NEW SPECIES OF NORTH AMERICAN ACMAEODERA (COLEOPTERA : BUPRESTIDAE) (1)

Transcription

1 NEW SPECIES OF NORTH AMERICAN ACMAEODERA (COLEOPTERA : BUPRESTIDAE) (1) by WILLIAM F. BARR University of ldaho Moscow, ldaho ABSTRACT: Twenty four new species of Acrnaeodera are described. These species have restricted distributional ranges in portions of Arizona, California, Nevada, New Mexico, Oklahoma, and Texas of the United States and in Baja California, Colima, Guerrero, Michoacan, Oaxaca and San Luis Potosi of Mexiw. Study of large collections of North American Acmaeodera including previously unobtainable series of the majority of the described species and examination of all the primary type specimens that have been deposited in collections in the United States have indicated the presence of a number of new species and the need for some taxonomic changes in the genus. As an initial contributory step towards a classification of the North American species of Acmaeodera the present paper is concerned with the naming and describing of 24 new species. Appreciation is expressed to the many individuals and organizations who are listed as recipients of paratype material for sending me specimens for study. Special acknowledgment also is given to M. A. Cazier, formerly of the American Museum of Natural History; the late Harold Grant, Philadelphia Academy of Sciences; J. F. Gates Clarke, U. S. National Museum; P. J. Darlington, Museum of Comparative Zoology; J. N. Knull, Ohio State University; and H. B. Leech, California Academy of Sciences for allowing (1) Published with the approval of the Director of the Idaho Agricultural Experiment Station as research paper no. 788.

2 William F. Barr me to study type specimens in their care. I am further indebted to tbc American Philosophical Society and the National Science Foundation for 6namial assistance under Penrose Fund grant no and Science Faculty Felloarhip no , respectively, that afforded me the opportunity of travel to & certain field studies and to make examination of type specimens. Paratypes of several species described herein are deposited in the British Museum (Natural History), Museu Bocage, Museum G. Frey and Mmenm National dlhistorie Naturelle. Acmaeodera chiricahuae Barr, new species 1) Male: Medium-sized, somewhat elongate; black with a bronzy cast: pronotum with a narrow, submarginal yellow stripe extending from base to anterior third; elytra with a narrow, lateral yellow stripe extending from base to behind middle and with a pair of narrow, transverse reddish spots at about apical fourth and in front of apices that extend from lateral margin to war suture, the lateral yellow stripe expanded at middle of elytra, left elytron with an additional tiny yellow spot near suture at about apical third; upper surface clothed with moderately long, erect dark hairs; ventral surface and legs irregularly clothed with erect and suberect, fine silvery hairs. Head rather finely, densely punctured; vertex with a faintly indicated, longitudinal median carina; front transversely convex except for upper portion which is subflattened at middle and the area above clypeus which is feebly depressed; clypeus densely punctured, front margin deeply and broadly, triangularly emarginate; antenna shining, extending to middle of pronotum, serrate from fifth segment, outer segments nearly twice as wide as long. Pronoturn slightly more than twice as wide as long, wider than elytra; disk subflattened except for a broad and shallow median depression in front of base that extends anteriorly as a faintly indicated, shallow median groove, and for a shallow depression extending obliquely from basal pits, transverse depression behind front margin very feebly indicated on either side of middle; sides widest in front of base, broadly, arcuately narrowing to front angles; lateral margins strongly reflexed, visible from above throughout; front angles strongly produced, acute; front margin very feebly lobed at middle; hind margin feebly emarginate; surface rather finely, sparsely punctured at middle, becoming very coarsely and densely punctured laterally. Elytra subconvex except for a broad, shallow median depression at base; sides more or less vertical on posterior half, when viewed from above 'they are subparallel to middle and then broadly, arcuately converging to the conjointly, rather broadly rounded

