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1 Page 1 of 9 NOAA Fisheries Office of Protected Resources OPR Home About OPR Species Permits Laws & Policies Programs Education Publications Species Marine Mammals Cetaceans Pinnipeds Marine Turtles Marine & Anadromous Fish Marine Invertebrates & Plants Species of Concern Threatened & Endangered Species Critical Habitat Maps Contact OPR Glossary OPR Site Map Search OPR Blue Whale (Balaenoptera musculus) Status Taxonomy Species Description Habitat Distribution Population Trends Threats Conservation Efforts Regulatory Overview Key Documents More Info Status ESA Endangered - throughout its range MMPA Depleted - throughout its range Taxonomy Kingdom: Animalia Phylum: Chordata Class: Mammalia Order: Cetacea Family: Balaenopteridae Genus: Balaenoptera Species: musculus Subspecies of the blue whale include: Northern Hemisphere blue whale (B. m. musculus) Antarctic blue whale (B. m. intermedia) Pygmy blue whale (B. m. brevicauda) - a significantly smaller and morphologically distinct form found in the subantarctic zone of the southern Indian Ocean and southwestern Pacific Ocean. Species Description Physical characteristics The blue whale, Balaenoptera musculus (Linnaeus 1758), is a cosmopolitan species of baleen whale. Blue whales in the Northern Hemisphere are generally smaller than those in the Southern Ocean. Maximum body length in the North Atlantic was about 88.5 feet (27 m) and the Blue Whale (Balaenoptera musculus) Photo: NMFS Northeast Fisheries Science Center Did You Know? The blue whale is the largest animal ever known to have lived on Earth; they can weigh over 330,000 pounds (150,000 kg). Despite their massive size, blue whales feed almost exclusively on krill, tiny shrimp-like animals about the size of a jelly bean! largest blue whale reported from the North Pacific was about 88 feet (26.8 m). Adults in the Antarctic can reach a maximum body length of about 108 feet (33 m) and can weigh more than 330,000 pounds (150,000 kg). As is true of other baleen whale species, female blue whales are somewhat larger than males. Blue whales are identified by the following characteristics: a long-body and comparatively slender shape; a broad, flat "rostrum" when viewed from above; a proportionately smaller dorsal fin than other baleen whales; and a mottled gray color pattern that appears light blue when seen through the water. Diet The primary and preferred diet of blue whales is krill (euphausiids). In the North Atlantic, blue whales feed on two main euphausiid species: Thysanoëssa inermisand Meganyctiphanes norvegica. In addition, T. raschiiand M. norvegicahave been recorded as important food sources of blue whales in the Gulf of St. Lawrence. In the North Pacific, blue whales prey mainly on Euphausia pacificaand secondarily on T. spinifera. While other prey species, including fish and copepods, have been mentioned in the scientific literature, these are not likely to contribute significantly to the diet of blue whales. Life History

2 Page 2 of 9 Scientists have yet to discern many details regarding the life history of the blue whale. The best available science suggests the gestation period is approximately months and that blue whale calves are nursed for about 6-7 months. Most reproductive activity, including births and mating, takes place during the winter. Weaning probably occurs on, or en route to, summer feeding areas. The average calving interval is probably two to three years. The age of sexual maturity is thought to be 5-15 years. There are no known differences in the reproductive biology of blue whales in the North Pacific and North Atlantic oceans. Blue Whale video (Balaenoptera musculus) Photo: NOAA's National Marine Sanctuaries Habitat Blue whales inhabit sub-polar to sub-tropical latitudes. Poleward movements in spring allow the whales to take advantage of high zooplankton production in summer. Movement towards the subtropics in the fall allows blue whales to reduce their energy expenditure while fasting, avoid ice entrapment in some areas, and engage in reproductive activities in warmer waters of lower latitudes. Although the species is often found in coastal waters, blue whales are thought to occur generally more offshore than humpback whales, for example. Distribution Global Blue whales are found in oceans worldwide and are separated into populations by ocean basin in the North Atlantic, North Pacific, and Southern Hemisphere. They follow a seasonal migration pattern between summering and wintering areas, but some evidence suggests that individuals remain in certain areas yearround. The extent of knowledge concerning distribution Blue Whale Range Map (click for larger view PDF) and movement varies with area and migratory routes are not well known but, in general, distribution is driven largely by food requirements. For example, in the Gulf of St. Lawrence blue whales prefer deep waters where krill is concentrated. North Atlantic The overall distribution of blue whales in the North Atlantic extends from the subtropics to Baffin Bay and the Greenland Sea. Blue whales are most frequently sighted in the waters off eastern Canada, with the majority of recent records from the Gulf of St. Lawrence, where they are present throughout most of the year. They are most common during the summer and fall feeding seasons and typically