Islands, 145 mm carapace length. A, dorsal view of abdomen; B, lateral view of abdomen; C, lateral view of carapace; D, dorsal view of carapace; E, antennular flagella (after Sekiguchi 1991).

7 Sekiguchi & George Description of Panulirus brunneiflagellum new species 569 4C,D); they are distinguished by the broad pale stripe along the legs of P. l. bispinosus and the broad pale stripe broken by three white spots along the legs of P. l. longipes. P. femoristriga has plain (one white, one dark ramus) antennular flagella, a wishbone mark behind the cervical groove, two pale stripes along the lateral carapace, and narrow white stripes along the legs. As mentioned above, P. brunneiflagellum is morphologically close to P. japonicus. However, as part of his study of the phyllosoma larvae of all the species in the P. japonicus group that occur in the Kuroshio subgyre, Dr T. Yoshimura (pers. comm.) used the Internet web DNA database for identification and concluded that P. brunneiflagellum has the closest genetic affinity with P. femoristriga. It is not apparent at this time whether the morphological or the molecular data provide a better appraisal of the true relationships of P. brunneiflagellum. P. japonicus have ever been recorded there and that only a few larvae of P. l. bispinosus and even fewer larvae of P. penicillatus settle from the nearby west Philippine Basin suggest that a discreet current system circulates around the volcanic island chain (Sekiguchi 1991). This current system probably developed from a major deflection of the Kuroshio subgyre by the elevation of the Iwo Jima Ridge and it is now responsible for the transport and circulation of P. brunneiflagellum phyllosoma larvae away from and back to the Ogasawara Islands. Although the Ogasawara Islands may have become subaerial only c. 2 Ma, a significant deflection of the oceanic currents and the subsequent genetic separation of P. brunneiflagellum may have been possible at an earlier time since both P. brunneiflagellum and P. japonicus would be capable of living on any of the ridges or seamounts as they gradually came closer to the surface at depths of at least 90 m. SPECIATION When suggesting possible evolutionary processes to explain the isolation of P. brunneiflagellum, volcanic ridge uprisings in the north-west Pacific are important events contributing to their subsequent geographic separation. According to Kennet (1982), from late Oligocene (30 Ma) a major pulse of volcanism occurred near the continental margin of Japan, peaking about middle Miocene (12 Ma) and this was followed in late Miocene (9 Ma) by a dramatic cold water upwelling and increased oceanic turnover, indicated by the high productivity and altered composition of diatoms. It was into this environment that P. japonicus evolved as it moved north from its equatorial origin which it had shared with the incipient P. cygnus, and it biologically responded to the Kuroshio subgyre which is the dominant oceanic current system of the region (Sekiguchi 1991). According to Springer (1982), in the middle Miocene a period of volcanism along the Palau- Kyushu Ridge resulted in the formation of the eastward moving West Mariana Ridge. Sundering along the West Mariana Ridge c. 5 Ma years ago formed the Mariana Trough and produced the volcanically active Mariana Island Ridge to the east. The Iwo Jima Ridge which contains the Ogasawara Islands is a Pliocene (2 Ma) volcanic extension of this Mariana Trench system. The fact that P. brunneiflagellum is the clearly dominant species of Panulirus around the Ogasawara Islands, that no ACKNOWLEDGMENTS We are grateful to Mr K. Nishikiori, Ogasawara Branch of Tokyo Meteropolitan Fisheries Laboratory for helping collecting Aka-ebi specimens at Ogasawara Islands. Thanks to anonymous referees for their comments on the early version of the manuscript. REFERENCES Chan, T. Y.; Chu, K. H. 1996: On the different forms of Panulirus longipes femoristriga (von Martens, 1872) (Crustacea: Decapoda: Palinuridae) with a description of a new species. Journal of Natural History 30: Chan, T. Y.; Ng, P. K. L. 2001: On the nomenclature of the commercially important spiny lobsters Panulirus longipes femoristriga (Von Martens, 1872), P. bispinosus Borradaile, 1899, and P. albiflagelum Chan & Chu 1996 (Decapoda, Palinuridae). Crustaceana 74: George, R. W. 1972: South Pacific islands rock lobster resources. Report prepared for the South Pacific Development Agency. Document WS/C7959. Rome, Food and Agriculture Organisation of the United Nations. 42 p. George, R. W. 1997: Tectonic plate movements and the evolution of Jasus and Panulirus spiny lobsters (Palinuridae). Marine and Freshwater Research 48: George, R. W.; Carlberg, K. 1995: Variation in the male pleopods of Panulirus japonicus (von Siebold, 1824). Crustaceana 68:

8 570 New Zealand Journal of Marine and Freshwater Research, 2005, Vol. 39 George, R. W.; Holthuis, L. B. 1965: A revision of the Indo-West Pacific spiny lobsters of the Panulirus japonicus group. Zoologische Verhandendellingen 72: George, R. W.; Main, A. R. 1967: The evolution of spiny lobsters (Palinuridae): a study of evolution in the marine environment. Evolution 21: Juinio, A. R.; Macaranas, J. M.; Gomez, E. D. 1991: Sympatric occurrence of two subspecies of Panulirus longipes Milne Edwards, 1868 (Decapoda, Palinuridae) and biochemical evidence of interbreeding. Memoirs of Queensland Museum 31: 209. Kennet, J. P. 1982: Marine geology. Englewood Cliffs, New Jersey, Prentice-Hall. 813 p. Nishikiori, K.; Sekiguchi, H. 2001: Spiny lobster fishery in Ogasawara (Bonin) Islands, Japan. Bulletin of Japanese Society of Fisheries Oceanography 65: (In Japanese.) Sekiguchi, H. 1991: Two forms of Panulirus longipes femoristriga (Crustacea: Palinuridae) from Ogasawara waters, Japan. Proceedings of Japanese Society of Systematic Zoology 44: Springer, V. G. 1982: Pacific plate biogeography with special reference to shorefishes. Smithsonian Contributions to Zoology 367:

Panulirus penicillatus (Olivier, 1791)