3 New species of North American Acmaeodera 147 apices; front margin slightly beaded at middle; front angles from side view appearing to be broadly rounded, but actually produced ventrally, acute; lateral margins when viewed from the side feebly bisinuate to basal fourth where they are slightly emarginate and then very broadly and shallowly sinuate to apices, apical half very coarsely and sharply serrate; stria1 punctures circular and deep, small near suture, becoming elongate apically and slightly coarser laterally, striae feebly impressed behind middle and near sides; interstrial spaces subflattened, each with a row of tiny punctures, first, second and third interstrial spaces feebly and uniformly elevated, fifth interstrial space very feebly elevated to about middle, seventh interstrial space slightly elevated, ninth interstrial space distinctly elevated to about basal fourth, suture strongly elevated except on about basal third. Ventral surface very coarsely, densely and very shallowly punctured at sides of thoracic sternites, more sparsely punctured at middle, more finely punctured on metacoxa, abdominal sternites with densely placed, shallow crescent-shaped punctures, middle of first sternite sparsely punctured; front margin of prosternum strongly retracted, not reflexed, shallowly emarginate, with a small rounded tooth on either side of middle; last visible abdominal sternite broadly rounded, sides distinctly reflexed, with a very large and smooth subapical plate that is strongly undercut and very broadly rounded at apex. Length: 9.5 mm. Width: 3.8 mm. Holotype male (California Academy of Sciences) and 4 paratypes from 5 miles west of Portal, Cochise County, Arizona, September 1, 1957 (W. F. Barr), September 30, 1957 (W. Rosenberg). Additional paratypes from localities in Cochise County, Arizona as follows: 64 specimens from Southwest Research Station, 5 miles west of Portal, 5400 ft., August 19, 21, 24, 26, 27, 29, September 26, 30, October 5 and 11, 1956 and October 3, 1955 (E. Ordway), August 23, 1961 and 26, 1957 (M. Cazier) and August 26, 1959 (R. S. Beal); 5 specimens from 3 miles west of Portal, August 28, 1957 (W. F. Barr); 11 specimens from 8 miles west of Portal, September 5, 1957 (W. F. Barr); 1 specimens from Portal, August 16, 1964 (L. D. Anderson); 11 specimens from Pinery Canyon, Chiricahua Mountains, August 29, 1957 (W. F. Barr), September 5, 1958 (W. F. Barr), September 8, 1950 (T. Cohn, P. Boone and M. Cazier), October 8, 1955 (Truxal and Martin); 4 specimens from Pinery creek, Chiricahua Mountains, August 27, 1952 (B. Malkin); 15 specimens from Rucker Canyon, Chiricahua Mountains, September 7, 1959 (F. H. Parker); 22 specimens from Tex Canyon, Chiricahua Mountains ft., October 7, 1927 (J. A. Kusche); 3 specimens from Paradise, Chiricahua Mountains, ft., August 22,1927 (J. A. Kusche), August 23,1966 (G. R. Ballmer); 7 specimens from 4 miles west of Paradise, Chiricahua

4 148 William F. Barr Mountains, ft., September 9, 1950 (W. Gertsch and M. Cazier); 1 specimen from Sunny Slope, Cave Creek, Chiricahua Mountains, August 26, 1964 (R. A. Rauch); 16 specimens from Cave Creek, Chiricahua Mountains (W. H. Nutting), 5400 ft., September 9, 1958 and September 11, 1954 (P. A. Timberlake); 1 specimen from Cave Creek Cayon, Chiricahua Mountains, September 11, 1954 (J. C. Hall); 4 specimens from Upper Cave Creek, Chiricahua Mountains, August 29, 1957 (W. F. Barr); 1 specimen from South Fork of Cave Creek, Chiricahua Mountains, August 27, 1965 (E. I. Schlinger); 28 specimens from Onion Saddle, Chiricahua Mountains, 7600 ft., August 15, 1965 (G. W. Forister), August 16, 1969 and August 31, 1965 (B. Tilden), August 29,1966 (M. A. Cazier) and September 10,1954 (J. C. Hall, P. A. Timberlake); 2 specimens from 9 miles west of Onion Saddle, Chiricahua Mountains, September 10, 1954 (P. A. Timberlake); 5 specimens from Rustlers Park, Chiricahua Mountains, 8300 and 9000 ft., August 5, 1948 (G. E. Ball), August 12, 1940 (P. A. Timberlake), August 25 and 26, 1952 (B. Malkin) and August 27, 1958 (W. R. Bowen); 1 specimen from Chiricahua National Monument, September 27, 1960 (D. C. Heninger); 1 specimen from Douglas, October 15, 1944 (R. A. Martin); 73 specimens from Chiricahua Mountains, August 1961 (W. Nutting), August 22, 1916 (V. Owen), August 28,1954 (R. M. Bohart), August 28, 1961 (W. Nutting, R. M. Bohart), August 29, 1954 (R. M. Bohart), September 9, 1916, September 14, 1938 and 19 and 29, 1947 (D. J. and J. N. Knull), September 15, 1935 (F. H. Parker), September 20, 1949 (F. H. Parker), September 28, 1960 (D. C. Heninger), and October 14, 1935 (0. Bryant). Paratypes in the collections of the American Museum of Natural History, California Academy of Sciences, Los Angeles County Museum, University of California at Riverside, Arizona State University, University of Washington, University of Idaho, F. H. Parker, D. C. Heninger, W. H. Nutting, B. Tilden, G. H. Nelson and W. F. Barr. This species which appears to be restricted to the Chiricahua Mountains area of southeastern Arizona is most closely related to A. amabilis Horn which is not known from these mountains. These species can be most readily recognized by their different elytral markings. On the other hand, A. chiricahuae closely resembles A. jlavomarginata Gray, under which name it is found in many collections. However, these species are separated on the basis of several characters. In A. chiricahuae the red elytral fasciae usually are interrupted at the middle, the marginal yellow stripe of the elytra is usually distinctly expanded medially, a well-developed subapical plate is present on the last abdominal sternite, the submarginal yellow marking of the pronotum extends only to slightly in front of the middle, and the punctuation of the