leave by early winter to avoid ice entrapment. Although they are rare in the shelf waters of the eastern U.S., occasional sightings of blue whales have been made off Cape Cod, Massachusetts. It is believed this region may represent the current southern limit of the blue whales' feeding range. In addition, there is some evidence to suggest that blue whales occasionally stray into the Gulf of Mexico and the Caribbean, but they are less common in these waters. Some scientists believe blue whales in the North Atlantic occur in relatively discrete feeding populations (Sigurjónsson and Gunnlaugsson, 1990) whereas other evidence suggests blue whales may comprise one "panmictic" population (Clark, 1994). North Pacific The blue whale's range is known to encompass much of the North Pacific Ocean, from Kamchatka to southern Japan in the west, and from the Gulf of Alaska and California south to at least Costa Rica in the east. The species is found primarily south of the Aleutian Islands and the Bering Sea. Whaling and sighting data suggest the existence of at least five subpopulations of blue whales, with an unknown degree of mixing among them: 1. southern Japan (which appears to have been virtually "extirpated" by whaling) 2. northern Japan/Kurils/Kamchatka 3. Aleutian Islands (the central stock, which may winter in deep water north of Hawaii) 4. eastern Gulf of Alaska 5. California/Mexico

3 Page 3 of 9 The International Whaling Commission (IWC), however, formally considers only one management stock for blue whales in the North Pacific. For management purposes under the MMPA, blue whales inhabiting U.S. waters in the North Pacific are divided into two "stocks": Western and Eastern. Based on acoustic and whaling data, it is believed that the eastern stock winters off Mexico, central America, and as far south as 8 E S, and feeds during summer off the U. S. West Coast and to a lesser extent, in the Gulf of Alaska and in central North Pacific waters. The Western stock appears to feed in summer southwest of Kamchatka, south of the Aleutians, and in the Gulf of Alaska (Stafford, 2003 and Watkins et al., 2000); in winter they migrate to lower latitudes in the western Pacific and less frequently in the central Pacific, including Hawaii (Stafford et al., 2001). Blue whales accompanied by young calves have been observed often in the Gulf of California from December through March, and, thus, at least some calves may be born in or near the Gulf (Sears, 1990). Therefore, this area is probably an important calving and nursing area for the species. Northern Indian Ocean There is a "resident" population of unknown taxonomic status present in the northern Indian Ocean. Blue whales sightings have been reported from the Gulf of Aden, Persian Gulf, Arabian Sea, and across the Bay of Bengal to Burma and the Strait of Malacca. The migratory movements of these whales are unknown. Southern Hemisphere In the Southern Hemisphere distributions of B. m. intermedia and B. m. brevicauda may be segregated. B. m. intermedia occurs mainly in relatively high latitudes south of the "Antarctic Convergence" and close to the ice edge. B. m. brevicada is typically distributed north of the Antarctic Convergence. Changes in distribution There is some evidence which suggests the distribution and migratory patterns of blue whales may have changed in at least four areas: northern Norway, southern Japan, eastern Aleutian Islands, and northern California. In northern Norway (ie. Finnmark, Bear Island, and Svalbard) the paucity of sightings during recent surveys along the coast where blue whales were common in the late 1800s and early 1900s, may suggest that the historic distribution has changed (Christensen et al., 1992). However, it could also indicate depletion of the population by whaling. In the western North Pacific, the lack of blue whales off southern Japan today may also suggest that the distribution of these animals has changed or that the animals of this region have been extirpated. South of the eastern Aleutian Islands (160ºW and 180ºW), relatively large concentrations of blue whales were documented in the 1970s but the species appears rare there today, suggesting that illegal and unreported whaling depleted the population (Stewart et al., 1987 and Forney and Brownell, 1997). Off northern California (eg. Farallon Islands, Moss Landing, and Trinidad), the recent appearance of numerous blue whales is noteworthy in light of their rarity in these regions prior to the late 1970s. Calambokidis (1995) concluded that such changes in distribution reflect a shift in feeding from the more offshore euphausiid, Euphausia pacifica, to the primarily "neritic" euphausiid, Thysanoëssa spinifera. Population Trends Global Blue whales were significantly depleted by commercial whaling activities worldwide. In the Southern Hemisphere, pre-exploitation population estimates range from 150,000 to 210,000 whales; recent abundance estimates range between 400 and 1,400 whales. In the North Pacific, pre-exploitation population size is estimated as approximately 4,900 blue whales, whereas the current population estimate is a minimum of 3,300 blue whales. In the North Atlantic, estimates for the entire basin are considered unreliable, but range from 1,100 to 1,500 blue whales preexploitation, and 100 to 555 whales currently (IWC, 2007; see Table 1 below).