Panulirus penicillatus (Olivier, 1791) click for previous page Palinuridae 1019 Panulirus penicillatus (Olivier, 1791) Frequent synonyms / misidentifications: None / None. FAO names: En - Pronghorn spiny lobster; Fr - Langouste fourchette;

More information

INVERTEBRATE ZOOLOGY/ Crustacea

INVERTEBRATE ZOOLOGY/ Crustacea KONINKLIJKE NEDERLANDSCHE AKADEMIE VAN WETENSCHAPPEN Note on the Species of Palaemonetes (Crustacea Decapoda) found in the United States of America BY L. B. HOLTHUIS INVERTEBRATE ZOOLOGY/ Crustacea Reprinted

More information

ZOOLOGISCHE MEDEDELINGEN

ZOOLOGISCHE MEDEDELINGEN MINISTERIE VAN ONDERWIJS, KUNSTEN EN WETENSCHAPPEN ZOOLOGISCHE MEDEDELINGEN UITGEGEVEN DOOR HET RIJKSMUSEUM VAN NATUURLIJKE HISTORIE TE LEIDEN DEEL XXXIV, No. 11 1 Februari 1956 THREE SPECIES OF CRUSTACEA

More information

New records of Panulirus femoristriga Von Martens, 1872 (Crustacea Achelata Palinuridae) from Celebes and Seram Islands, Indonesia

New records of Panulirus femoristriga Von Martens, 1872 (Crustacea Achelata Palinuridae) from Celebes and Seram Islands, Indonesia Biodiversity Journal, 2016, 7 (4): 901 906 New records of Panulirus femoristriga Von Martens, 1872 (Crustacea Achelata Palinuridae) from Celebes and Seram Islands, Indonesia Rudi Alek Wahyudin 1, Agus

More information

THEJ FIRST ZOEA OF PORCELLANA. By W K. BROOKS and E. B. WILSON. With Plate* VI and VII.

THEJ FIRST ZOEA OF PORCELLANA. By W K. BROOKS and E. B. WILSON. With Plate* VI and VII. 7 "83?* {Jn-fJatoj 11 "T THEJ FIRST ZOEA OF PORCELLANA. By W K BROOKS and E. B. WILSON. With Plate* VI and VII. 7z O THE FIRST ZOEA OF PORCELLANA. By W K. BROOKS and E. B. WILSON. With Plates VI and VII.

More information

Vertical Distributions of Phyllosoma Larvae of Palinurid and Scyllarid Lobsters in the Western North Pacific

Vertical Distributions of Phyllosoma Larvae of Palinurid and Scyllarid Lobsters in the Western North Pacific Journal of Oceanography, Vol. 57, pp. 743 to 748, 2001 Short Contribution Vertical Distributions of Phyllosoma Larvae of Palinurid and Scyllarid Lobsters in the Western North Pacific HIDEKI MINAMI, NARIAKI

More information

L.B. Holthuis. Introduction

L.B. Holthuis. Introduction Cherax (Astaconephrops) minor new species, a parastacid from the mountains of Irian Jaya (W. New Guinea), Indonesia (Crustacea: Decapoda: Parastacidae) L.B. Holthuis Holthuis, L.B. Cherax (Astaconephrops)

More information

Slide 1. Slide 1. Next. 5:30:08 AM

Slide 1. Slide 1. Next.  5:30:08 AM Slide 1 Slide 1 http://www3.utep.edu/leb/mosquito/larvslide1.htm10/27/2004 5:30:08 AM Slide 1 Slide 2 Recognition that the specimens are mosquito larvae is a prerequisite to identification of the genera.

More information

TWO NEW SPECIES OF THE GENUS DYNAMENELLA FROM THE NORHTERN ARABIAN SEA (ISOPODA)

TWO NEW SPECIES OF THE GENUS DYNAMENELLA FROM THE NORHTERN ARABIAN SEA (ISOPODA) TWO NEW SPECIES OF THE GENUS DYNAMENELLA FROM THE NORHTERN ARABIAN SEA (ISOPODA) BY WAQUAR JAVED and ROSHAN AHMED Department of Zoology, University of Karachi, Karachi-32 RESUME Deux nouvelles espèces,

More information

THE GENUS PHOTIS (CRUSTACEA: AMPHIPODA) FROM THE TEXAS COAST WITH THE DESCRIPTION OF A NEW SPECIES, PHOTIS MELANICUS

THE GENUS PHOTIS (CRUSTACEA: AMPHIPODA) FROM THE TEXAS COAST WITH THE DESCRIPTION OF A NEW SPECIES, PHOTIS MELANICUS THE GENUS PHOTIS (CRUSTACEA: AMPHIPODA) FROM THE TEXAS COAST WITH THE DESCRIPTION OF A NEW SPECIES, PHOTIS MELANICUS L. D. McKinney Texas A&M University at Galveston, Moody College of Marine Technology,

More information

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi For office use: MARINE BIORESOURCES FORMS DATA ENTRY: Form- 1(general) (please answer only relevant fields;

More information

Three New Copepods from Brackish-Water Lakes of Japan

Three New Copepods from Brackish-Water Lakes of Japan Three New Copepods from Brackish-Water Lakes of Japan T AKASHI I TOl THE PRESENT PAPER deals with three new copepods found in brackish-water lakes of Japan. Two of them were obtained from the brackish

More information

TOMINEMOURA, A NEW STONEFLY GENUS FROM SABAH, EAST MALAYSIA (PLECOPTERA: NEMOURIDAE)

TOMINEMOURA, A NEW STONEFLY GENUS FROM SABAH, EAST MALAYSIA (PLECOPTERA: NEMOURIDAE) TOMINEMOURA, A NEW STONEFLY GENUS FROM SABAH, EAST MALAYSIA (PLECOPTERA: NEMOURIDAE) Ignac Sivec 1 and Bill P. Stark 2 1 Slovenian Museum of Natural History, Prešernova 20, P.O. Box 290, SLO-1001 Ljubljana,

More information

FAO SPECIES IDENTIFICATION SHEETS SYNODONTIDAE. Lizardfishes

FAO SPECIES IDENTIFICATION SHEETS SYNODONTIDAE. Lizardfishes click for previous page SYNOD 1474 FAO SPECIES IDENTIFICATION SHEETS FISHING AREAS 57,71 (E Ind. Ocean) (W Cent. Pacific) SYNODONTIDAE Lizardfishes Body elongate, usually cylindrical and with adipose fin.