5 New species of North American Acmaeodera 149 pronotal disk is more or less uniform. In A. Javomarginata the red elytral fasciae usually are joined at the middle, the marginal yellow stripe of the elytra is very slightly expanded, the last abdominal sternite lacks a subapical plate, the submarginal yellow marking of the pronotum extends to or nearly to the anterior angles, and the punctures of the median depression of the pronotum are more densely placed than on the surrounding areas of the disk. Specimens of A. chiricahuae have been collected from the flowers of Verbesina enceloides, Artemia sp., Cirsium sp., Heliopsis parvijora, and Heterotheca subaxillaris. Acmaeodera inopinata Barr, new species (Figure 2) Male: Medium-sized, rather robust, subflattened, shining blackishbronze; elytra with small reddish spots on third interstrial space in front of basal fourth, in front of middle, at about apical third and behind apical fourth, with small elongate yellow spots on seventh interstrial space behind basal third and at apical third, with small yellow spots on ninth interstrial space in front of basal fourth and behind apical fourth, and with a tiny reddish spot at apex, median reddish spot on third interstrial space transverse and margined with yellow internally and externally; head and pronotum densely clothed with long, erect brown hairs that are sparsely intermixed with white hairs; elytra moderately clothed with long, suberect brown hairs that are sparsely intermixed with white hairs; ventral surface moderately clothed with suberect, rather short white hairs that become denser and longer at sides of thoracic sternites and abdomen. Head rather finely, densely punctured; vertex with a distinct, longitudinal median carina; front transversely convex, flattened on upper portion at middle and transversely depressed above clypeus; clypeus finely, densely punctured, roughened, front margin very broadly and deeply, arcuately emarginate; antenna feebly shining, extending to about middle of pronotum, serrate from fifth segment, outer segments slightly wider than long. Pronotum slightly more than one and three-fourths times wider than long, very slightly narrower than elytra, moderately convex with a shallow median depression at base, an indistinct median groove in front of middle and a strongly indicated, broad depression in front of the distinct basal pits, transverse depression behind front margin feebly indicated on either side of middle; sides shallowly arcuately rounded, indistinctly widened at about basal third; lateral margins narrowly reflexed on about anterior third; front angles feebly produced, subrectangular; front margin broadly, shallowly lobed at middle; hind margin broadly, shallowly emar-