4 Page 4 of 9 Southern Hemisphere North Pacific North Atlantic Pre-exploitation 150, ,000 4,900 1,100-1,500 Current 400-1,400 3, Table 1. Global abundance estimates for blue whales by ocean basin (IWC, 2007). Western North Atlantic Based on data from individuals found only in the Gulf of St. Lawrence, the current minimum population estimate for the western North Atlantic stock is 308 whales. Mitchell (1974) estimated that the blue whale population in the western North Atlantic during the late 1960s and early 1970s was in the very low hundreds at most, and according to its most recent stock assessment, NMFS has no evidence to refute this estimate. More than 320 individual blue whales have been photoidentified in the Gulf of St. Lawrence and the total number of photo-identified individuals for eastern Canada and New England is 352 as of Fall According to the most recent NMFS stock assessment, there is insufficient data to determine population trends for this species. There are no accurate estimates of pre-exploitation population sizes available for the western North Atlantic ; however, several rough estimates have been made by scientists. For example, Sergeant (1966) approximated that there were 1,500 blue whales in eastern Canadian waters and Allen (1970) inferred a population greater than 1,100. Eastern North Atlantic There is no current estimate for the number of blue whales in eastern North Atlantic waters. However, some data have been collected for blue whales in Icelandic waters. As of autumn 1997, 32 individuals had been photo-identified in Icelandic waters. Additional studies have suggested that the population in Icelandic and neighboring waters may be in the high hundreds (Gunnlaugsson and Sigurjónsson, 1990 and Sigurjónssonand Gunnlaugsson, 1990) or greater than 1,000 (Christensen et al., 1992). Sightings data off the west and southwest coasts of Iceland suggest the population has been increasing at about five percent per year since the late 1960s (Sigurjónsson and Gunnlaugsson, 1990). Despite these differences in pre-exploitation estimates and the lack of estimates for current population abundance, it is clear that blue whale stocks in the western, eastern, and central North Atlantic were severely depleted by the time that legal protection was introduced in Western North Pacific There is insufficient data to evaluate the current abundance or population trends of blue whale stocks in the western North Pacific, but blue whales in Hawaiian waters are considered extremely rare. Eastern North Pacific Blue whale abundance in the eastern Pacific is about 1,700. Along the California coast blue whale abundance has been increasing during the past two decades (Calambokidis et al., 1990; Barlow, 1994 and Calambokidis, 1995). The magnitude of this apparent increase is too large to be accounted for by population growth alone and, therefore, it is assumed that a shift in distribution has occurred. Although the population in the North Pacific is expected to have grown since protection began in 1966, the possibility of continued unauthorized takes, incidental ship strikes and mortality, and serious injury in fishing gear makes this uncertain. Also, the evident scarcity of blue whales in areas of former abundance (e.g., Gulf of Alaska and near the Aleutian Islands) suggests that the increasing trend does not apply to the species' entire range in the eastern North Pacific. Threats The primary threats currently facing blue whales are:

5 Page 5 of 9 vessel strikes fisheries interactions Additional threats that have not been documented but could potentially affect these populations include: natural mortality anthropogenic noise competition for prey resources habitat degradation vessel disturbance Blue Whale (Balaenoptera musculus) Photo: NOAA Although whaling substantially reduced blue whale populations worldwide during the first half of the 20th century, whaling is no longer considered a threat today. Ship strikes A primary threat to blue whales is mortality and serious injury caused by ship strikes. In the eastern North Pacific, ship strikes were implicated in the deaths of blues whales in 1980, 1986, 1987, 1993 and The average number of blue whale mortalities in California attributed to ship strikes was 0.2 per year from and from In September 2007, three blue whale mortalities were confirmed to be caused by ship strikes in the Santa Barbara Channel off Southern California. These deaths were