More information

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi MARINE BIORESOURCES FORMS DATA ENTRY: Form- 1(general ) (please answer only relevant fields;add additional fields

More information

Longnose Skate Raja rhina

Longnose Skate Raja rhina Longnose Skate Raja rhina 42 dorsal ventral egg case Nuchal Orbital Dorsal diagram showing thorns on disc. Diagnosis The only Alaska species of Raja with an elongate snout and deeply notched pelvic fins.

More information

Four Undescribed Species of Waterstriders (Hemip.-Gerridae)

Four Undescribed Species of Waterstriders (Hemip.-Gerridae) The Ohio State University Knowledge Bank kb.osu.edu Ohio Journal of Science (Ohio Academy of Science) Ohio Journal of Science: Volume 25, Issue 1 (January, 1925) 1925-01 Four Undescribed Species of Waterstriders

More information

Habitat and Biology : Inhabits shallow waters, between 1 and 15 m depth on rocky bottoms.

Habitat and Biology : Inhabits shallow waters, between 1 and 15 m depth on rocky bottoms. click for previous page 144 Habitat and Biology : Inhabits shallow waters, between 1 and 15 m depth on rocky bottoms. Size : Maximum total body length 30 cm; common length up to 25 cm. Interest to Fisheries

More information

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi For office use: MARINE BIORESOURCES FORMS DATA ENTRY: Form- 1(general ) Ref. No.: (please answer only relevant

More information

DROPLETS FROM THE PLANKTON NET. XXI Title.

DROPLETS FROM THE PLANKTON NET. XXI Title. DROPLETS FROM THE PLANKTON NET. XXI Title RECORD OF SAPPHIRINA SALPAE GIESBRE THE NORTH PACIFIC, WITH NOTES ON IT COPEPODITE STAGES- Author(s) Furuhashi, Kenzo Citation PUBLICATIONS OF THE SETO MARINE

More information

LYSIOSQUILLOIDES MAPIA, A NEW SPECIES OF STOMATOPOD CRUSTACEAN FROM NORTHERN SULAWESI (STOMATOPODA: LYSIOSQUILLIDAE)

LYSIOSQUILLOIDES MAPIA, A NEW SPECIES OF STOMATOPOD CRUSTACEAN FROM NORTHERN SULAWESI (STOMATOPODA: LYSIOSQUILLIDAE) THE RAFFLES BULLETIN OF ZOOLOGY 2003 51(1): 43-47 National University of Singapore LYSIOSQUILLOIDES MAPIA, A NEW SPECIES OF STOMATOPOD CRUSTACEAN FROM NORTHERN SULAWESI (STOMATOPODA: LYSIOSQUILLIDAE) Mark

More information

Synaxidae 311 SYNAXIDAE. Furry lobsters

Synaxidae 311 SYNAXIDAE. Furry lobsters click for previous page Synaxidae 311 A single species occurring in the area. Palinurellus gundlachi (Von Martens, 1878) SYNAXIDAE Furry lobsters Frequent synonyms / misidentifications: None / None. FAO

More information

Chapter 10 Lecture Outline. The Restless Oceans

Chapter 10 Lecture Outline. The Restless Oceans Chapter 10 Lecture Outline The Restless Oceans Focus Question 10.1 How does the Coriolis effect influence ocean currents? The Ocean s Surface Circulation Ocean currents Masses of water that flow from one

More information

34. Records of the Zoological Surt'ey 01 India

34. Records of the Zoological Surt'ey 01 India 34. Records of the Zoological Surt'ey 01 India study, the occurrence and density of different groups seems to be influenced by the temperature. The temperature variance causes presence of less population,

More information

- 7 - DESCRIPTION OF SPECIES

- 7 - DESCRIPTION OF SPECIES I - 7 - DESCRIPTION OF SPECIES./' Anguilla bicolor McClelland ' Level-finned eel (Figs.i & 2) Length of head 6-8 times in length of body; Diameter of eye 8-10 times, Inter-orbital length 2-2.5 times, Gape

More information

Beaufortia. Redescriptions of Oratosquilla indica (Hansen) (Hansen), with accounts of a new genus. species (Crustacea, Stomatopoda)

Beaufortia. Redescriptions of Oratosquilla indica (Hansen) (Hansen), with accounts of a new genus. species (Crustacea, Stomatopoda) Beaufortia SERIES OF MISCELLANEOUS PUBLICATIONS INSTITUTE OF TAXONOMIC ZOOLOGY (ZOOLOGICAL MUSEUM) UNIVERSITY OF AMSTERDAM No. 318 Volume 25 September 1, 1976 Redescriptions of Oratosquilla indica (Hansen)

More information

OCEANOGRAPHY STUDY GUIDE

OCEANOGRAPHY STUDY GUIDE OCEANOGRAPHY STUDY GUIDE Chapter 2 Section 1 1. Most abundant salt in ocean. Sodium chloride; NaCl 2. Amount of Earth covered by Water 71% 3. Four oceans: What are they? Atlantic, Pacific, Arctic, Indian

More information

Crayfish Dissection. Materials:gloves, preserved crayfish, paper towel, dissecting pan, scissors, forceps, dissecting. Background: LME-305

Crayfish Dissection. Materials:gloves, preserved crayfish, paper towel, dissecting pan, scissors, forceps, dissecting. Background: LME-305 Living Science LME-305 Crayfish Dissection Materials:gloves, preserved crayfish, paper towel, dissecting pan, scissors, forceps, dissecting needle, dissecting pins, and pen or pencil Background: Like all

More information

1ESTERN ROCK LBSTER. Panulirus cygnus (George, 1962) L:1:ioPsIs OF BIOLOGICAL DATA ON THE. FAO Fisheries Synopsis N SAST