6 150 William F. Barr ginate; surface finely, very sparsely punctured medially, abruptly becoming coarsely and densely punctured towards sides. Elytra feebly convex except for a shallow median depression at base, subequal in width at base and at middle; sides more or less vertical on posterior half, when viewed from above they are very slightly and broadly constricted at about basal fourth and gradually arcuately converging from middle to the conjointly, rather narrowly rounded apices; front margin smooth at middle, not beaded; front angles in side view produced ventrally, acute; lateral margins when viewed from the side feebly sinuate on basal fourth then very broadly and shallowly arcuate to apices, moderately serrate on about apical two-thirds; stria1 punctures rather coarse, deep and circular near suture becoming slightly larger laterally, striae feebly impressed on apical third; interstrial spaces subflattened on disk, each with a row of tiny, sparsely placed punctures, interstrial spaces transversely ridged laterally and apically, distinctly roughened, third interstrial space elevated, seventh interstrial space feebly elevated, ninth interstrial space feebly elevated at base, suture strongly elevated behind middle. Ventral suflace coarsely, rather densely punctured at sides of thoracic sternites and on basal half of first abdominal sternite, sparsely punctured medially, finely, sparsely punctured on inner half of metacoxa and abdominal sternites two to five; front margin of prosternum distinctly retracted, emarginate, not reflexed, with a small rounded tooth on either side of middle; last visible abdominal sternite with sides not conspicuously reflexed, hind margin broadly rounded, disk distinctly convex with a large, densely punctured, subapical elevation that is narrowly rounded and distinctly undercut at apex and which extends slightly beyond hind margin of sternite. Length: 8.7 mm. Width: 3.2 mm. Holotype male (California Academy of Sciences) from Tanbark Flat, Los Angeles County, California, July 12, 1950 (M. F. McClay). Thirteen paratypes from same locality June 20 and 28 and July 8 and 12, 1950 (A.T. McClay), June 23,1950 (W. 0. Marshall), July 1 and 2,1950 (P. D. Hurd), July 2, 1950 (E. G. Linsley), July 4, 1950 (T. R. Haig), July 11, 1952 (J. K. Hester), July 12 and 14, 1950 (M. F. McClay), and July 12, 1950 (H. L. Hansen). Additional paratypes as follows: 81 specimens from Boulevard, San Diego County, California, May 14, 1962, on Haplopappus pingolius (D. S. Verity); 3 specimens from 3 miles east of Etiwanda, San Bernardino County, California, March 31, 1963, reared from Haplopappus pingolius (G. H. Nelson); 24 specimens from Etiwanda, San Bernardino County, California, May 14, 1961, on flowers of Salvia S. Verity); 1 specimen from 4 miles northwest of Cajon Junction, San Bernardino County, California, July 4, 1950 (E. I.

7 New species of North American Acmaeodera 151 Schlinger); 3 specimens from Camp Baldy, Los Angeles County, California, July 7, 1952 (A. T. McClay) and July 14, 1950 (J. W. MacSwain); 1 specimen from Crystal Lake, Los Angeles County, California, June 29, 1950 (5. C. Hall); 1 specimen from Westwood Hills, Los Angeles County, California, May 27, 1934 (G. F. Smith); 15 specimens from 5 miles south of Simi, Ventura County, California, May 4, 1959 (D. L. Mays); 1 specimen from Anza, California, June 28, 1949 (W. E. Simonds); 1 specimen from Santa.Paula, California, June 15, 1957 (W. E. Simonds). Paratypes in the collections of the American Museum of Natural History, California Academy of Sciences, University of California, California Department of Agriculture, G. H. Nelson, D. S. Verity, R. L. Westcott and W. F. Barr. This species is related to and overlaps the distributional range of A. fenyesi Fall. A..inopinata can be recognized by the distinct bronze coloration of the elytra, the more numerous and larger elytral markings and by the sparsely punctured pronotal disk. Acmaeodera deviata Barr, new species (Figure 3) Male: Medium-sized, slender, subcylindrical, shining black; elytra with a tiny yellow spot on third interstrial space, slightly in front of middle; head and pronotum rather densely clothed with long brown hairs that are replaced by irregular white hairs above clypeus and on posterior half of pronotum, elytra sparsely clothed with long, erect brown hairs, rather densely intermixed with irregular, suberect white hairs; ventral surface moderately clothed with suberect white hairs that become dense at sides of thoracic sternites and abdomen. Head rather finely, densely punctured; vertex with a faintly indicated, small, median longitudinal carina; front convex except subflattened at middle and moderately, transversely depressed above clypeus; clypeus finely, densely punctured, roughened, front margin very broadly and deeply, arcuately emarginate; antenna feebly shining, extending to middle of pronotum, serrate from fifth segment, outer segments slightly wider than long. Pronotum nearly twice as wide as long, widest slightly behind middle, subequal in width to elytra, strongly conkex except for a small median depression at base, and a very shallow, feebly indicated depression in front of the distinct basal pits, transverse groove behind front margin not evident; sides arcuately rounded; lateral margins narrowly reflexed throughout, visible from above on anterior two-thirds; front angles subrectangular; front margin very broadly arcuate, not distinctly lobed; disk sparsely, rather coarsely punctured, punctu-