part of a larger Unusual Mortality Event, declared by the Working Group on Marine Mammal Unusual Mortality Events. In the western North Atlantic, at least nine percent of the whales in the Gulf of St. Lawrence have injuries or scars attributed to contact with ships (Sears et al., 1990). This area has a relatively high risk of ship strikes because the St. Lawrence Seaway has heavy ship traffic during the time of year when blue whales are relatively abundant. In March 1998, a dead 20 m (66ft) male blue whale was brought into Rhode Island waters on the bow of a tanker. The cause of death was determined to be ship strike. There are no records of ship strikes for blue whales in the western North Pacific but mortalities caused by ship strikes have likely occurred without being reported. Fisheries interactions Despite the lack of observed fisheries interactions in the last decade, incidental take in fisheries threaten blue whales for two reasons. First, past records of entanglements suggest that interaction with fishing gear may affect blue whales. Second, entanglement rates may be underestimated because blue whales may break through or carry away fishing gear, perhaps suffering unrecorded subsequent mortalities or serious injuries. It is also likely that stranding data underestimate the number of whales killed by fishing gear, because most whales do not drift far enough to strand on beaches or to be detected floating in the nearshore corridor. Although direct observation of mortality is rare, at least two documented cases of dead blue whales are apparently from the effects of entanglement in fishing gear (one in 1987 off Stellwagen Bank and the other in the 1990s in the Gulf of St. Lawrence). Natural mortality Little is known about natural mortality of blue whales in the North Atlantic. Ice entrapment is known to injure and kill some blue whales, particularly along the southwest coast of Newfoundland during late winter and early spring. In the Gulf of St. Lawrence, scarring on the dorsal surface of some whales is thought to be from contact with ice. Also, two blue whales bore rake-like markings assumed to be from the teeth of killer whales (Orcinus orca) (Sears et al., 1990). However, no direct evidence of predation on blue whales has been reported in this area. A well-documented observation of killer whales attacking a blue whale off Baja California proves that blue whales are at least occasionally vulnerable to these predators (Tarpy, 1979). A high proportion of the blue whales in the Gulf of California bear injuries or rake-like scars that are the result of encounters with killer whales (Sears, 1990), although the extent to which such attacks are fatal is unknown. Unlike in the western North Atlantic, injury or suffocation from ice entrapment is not known to affect blue whales in the North Pacific. Other potential non-primary threats Anthropogenic noise, competition for prey resources, habitat degradation, and vessel

6 Page 6 of 9 disturbance are additional concerns; however, there is little evidence available to describe or quantify the impacts of these threats on blue whales. For example, while anthropogenic noise is known to be a threat for other cetaceans little is known about whether, or how, vessel noise affects blue whales. Competition for prey resources is possible between blue whales and other sympatric whale species but there is no direct evidence for interspecific competition (Clapham and Brownell, 1996) and it is unlikely blue whales compete with humans for prey resources. Habitat degradation (e.g., chemical pollution) has occurred in some areas of the North Atlantic (e.g., St. Lawrence River) but the impacts of this degradation have not been proven to affect blue whales (O'Shea and Brownell, 1994) and are understudied. Lastly, the effect of vessel disturbance (e.g., whalewatching boats) is of concern but there is no direct evidence to demonstrate that persistent close approaches by tour boats has a negative effect on blue whales. Whaling Blue whales were hunted as early as the 19th century but were not intensively hunted until the turn of the 20th century. From the early 1900s to the mid-1960s blue whales were hunted in all the world's oceans and their populations significantly reduced. At least 9,500 blue whales were taken by commercial whalers throughout the North Pacific between 1910 and 1965 (Ohsumi and Wada, 1972) and at least 11,000 were taken in the North Atlantic from