1ESTERN ROCK LBSTER. Panulirus cygnus (George, 1962) L:1:ioPsIs OF BIOLOGICAL DATA ON THE. FAO Fisheries Synopsis N SAST FAO Fisheries Synopsis N SAST L:1:ioPsIs OF BIOLOGICAL DATA ON THE 1ESTERN ROCK LBSTER Panulirus cygnus (George, 1962) F6 FOOD AND AGRICULTURE ORGANIZATION OF THE UNI1ED NATIONS FISEU SYNOPSES This series

More information

FAO SPECIES IDENTIFICATION SHEETS. FAMILY: SIGANIDAE FISHING AREA 51 (W. Indian Ocean) Siganis rivulatus Forsskål, 1775

FAO SPECIES IDENTIFICATION SHEETS. FAMILY: SIGANIDAE FISHING AREA 51 (W. Indian Ocean) Siganis rivulatus Forsskål, 1775 click for previous page SIGAN Sigan 1 1983 FAO SPECIES IDENTIFICATION SHEETS FAMILY: SIGANIDAE FISHING AREA 51 (W. Indian Ocean) Siganis rivulatus Forsskål, 1775 OTHER SCIENTIFIC NAMES STILL IN USE : Teuthis

More information

C. Lukhaup & R. Pekny

C. Lukhaup & R. Pekny Cherax (Astaconephrops) boesemani, a new species of crayfish (Crustacea: Decapoda: Parastacidae) from the centre of the Vogelkop Peninsula in Irian Jaya (West New Guinea), Indonesia C. Lukhaup & R. Pekny

More information

An Annotated and Illustrated Key to Multistage Larvae of Ohio Salamanders

An Annotated and Illustrated Key to Multistage Larvae of Ohio Salamanders The Ohio State University Knowledge Bank kb.osu.edu Ohio Journal of Science (Ohio Academy of Science) Ohio Journal of Science: Volume 64, Issue 4 (July, 1964) 1964-07 An Annotated and Illustrated Key to

More information

A HANDBOOK FOR LOBSTER FISHERMEN OF THE TROPICAL PACIFIC ISLANDS

A HANDBOOK FOR LOBSTER FISHERMEN OF THE TROPICAL PACIFIC ISLANDS Handbook No. 19 (1980) A HANDBOOK FOR LOBSTER FISHERMEN OF THE TROPICAL PACIFIC ISLANDS by J. PRESCOTT (SPC Lobster Project Officer) SOUTH PACIFIC COMMISSION NOUMEA, NEW CALEDONIA CONTENTS PREFACE 1 INTRODUCTION

More information

Platybema rugosus Bate, 1888:579, pl. 104, )' the IJSNM and Santa Marta specfig.

Platybema rugosus Bate, 1888:579, pl. 104, )' the IJSNM and Santa Marta specfig. REDESCRIPTION OF THE HIPPOLYTID SHRIMP TRACHYCARIS R UGOSA (BATE) (CRUSTACEA: DECAPODA: CARIDEA) FROM THE WESTERN ATLANTIC, WITH NOTES ON SEXUAL DIMORPHISM Maria M. Criales 15 October 1992 PROC. BIOL.

More information

Tips for Identifying Common Fish Species in the Bush River

Tips for Identifying Common Fish Species in the Bush River Tips for Identifying Common Fish Species in the Bush River Juvenile Largemouth bass Spot Threadfin shad Juvenile White perch The Herrings (Family Clupeidae) Alewife American shad Blueback herring Atlantic

More information

Musbir*, Sudirman, and Ridwan Bohari

Musbir*, Sudirman, and Ridwan Bohari Catch Composition and Size distribution of the spiny lobster (Panulirus spp) Related to Oceanography Parameters of the Southern Coast of South Sulawesi Musbir*, Sudirman, and Ridwan Bohari Fisheries Department,

More information

Cherax (Astaconephrops) pulcher, a new species of freshwater crayfish... 1 RESEARCH ARTICLE

Cherax (Astaconephrops) pulcher, a new species of freshwater crayfish... 1 RESEARCH ARTICLE ZooKeys 502: 1 10 (2015) doi: 10.3897/zookeys.502.9800 http://zookeys.pensoft.net Cherax (Astaconephrops) pulcher, a new species of freshwater crayfish... 1 RESEARCH ARTICLE A peer-reviewed open-access

More information

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi MARINE BIORESOURCES FORMS DATA ENTRY: Form- 1(general ) Ref. No.: (please answer only relevant fields;add additional

More information

Lecture Outlines PowerPoint. Chapter 15 Earth Science, 12e Tarbuck/Lutgens

Lecture Outlines PowerPoint. Chapter 15 Earth Science, 12e Tarbuck/Lutgens Lecture Outlines PowerPoint Chapter 15 Earth Science, 12e Tarbuck/Lutgens 2009 Pearson Prentice Hall This work is protected by United States copyright laws and is provided solely for the use of instructors

More information

Description of five new species of marine gobies (Teleostei: Gobiidae) of the genus Grallenia from the tropical western Pacific Ocean

Description of five new species of marine gobies (Teleostei: Gobiidae) of the genus Grallenia from the tropical western Pacific Ocean Description of five new species of marine gobies (Teleostei: Gobiidae) of the genus Grallenia from the tropical western Pacific Ocean Abstract GERALD R. ALLEN Department of Aquatic Zoology, Western Australian

More information

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi MARINE BIORESOURCES FORMS DATA ENTRY: Form- 1(general ) (please answer only relevant fields;add additional fields

More information

Recruitment processes of jack mackerel (Trachurus

Recruitment processes of jack mackerel (Trachurus Recruitment processes of jack mackerel (Trachurus japonicus) ) in the East China Sea (ECS) in relation to environmental conditions Chiyuki Sassa 1, Youichi Tsukamoto 1, Yoshinobu Konishi 1* Songguang Xie

More information

SEA GRANT PROGRESS REPORT

SEA GRANT PROGRESS REPORT SEA GRANT PROGRESS REPORT Project Title: The relationship between seasonal migrations of berried female lobster Homarus americanus, egg development and larval survival. Principal Investigator: Winsor H.