8 152 William F. Barr res becoming coarser and denser toward sides. Elytra feebly convex except for a shallow, transverse median depression at base; sides more or less vertical on posterior half, when viewed from above they are very slightly expanded immediately behind base, then subparallel to middle where they are somewhat arcuately narrowed to the broadly conjointly rounded apices; front singles from side view slightly produced ventrally, acute; lateral margins when viewed from the side broadly, rather deeply, arcuately emarginate from about basal foulth, then very broadly and shallowly arcuate to apices, coarsely, acutely serrate on apical half; stria1 punctures somewhat elongate, deep and rather coarse near suture, becoming more circular and coarser laterally and towards base, striae slightly impressed on apical half; interstrial spaces subflattened on disk, each with a row of tiny distinct punctures, lateral interstrial spaces subserrate apically, third interstrial space elevated, ninth interstrial space strongly elevated at base and then feebly elevated to about middle. Ventral surface coarsely, rather densely punctured at sides of thoracic sternites and first abdominal sternite, more sparsely punctured medially, finely, sparsely punctured on abdominal sternites two to five; front margin of prosternum slightly retracted, shallowly, broadly arcuately emarginate, with a small lobe on either side of middle; last visible abdominal sternite with sides not conspicuously reflexed, hind margim very broadly rounded, disk feebly convex with a conspicuous, sparsely punctured, subapical elevation that is broadly rounded and distinctly undercut at apex. Length: 9 mm. Width: 3.2 mrn. Holotype male (California Academy of Sciences) and 22 paratypes from Desert Springs, San Bernardino County, California, June 11, 1962, on Salvia dorii (G. H. Nelson). Eleven paratypes from Cajon Pass, San Bernardino County, California, May 27, 1962 (D. S. Verity) and June 11, 1962 (G. H. Nelson); 1 paratype from Quail Guzzler, Joshua Tree National Monument, California, June 10, 1966 (E. L. Sleeper and S. L. Jenkins); 1 paratype from Littlerock, Los Angeles County, California, June 17, 1967, on Encelia flower (D. S. Verity); 2 paratypes from Palmdale, Los Angeles County, California, emerged from Juniperus californicus May 20, 1971 (D. E. Rich); 1 paratype from Pasadena, California; 1 paratype from Highway 74, Santa Rosa Mountains, Riverside County, California, 3000 ft., June 12, 1965; and 13 paratypes from Pinyon Flats, San Jacinto Mountains, California, June 13, 1964 and June 18 and 21, 1962 on flowers of Cactus, Salvia apiana and Sphaeralcea ambigua (G. H. Nelson). Paratypes in the collections of the California Academy of Sciences, G. H. Nelson, D. S. Verity and W. F. Barr.

9 New species of North American Acmaeodera 153 This species is close to A. perlanosa Timberlake in appearance and structure. It is most readily recognized by the distinctively elevated ninth interstrial space of the elytra and by the shining black body. A. deviata also tends to be more cylindrical, the body is less pubescent and the elytral striae larger and deeper than with that species. A. deviata does present an unusual variation in markings. Most specimens examined have one or two pairs of small yellowish spots at about the middle of the elytra, some specimens are immaculate, but other specimens may have up to seven pairs of small spots scattered over the elytral surface. Occasionally two spots at the middle of each elytron are joined. Females show a greater degree of maculation than do the males. Acmaeodera verecunda Barr, new species (Figure 4) Male: Medium-sized, convex, shining brown with a purplish-bronze luster; elytra with a rounded, yellowish spot at middle that extends from lateral margin to fourth interstrial space and which is faintly tinged with red posteriorly, with a smaller red spot on lateral margin at about apical third and with a short red stripe on lateral margin in front of apex, right elytron with a tiny yellow spot on lateral margin at about basal fourth and a small red and yellow discal spot at apical third; upper surface rather densely clothed with moderately long, erect brown hairs, that are sparsely intermixed with fine silvery hairs on posterior half of pronotum and on entire elytra; ventral surface moderately clothed with short, suberect, fine silvery hairs which become longer and more densely placed toward sides. Head finely, densely punctured; vertex with a small, distinct, longitudinal median carina; front convex except for a slight median depression and a distinct transverse depression above clypeus; clypeus finely, densely punctured, roughened, front margin deeply and broadly, arcuately emarginate; antenna feebly shining, extending nearly to base of pronotum, serrate from fifth segment, outer segments distinctly wider than long. Pronotum nearly twice as wide as long, slightly narrower than elytra, convex except for a broad shallow median depression at base, a faint median depression near front margin and a very feebly indicated depression in front of basal pits, transverse depression behind front margin feeebly indicated on either side of middle; sides broadly, arcuately narrowing to front angles, slightly widened behind middle; lateral margins narrowly reflexed on anterior two-thirds, feebly reflexed posteriorly, visible from above on anterior half; front angles slightly produced, more or less acute; front margin broadly lobed at middle; hind