the late 19th to mid 20th centuries (Sigurjónsson and Gunnlaugsson, 1990). In 1966, the IWC banned commercial whaling for blue whales and no whaling (either "aboriginal subsistence" or commercial) occurs at present. Illegal whaling for blue whales has been documented or is likely to have occurred in the North Pacific, North Atlantic, and Southern Hemisphere. A small number of illegal kills of blue whales have been documented in the Northern Atlantic off Canada and Spain, and in the eastern North Atlantic. Blue whales were also killed in the Southern Hemisphere by the Soviet Union after 1966 (Zemsky et al., 1995a, 1995b). Some illegal whaling by the USSR also occurred in the North Pacific (Yablokov, 1994); it is likely that blue whales were among the species taken by these operations, but the extent is not known. No aboriginal whaling for blue whales is known to exist. Aboriginal hunters who target whales off the coast of the Lesser Antilles and Greenland do not target blue whales. Further, Norwegian whaling operations target only minke whales, and the commercial whaling stations in Iceland, Spain, and the Portugese islands of the Azores and Madeira remain officially closed. Conservation Efforts Conservation actions for the blue whale are on-going, and the most recent efforts by the U.S. include the following: Monitoring the status of the California/Oregon/Washington stock of blue whales via shipboard surveys, conducted every 3 years. Placing observers onboard vessels in the California/Oregon swordfish/thresher shark drift gillnet fishery to monitor the take of protected species, including marine mammals. Blue Whale (Balaenoptera musculus) Photo: NOAA Implementing marine mammal take reduction measures identified in the Pacific Offshore Cetacean Take Reduction Plan (including the use of acoustic pingers) to reduce the bycatch of marine mammals. In 1998, NMFS published the Blue Whale Recovery Plan [pdf]. The Plan details the comprehensive and long-term conservation efforts for blue whales and is available below under the Key Document section. Regulatory Overview Within the U.S. the blue whale is listed as endangered throughout its range under the Endangered Species Act of 1973 and is listed as "depleted" throughout its range under the Marine Mammal Protection Act of Internationally, blue whales received complete legal protection from commercial whaling in 1966 under the

7 Page 7 of 9 International Convention for the Regulation of Whaling. In addition, the 2007 IUCN Red List of Threatened Species populations of blue whales, as follows: lists four geographical 1. B.m. musculus North Atlantic stock as "vulnerable" 2. B.m. musculus North Pacific stock as "Lower Risk/conservation dependent" 3. B.m. intermedia as "endangered" 4. B.m. brevicauda as data deficient However, these IUCN listings are not regulatory for blue whales in U.S. waters. Key Documents (All documents are in PDF format.) Title Federal Register Date Recovery Plan n/a 07/1998 ESA Listing Rule 35 FR /02/1970 Stock Assessment Reports n/a various More Information NMFS National Marine Mammal Laboratory Blue Whale Information and Research NMFS Southwest Fisheries Science Center Blue Whale Information Kids' Times: Blue Whale [pdf] NOAA News: NOAA Scientists Sight Blue Whales in Alaska: Critically Endangered Blue Whales Rarely Seen in Alaska Waters (2004) NOAA's National Marine Sanctuaries Channel Islands Sanctuary Blue Whale Species Card with video Cordell Bank Sanctuary Blue Whale Species Card with video Gulf of the Farallones Sanctuary Blue Whale Species Card Monterey Bay Sanctuary Blue Whale Species Card with video Sounds of the Blue Whale from NOAA's Pacific Marine Environmental Laboratory U.S. Fish & Wildlife Service Blue Whale Species Profile International Whaling Commission IUCN Red List of Threatened Species References Allen, K.R A note on baleen whale stocks of the north west Atlantic. Report to the International Whaling Commission. 20: Barlow, J Abundance of large whales in California coastal waters: a comparison of ship surveys in 1979/80 and in Report to the International Whaling Commission. 