More information

THE RAFFLES BULLETIN OF ZOOLOGY 2009

THE RAFFLES BULLETIN OF ZOOLOGY 2009 THE RAFFLES BULLETIN OF ZOOLOGY 2009 THE RAFFLES BULLETIN OF ZOOLOGY 2009 57(2): 343 452 Date of Publication: 31 Aug.2009 National University of Singapore RADIATION OF ENDEMIC SPECIES FLOCKS IN ANCIENT

More information

Article. A new species of Glyphocrangon (Decapoda: Caridea: Glyphocrangonidae) from off the coast of western Mexico*

Article. A new species of Glyphocrangon (Decapoda: Caridea: Glyphocrangonidae) from off the coast of western Mexico* Zootaxa 2372: 358 366 (2010) www.mapress.com/zootaxa/ Copyright 2010 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) A new species of Glyphocrangon (Decapoda:

More information

Checklist of the Collembola: Key to common surface dwelling species of Collembola from the UK

Checklist of the Collembola: Key to common surface dwelling species of Collembola from the UK http://www.collembola.org/key/fkuk.pdf - Last updated on 2009.03.14 by Frans Janssens Checklist of the Collembola: Key to common surface dwelling species of Collembola from the UK This key is still under

More information

Noldo kaprusii n. sp., a new species from Crimea (Protura: Nipponentomidae)

Noldo kaprusii n. sp., a new species from Crimea (Protura: Nipponentomidae) Genus Vol. 17(2): 151-15 Wroc³aw, 30 VI 2006 Noldo kaprusii n. sp., a new species from Crimea (Protura: Nipponentomidae) JULIA SHRUBOVYCH 1 & ANDRZEJ SZEPTYCKI 2 1 State Museum of Natural History, Ukrainian

More information

Identifying Invasive Freshwater Shrimps and Isopods

Identifying Invasive Freshwater Shrimps and Isopods Identifying Invasive Freshwater Shrimps and Isopods Michael Dobson Freshwater Biological Association March 2012 Identifying Invasive Freshwater Shrimps and Isopods Michael Dobson Freshwater Biological

More information

Key to the Species of Anamixis

Key to the Species of Anamixis Key to the Species of Anamixis (Except A. stebbingi*) 1. Article 2 of gnathopod 2 with 3 or more long setae on anterior margin (1)...2 1. Article 2 of gnathopod 2 lacking long setae on anterior margin...3

More information

Commensal Leucothoidae (Crustacea, Amphipoda) of the Ryukyu Archipelago, Japan. Part III: coral rubble-dwellers

Commensal Leucothoidae (Crustacea, Amphipoda) of the Ryukyu Archipelago, Japan. Part III: coral rubble-dwellers ZooKeys 173: 11 50 (2012) Commensal Leucothoidae (Crustacea, Amphipoda) of the Ryukyu Archipelago, Japan. 11 doi: 10.3897/zookeys.173.2498 www.zookeys.org Research article A peer-reviewed open-access journal

More information

Revision of the Siriella brevicaudata species group (Crustacea: Mysida: Mysidae) from the West Indo-Pacific

Revision of the Siriella brevicaudata species group (Crustacea: Mysida: Mysidae) from the West Indo-Pacific European Journal of Taxonomy 426: 1 80 ISSN 2118-9773 https://doi.org/10.5852/ejt.2018.426 www.europeanjournaloftaxonomy.eu 2018 Daneliya M. et al. This work is licensed under a Creative Commons Attribution

More information

Received : Accepted : ABSTRACT

Received : Accepted : ABSTRACT Short Communication A NEW RECORD OF AXIOPSIS CONSOBRINA (DE MAN, 1905) (FAMILY: AXIIDAE : BORRADAILE, 1903) (CRUSTACEA : DECAPODA : THALASSINIDEA) OFF THOOTHUKUDI, SOUTHEAST COAST OF INDIA (08º 53.6 N

More information

Article.

Article. Zootaxa 3857 (2): 183 206 www.mapress.com/zootaxa/ Copyright 2014 Magnolia Press Article http://dx.doi.org/10.11646/zootaxa.3857.2.2 http://zoobank.org/urn:lsid:zoobank.org:pub:b0df10be-b582-4ed9-83ca-89729b84a574

More information

Sustainable Recreational Fishing Student Activity Workbook 6.2. Sustainable Recreational Fishing

Sustainable Recreational Fishing Student Activity Workbook 6.2. Sustainable Recreational Fishing Sustainable Recreational Fishing! Activity zone 1 - Recreational Fishing Panel Fishing is a popular pastime for more millions Australians each year but is not the only way to enjoy our lakes and coastline.

More information

Department of Marine and Environmental Sciences, University of the Ryukyus, 1 Senbaru, Nishihara-cho, Okinawa , Japan. 2

Department of Marine and Environmental Sciences, University of the Ryukyus, 1 Senbaru, Nishihara-cho, Okinawa , Japan. 2 Scientia Marina 72(1) March 2008, 37-57, Barcelona (Spain) ISSN: 0214-8358 Species of Galacantha and Munidopsis (Crustacea: Decapoda: Anomura: Galatheidae) from the deep-waters off Taiwan, with the description

More information

Using a Dichotomous Classification Key to Identify Common Freshwater Fish of New York State

Using a Dichotomous Classification Key to Identify Common Freshwater Fish of New York State D3()O!PPPPPPPPPPPPPPPPPPPPPPPPPPPPPPPPPP!Q3*)!&'(N7)*)LO!PPPPPPPPPPPPPPPPPPPPPPPPPPPPP! &73,,O!PPPPPPPPPPPP!-34!#.06*),O!PPPPPPPPPPPPPPP!C)3B?)$O!PPPPPPPPPPPPPPPPPPPPPPPPPPPPPPP! Using a Dichotomous Classification

More information

RESEARCHES ON THE COAST OF SOMALIA PSEUDOCERCEIS SELENEIDES N. SP., A NEW MARINE ISOPOD (SPHAEROMATIDAE DYNAMENINAE) FROM THE SOMALIAN COAST

RESEARCHES ON THE COAST OF SOMALIA PSEUDOCERCEIS SELENEIDES N. SP., A NEW MARINE ISOPOD (SPHAEROMATIDAE DYNAMENINAE) FROM THE SOMALIAN COAST RESEARCHES ON THE COAST OF SOMALIA PSEUDOCERCEIS SELENEIDES N. SP., A NEW MARINE ISOPOD (SPHAEROMATIDAE DYNAMENINAE) FROM THE SOMALIAN COAST BY GIUSEPPE MESSANA Centro di Studio per la Faunistica ed Ecologia

More information

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi MARINE BIORESOURCES FORMS DATA ENTRY: Form- 1(general ) (please answer only relevant fields;add additional fields

More information

First Record of Two Siriella Species of Mysids (Crustacea: Mysida) from the Korean Waters

First Record of Two Siriella Species of Mysids (Crustacea: Mysida) from the Korean Waters http://e-fas.org Original rticle Fish quat Sci 17(3), 355-360, 2014 First Record of Two Siriella Species of Mysids (rustacea: Mysida) from the Korean Waters Soo-Gun Jo * Department of Marine iotechnology,

More information

TitleA New Species of Deep-dwelling Razo.

TitleA New Species of Deep-dwelling Razo. TitleA New Species of Deep-dwelling Razo Author(s) Araga, Chuichi; Yoshino, Tetsuo Citation PUBLICATIONS OF THE SETO MARINE BIO LABORATORY (1986), 31(1-2): 75-79 Issue Date 1986-03-25 URL http://hdl.handle.net/2433/176116

More information

Captive breeding and larval development of the scyllarine lobster Petrarctus rugosus

Captive breeding and larval development of the scyllarine lobster Petrarctus rugosus New Zealand Journal of Marine and Freshwater Research ISSN: 002-330 (Print) 1175-05 (Online) Journal homepage: http://www.tandfonline.com/loi/tnzm20 Captive breeding and larval development of the scyllarine

More information

Neostrengeria binderi, a new species of pseudothelphusid crab from the eastern Andes of Colombia (Crustacea: Decapoda: Brachyura)

Neostrengeria binderi, a new species of pseudothelphusid crab from the eastern Andes of Colombia (Crustacea: Decapoda: Brachyura) DdwMagalhOes ' ' - ' Pesqutador 31 July 2000 Iffl* CPBA PROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON 113(2):401-405. 2000. Neostrengeria binderi, a new species of pseudothelphusid crab from the

More information

Contribution to the morphology of the third-instar larvae of Laccophilus poecilus KLUG (Coleoptera: Dytiscidae)

Contribution to the morphology of the third-instar larvae of Laccophilus poecilus KLUG (Coleoptera: Dytiscidae) Genus Vol. 15(1): 31-36 Wroc³aw, 30 III 2004 Contribution to the morphology of the third-instar larvae of Laccophilus poecilus KLUG (Coleoptera: Dytiscidae) EUGENIUSZ BIESIADKA and IWONA KA KAŹMIERSKA

More information

NOTONYX GUINOTAE, A NEW SPECIES OF GONEPLACID CRAB (BRACHYURA, GONEPLACIDAE) FROM LOMBOK ISLAND, INDONESIA

NOTONYX GUINOTAE, A NEW SPECIES OF GONEPLACID CRAB (BRACHYURA, GONEPLACIDAE) FROM LOMBOK ISLAND, INDONESIA NOTONYX GUINOTAE, A NEW SPECIES OF GONEPLACID CRAB (BRACHYURA, GONEPLACIDAE) FROM LOMBOK ISLAND, INDONESIA BY DWI LISTYO RAHAYU 1,3 ) and PETER K. L. NG 2,4 ) 1 ) Marine Bio-Industry Implementation Unit

More information

Cutthroat trout genetics: Exploring the heritage of Colorado s state fish

Cutthroat trout genetics: Exploring the heritage of Colorado s state fish Cutthroat trout genetics: Exploring the heritage of Colorado s state fish Metcalf et al. 2007 Molecular Ecology Metcalf et al. 2007 From Metcalf et al. 2007 But what about this one? Metcalf et al. 2007

More information

2016 ANNUAL FISH TRAWL SURVEY REPORT

2016 ANNUAL FISH TRAWL SURVEY REPORT 216 ANNUAL FISH TRAWL SURVEY REPORT The University of Rhode Island Graduate School of Oceanography The Graduate School of Oceanography (GSO) Fish trawl survey began weekly sampling two stations in Narragansett

More information

Bulletin Zoölogisch Museum

Bulletin Zoölogisch Museum Bulletin Zoölogisch Museum UNIVERSITEIT VAN AMSTERDAM Vol.11 No. 2 1987 Neohyssura atlantica n.sp. from the Cape Verde Islands (Crustacea: Isopoda: Anthuridea) Johann Wolfgang Wägele Abstract A fourth

More information

A new species of Stratiotes Thomson, 1899 (Anomura, Paguroidea, Diogenidae) from the eastern tropical Pacific

A new species of Stratiotes Thomson, 1899 (Anomura, Paguroidea, Diogenidae) from the eastern tropical Pacific A new species of Stratiotes Thomson, 1899 (Anomura, Paguroidea, Diogenidae) from the eastern tropical Pacific Manuel AYON PARENTE Michel E. HENDRICKX Laboratorio de Invertebrados Bentónicos, Unidad Académica

More information

Parastenheliidae (Copepoda: Harpacticoida) from the Isles of Scilly

Parastenheliidae (Copepoda: Harpacticoida) from the Isles of Scilly Journal of Natural History, 2006; 40(47 48): 2611 2652 Parastenheliidae (Copepoda: Harpacticoida) from the Isles of Scilly J. MICHAEL GEE Plymouth Marine Laboratory, Prospect Place, Plymouth, UK (Accepted

More information

REICHENBACHI A STAATLICHES MUSEUM FUR TIERKUNDE IN DRESDE N

REICHENBACHI A STAATLICHES MUSEUM FUR TIERKUNDE IN DRESDE N REICHENBACHI A STAATLICHES MUSEUM FUR TIERKUNDE IN DRESDE N Bd. 13 Ausgegeben : 17. Oktober 197 0 Nr. 14 A new Phalangid from Cuba, Trinimontius darlingtoni gen. n., sp. n. (Opilionidea, Cosmetidae ) with

More information

Article. Opistognathus albicaudatus, a new species of jawfish (Teleostei: Opistognathidae) from the Andaman Islands

Article. Opistognathus albicaudatus, a new species of jawfish (Teleostei: Opistognathidae) from the Andaman Islands Zootaxa 3085: 34 40 (2011) www.mapress.com/zootaxa/ Copyright 2011 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) Opistognathus albicaudatus, a new species

More information

Directed Reading. Section: Ocean Currents. a(n). FACTORS THAT AFFECT SURFACE CURRENTS

Directed Reading. Section: Ocean Currents. a(n). FACTORS THAT AFFECT SURFACE CURRENTS Skills Worksheet Directed Reading Section: Ocean Currents 1. A horizontal movement of water in a well-defined pattern is called a(n). 2. What are two ways that oceanographers identify ocean currents? 3.