44: Bowles, A.E., M. Smultea, B. Würsig, D.P. DeMaster, and D. Palka Relative abundance and behavior of marine mammals exposed to transmissions from the Heard Island Feasibility Test. J. Acoust. Soc. America 96: Calambokidis, J Blue whales off California. Whalewatcher 29(1):3-7. Calambokidis, J., G.H. Steiger, J.C. Cubbage, K.C. Balcomb, C. Ewald, S. Kruse, R. Wells, and R. Sears Sightings and movements of blue whales off central California from photo-identification of individuals. Report to the International Whaling Commission, Special Issue 12: Christensen, I., T. Haug, and N. Øien Seasonal distribution, exploitation and present abundance of stocks of large baleen whales (Mysticeti) and sperm whales (Physeter macrocephalus) in Norwegian and adjacent waters. ICES J. mar. Sci. 49: Clapham, P.J., and R.L. Brownell, Jr Potential for interspecific competition in baleen whales. Report to the International Whaling Commission. 46: Clark, C.W Blue deep voices: insights from the Navy's Whales '93

8 Page 8 of 9 program. Whalewatcher 28(1):6-11. Forney, K.A., and R.L. Brownell, Jr Preliminary report of the 1994 Aleutian Island marine mammal survey. Paper SC/48/011 presented to the International Whaling Commission Scientific Committee, June 1996 (unpublished). Gunnlaugsson, T., and J. Sigurjónsson NASS-87: Estimation of whale abundance based on observations made on board Icelandic and Faroese survey vessels. Report to the International Whaling Commission. 40: IWC International Whaling Commission statistics. URL: Mitchell, E Present status of Northwest Atlantic fin and other whale stocks. Pp In: Schevill, W.E. (ed.) The whale problem: a status report. Harvard University Press, Cambridge, MA. 419pp. Ohsumi, S., and S. Wada Stock assessment of blue whales in the North Pacific. Unpublished working paper for the 24th meeting of the Scientific Committee of the International Whaling Commission, 20 pp. O'Shea, T.J., and R.L. Brownell, Jr Organochlorine and metal contaminants in baleen whales: a review and evaluation of conservation implications. Sci. Total Environment 154: Rice, D.W Blue whale. Pp In: D. Haley (ed.) Marine mammals of eastern North Pacific and Arctic waters. Second edition. Pacific Search Press. Sears, R The Cortez blues. Whalewatcher 24(2): Sears, R., F.W. Wenzel, and J.M. Williamson The Blue Whale: A Catalogue of Individuals from the Western North Atlantic (Gulf of St. Lawrence). Mingan Island Cetacean Study, St. Lambert, Quebec. 27 pp. Sears, R., J.M. Williamson, F.W. Wenzel, M. Bérubé, D. Gendron, and P. Jones Photographic identification of the blue whale (Balaenoptera musculus) in the Gulf of St. Lawrence, Canada. Report to the International Whaling Commission, Special Issue 12: Sergeant, D.E Populations of large whale species in the western North Atlantic with special reference to the fin whale. Fisheries Research Board of Canada, Arctic Biological Station, Circular No. 9. Sigurjónsson, J., and T. Gunnlaugsson Recent trends in abundance of blue (Balaenoptera musculus) and humpback whales (Megaptera novaeangliae) off west and southwest Iceland, with a note on occurrence of other cetacean species. Report to the International Whaling Commission. 40: Stafford, K.M Two types of blue whale calls recorded in the Gulf of Alaska. Marine Mammal Science. 19: Stafford, K.M., S.L. Nieukirk, and G.G. Fox Geographic and seasonal variation of blue whale calls in the North Pacific. Journal of Cetacean Research and Management 3(1): Stewart, B.S., S.A. Karl, P.K. Yochem, S. Leatherwood, and J.L. Laake Aerial surveys for cetaceans in the former Akutan, Alaska, whaling grounds. Arctic 40: Tarpy, C Killer whale attack! National Geographic 155: Watkins, W.A., J.E. George, M.A. Daher, K. Mullin, D.L. Martin, S.H. Haga, and N.A. DiMarzio Whale call data for the North Pacific November 1995 through July 1999: Occurrence of calling whales and source locations from SOSUS and other acoustic systems. Technical Report WHOI_00_02 available from Woods Hole Oceanographic Institution. 160pp. Yablokov, A.V Validity of whaling data. Nature 367:108. Yochem, P.K., and S. Leatherwood Blue whale Balaenoptera musculus (Linnaeus, 1758). Pp In: Ridgway, S.H. and R. Harrison (eds.), Handbook of Marine Mammals, Vol. 3: The Sirenians and Baleen Whales. Academic Press, London. 362 pp. Zemsky, V.A., A.A. Berzin, Y.A. Mikhaliev, and D.D. Tormosov. 1995a. Soviet Antarctic pelagic whaling after WWII: review of actual catch data. Report of the Sub-committee on Southern hemisphere baleen whales. Rep. int. Whal. Commn. 45: Zemsky, V.A., A.A. Berzin, Y.A. Mikhaliev, and D.D. Tormosov. 1995b. Antarctic whaling data ( ). Center for Russian Environmental Policy, Moscow. 320 pp. [In Russian with English summaries.]

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