More information

Two new species of the genus C y a n o se sia G o r b u n o v &

Two new species of the genus C y a n o se sia G o r b u n o v & N ach r. en to m o l. V er. A p o llo, S u p p l. 1 7 : 4 5 7-4 6 4 (1 9 9 8 ) 457 Two new species of the genus C y a n o se sia G o r b u n o v & A rita, 1995 (Lepidoptera, Sesiidae) from the Oriental

More information

By Charles Hawks Sponsored By The Angelfish Society

By Charles Hawks Sponsored By The Angelfish Society By Charles Hawks Sponsored By The Angelfish Society About this presentation In July, 2011, The Angelfish Society held its second annual Angelfish Show at the American Cichlid Association convention in

More information

Identifying Moths in Traps for Sweet Corn Pests

Identifying Moths in Traps for Sweet Corn Pests Bringing information and education into the communities of the Granite State Identifying Moths in Traps for Sweet Corn Pests Dr. Alan T. Eaton, Extension Specialist, Entomology We recommend control sprays

More information

NEW SPECIES OF OLETHREUTINE MOTHS (TORTRICIDAE) FROM TEXAS AND LOUISIANA

NEW SPECIES OF OLETHREUTINE MOTHS (TORTRICIDAE) FROM TEXAS AND LOUISIANA Journal of the Lepidopterists' Society 40(4), 1986, 322-326 NEW SPECIES OF OLETHREUTINE MOTHS (TORTRICIDAE) FROM TEXAS AND LOUISIANA EDW ARD C. KNUDSON 808 Woodstock, Bellaire, Texas 77401 Research Associate,

More information

Florida s Spiny Lobster Fishery

Florida s Spiny Lobster Fishery Florida s Spiny Lobster Fishery 2011 Seafood Webinar Series Presenters Dr. Lisa Krimsky, Miami-Dade Sea Grant Extension Agent Bryan Fluech, Collier County Sea Grant Extension Agent Using Elluminate Hand

More information

Goldfish control in the Vasse River: summary of the 2008 programme

Goldfish control in the Vasse River: summary of the 2008 programme Goldfish control in the Vasse River: summary of the 8 programme Prepared by Freshwater and Threatened Fish Research Facility Prepared for February 9 1 Goldfish control in the Vasse River: summary of the

More information

Atmospheric and Ocean Circulation Lab

Atmospheric and Ocean Circulation Lab Atmospheric and Ocean Circulation Lab name Key Objectives: The main goal of this lab is to learn about atmospheric and oceanic circulation and how these two processes are strongly inter-dependent and strongly

More information

Notes on Some ' Stomatopod / Crustacea from Southern Africa

Notes on Some ' Stomatopod / Crustacea from Southern Africa RAYMOND B. MANNI, Notes on Some ' Stomatopod / Crustacea from Southern Africa SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY* 969 NUMBER SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY NUMBER Raymond B. Manning Notes On Some

More information

Role of the oceans in the climate system

Role of the oceans in the climate system Role of the oceans in the climate system heat exchange and transport hydrological cycle and air-sea exchange of moisture wind, currents, and upwelling gas exchange and carbon cycle Heat transport Two Primary

More information

NEW SPECIES OF TROGLOBITIC SHRIMPS FROM MEXICO, WITH THE DESCRIPTION OF TROGLOMEXICANUS, NEW GENUS (DECAPODA: PALAEMONIDAE)

NEW SPECIES OF TROGLOBITIC SHRIMPS FROM MEXICO, WITH THE DESCRIPTION OF TROGLOMEXICANUS, NEW GENUS (DECAPODA: PALAEMONIDAE) JOURNAL OF CRUSTACEAN BIOLOGY, 19(1): 111-122, 1999 NEW SPECIES OF TROGLOBITIC SHRIMPS FROM MEXICO, WITH THE DESCRIPTION OF TROGLOMEXICANUS, NEW GENUS (DECAPODA: PALAEMONIDAE) Jose' Luis Villalobos, Fernando

More information

Faculty of Marine Science and Fisheries, Hasanuddin University, Makassar, Indonesia. Corresponding author: M. Musbir,

Faculty of Marine Science and Fisheries, Hasanuddin University, Makassar, Indonesia. Corresponding author: M. Musbir, Egg quantity of wild breeders of spiny lobster (Panulirus ornatus) caught from southern coastal waters of Bulukumba, South Sulawesi, Indonesia Musbir Musbir, Sudirman, Achmar Mallawa, Ridwan Bohari Faculty

More information

Shannon Point Marine Center, Western Washington University 1900 Shannon Point Road, Anacortes, WA and

Shannon Point Marine Center, Western Washington University 1900 Shannon Point Road, Anacortes, WA and Micronesica 22(2): 123-171, 1989 On the Status of Pylopaguropsis zebra (Henderson), P. magnimanus (Henderson), and Galapagurus teevanus Boone, with descriptions of seven new species of Pylopaguropsis (Crustacea:

More information

Lobsters, Crab and Shrimp

Lobsters, Crab and Shrimp Lobsters, Crab and Shrimp Crustaceans Phylum: Arthropoda ( jointed feet ) Invertebrate Exoskeleton (made of chitin) protects body and provides a place for muscles to attach to. Segmented body Jointed appendages

More information

ASIAPACIFIC-FISHWATCH CONCEPT AND THE WOLF HERRINGS (CHIROCENTRUS SPP) AS EXAMPLE. Meryl Williams Asian Fisheries Society

ASIAPACIFIC-FISHWATCH CONCEPT AND THE WOLF HERRINGS (CHIROCENTRUS SPP) AS EXAMPLE. Meryl Williams Asian Fisheries Society ASIAPACIFIC-FISHWATCH CONCEPT AND THE WOLF HERRINGS (CHIROCENTRUS SPP) AS EXAMPLE Meryl Williams Asian Fisheries Society Consumer information on fish is a well kept secret in Asia-Pacific Asian Fisheries

More information

New Calcinus Species (Decapoda: Anomura: Diogenidae) from Hawaii, with a Key to the Local Species 1

New Calcinus Species (Decapoda: Anomura: Diogenidae) from Hawaii, with a Key to the Local Species 1 'iv-» IN ST New Calcinus Species (Decapoda: Anomura: Diogenidae) from Hawaii, with a Key to the Local Species 1 JANET HAIG Allan Hancock Foundation, University of Southern California University Park, Los

More information

!"#$%&'() Mola mola *+,+-./

!#$%&'() Mola mola *+,+-./ Mola mola 2008 Summary A study on the reproductive biology of ocean sunfish Mola mola Toshiyuki akatsubo 1. Introduction Ocean sunfish, Mola mola, which belongs to the family Molidae in the order Tetraodontiformes,

More information

Cirolanid isopod crustaceans from the Townsville region of Queensland, Australia, with descriptions of six new species

Cirolanid isopod crustaceans from the Townsville region of Queensland, Australia, with descriptions of six new species JoiRNAL OF NATTRAI, HFSTORY, 1981, 15:555-605 Cirolanid isopod crustaceans from the Townsville region of Queensland, Australia, with descriptions of six new species D. M. HOLDICH and K. HARRISON Department

More information

Pre-Season Testing Protocol for the Tri-State Coastal Dungeness crab Commercial Fishery. Revised September 2013

Pre-Season Testing Protocol for the Tri-State Coastal Dungeness crab Commercial Fishery. Revised September 2013 Pre-Season Testing Protocol for the Tri-State Coastal Dungeness crab Commercial Fishery Revised September 2013 I. Purpose The Washington Department of Fish and Wildlife (WDFW), Oregon Department of Fish

More information

Significant Ecological Marine Area Assessment Sheet

Significant Ecological Marine Area Assessment Sheet Significant Ecological arine Area Assessment Sheet Name: ouhora arbour arine Values Summary: The entrance subtidal channel, tidal flats, seagrass mangrove and saltmarsh habitats of ouhora arbour have given

More information

A new killifish of the genus Melanorivulus from the upper Paraná river basin, Brazil (Cyprinodontiformes: Rivulidae)

A new killifish of the genus Melanorivulus from the upper Paraná river basin, Brazil (Cyprinodontiformes: Rivulidae) Senckenberg Gesellschaft für Naturforschung, 2013. 63 (3): 277 281 20.12.2013 A new killifish of the genus Melanorivulus from the upper Paraná river basin, Brazil (Cyprinodontiformes: Rivulidae) Wilson

More information

Orange County MPA Watch A n n u a l R e p o r t

Orange County MPA Watch A n n u a l R e p o r t Orange County MPA Watch 2 0 1 5 A n n u a l R e p o r t WHAT IS AN MPA? Marine Protected Areas (MPAs) are discrete geographic marine or estuarine areas designed to protect or conserve marine life and habitat.

More information

Article. https://doi.org/ /zootaxa

Article. https://doi.org/ /zootaxa Zootaxa 4318 (2): 295 311 http://www.mapress.com/j/zt/ Copyright 2017 Magnolia Press Article https://doi.org/10.11646/zootaxa.4318.2.4 http://zoobank.org/urn:lsid:zoobank.org:pub:f5b2a261-9454-4893-b055-5d2a483459b5

More information

A Survey of the North American Species of Merragata

A Survey of the North American Species of Merragata The Ohio State University Knowledge Bank kb.osu.edu Ohio Journal of Science (Ohio Academy of Science) Ohio Journal of Science: Volume 17, Issue 4 (February, 1917) 1917-02 A Survey of the North American

More information

CHAPTER 7 Ocean Circulation

CHAPTER 7 Ocean Circulation 1 2 3 4 5 6 7 8 9 10 11 12 13 14 CHAPTER 7 Ocean Circulation Words Ocean currents Moving seawater Surface ocean currents Transfer heat from warmer to cooler areas Similar to pattern of major wind belts

More information

THE BIOLOGY OF THE PRAWN, PALAEMON

THE BIOLOGY OF THE PRAWN, PALAEMON J. mar. bio!. Ass. U.K. (1959) 38 621-627 Printed in Great Britain 621 THE BOLOGY OF THE PRAWN PALAEMON (=LEANDER) SERRA TU S (PENNANT) BY G. R. FORSTER The Plymouth Laboratory n a recent paper Cole (1958)

More information

Striped Bass and White Hybrid (x) Striped Bass Management and Fishing in Pennsylvania

Striped Bass and White Hybrid (x) Striped Bass Management and Fishing in Pennsylvania Striped Bass and White Hybrid (x) Striped Bass Management and Fishing in Pennsylvania Prepared by R. Lorantas, D. Kristine and C. Hobbs PFBC Warmwater Unit 2005 (stocking numbers updated after 2005) Goal:

More information

Yellow Sea with contributions by John R. Apel

Yellow Sea with contributions by John R. Apel with contributions by John R. Apel Overview The is a shallow inland sea lying between northeastern China and the Korean Peninsula, with depths in its central north-south trough in excess of 60 to 80 m

More information

A review of Strobopagurus Lemaitre, 1989 (Crustacea: Decapoda: Paguroidea: Parapaguridae), with description of a new species*

A review of Strobopagurus Lemaitre, 1989 (Crustacea: Decapoda: Paguroidea: Parapaguridae), with description of a new species* SCI. MAR., 68 (3): 355-372 SCIENTIA MARINA 2004 A review of Strobopagurus Lemaitre, 1989 (Crustacea: Decapoda: Paguroidea: Parapaguridae), with description of a new species* RAFAEL LEMAITRE Department

More information