10 154 William F. Barr margin broadly emarginate; disk rather finely, very sparsely punctured, punctures becoming very coarse and dense laterally. Elytra convex except for a shallow median depression at base; sides more or less vertical on posterior half, when viewed from above they are slightly expanded immediately behind base and are broadly but feebly constricted at about basal third, from the slightly broadened middle they are very shallowly, arcuately narrowed to the conjointly, narrowly rounded apices; front angles from side view slightly produced ventrally, acute; lateral margins when viewed from the side broadly and shallowly, arcuately emarginate at about basal fourth and then broadly, feebly sinuate to apices, coarsely, acutely serrate on apical half; stria1 punctures circular and deep, small near suture, becoming elongate apically and coarser laterally, striae impressed behind apical third; interstrial spaces subflattened, each with a row of tiny shallow punctures, transversely roughened near lateral margins, third interstrial space slightly elevated, ninth interstrial space feebly elevated on basal fourth, suture strongly elevated on apical fourth. Ventral surface coarsely, rather densely punctured at sides of thoracic sternites and first abdominal sternite, more sparsely punctured at middle, metacoxa more finely punctured, very finely, shallowly punctured on abdominal sternites two to five; front margin of posternum distinctly retracted, not reflexed, with a very broad, slightly projectad subrectangular median lobe; last visible abdominal sternite subtriangular, broadly rounded at apex, sides distinctly reflexed, disk convex with a very large, densely punctured subapical elevation that is broadly rounded at apex. Length: 10.1 mm. Width: 3.8 mm. Holotype male (California Academy of Sciences) and 5 paratypes from Sunset Valley, Santa Barbara County, California, July 4, 1939, on Arctostaphylos glauca (W. F. Barr). Additional paratypes designated as follows: 7 specimens from type locality July 4, 1939 (B. E. White, E. C. VanDyke), July 12, 1941 (F. M. and V. S. Beer) and July 14, 1938 (M. A. Cazier); 1 specimen from Figueroa Mountain Guard Station, Santa Barbara County, California, July 6, 1939 (W. F. Barr); 1 specimen from Figueroa Mountain, Santa Barbara County, California, July 6, 1966 (C. W. Kirkwood); 9 specimens from Tanbark Flat, Los Angeles County, California, June 25, 1956 (H. W. Michalk), June 26, 1952 (S. Migagawa), July 9 and 14, 1956, on Lotus sp. (H. R. Moffitt), July 11, 1952 (A. R. Maggenti, B. Tinglof), July 16, 1956 (R. C. Bechtel); 1 specimen from San Gabriel Mountains, California; 1 specimen from Big Rock Creek, San Gabriel Mountains, Los Angeles County, California, August 11, 1962 (R. L. Westcott); 1 specimen from Mount Wilson, Los Angles County, California, August 3, 1941 (F. M. and

11 New species of North American Acmaeodera 155 V. S. Beer); 1 specimen from Highway 99, near Gorman, Los Angeles County, California, June 22, 1958 (W. E. Simonds); 1 specimen from Arroyo Seco, Monterey County, California, May 3, 1964 (W. E. Ferguson); 3 specimens from Idyllwild, San Jacinto Mountains, Riverside County, California, June 19, 1940, June 22, 1941 and June 28, 1928; 1 specimen from Keen Camp, Riverside County, California, June 6-12, 1917 (E. P. Van Duzee); 1 specimen from Herkey Creek, San Jacinto Mountains, California, June 4, 1940, on Arctostaphylos sp.; 2 specimens from 2.3 miles south of Camp Angeles, San Bernardino County, California, June 29, 1957 (R. L. Westcott); 5 specimens from 3 miles south of Camp Angeles, San Bernardino County, California, 5500 ft., July 6, 1960 (W. F. Barr) and August 6, 1955; 1 specimen from 4 miles south of Fallsvale, San Bernardino County, California, July 3, 1955; 3 specimens from lower Cajon Pass, California, July 10, 1949; 2 specimens from Santa Paula, California, March 21, Paratypes in the collections of the American Museum of Natural History, California Academy of Sciences, California Department of Agriculture, University of California at Riverside, F. M. Beer, G. H. Nelson, M. W. Stone, R. L. Westcott, D. S. Verity and W. F. Barr. A. verecunda can be found in many collections under the name A. plagiaticauda Horn, a species to which it is closely related. Although similar in form and strucuture, these two species have several distinctive anatomical features and different distributional ranges. The elytral markings of A. verecunda consist of spots that are usually restricted to the lateral margins and are, in part, bicolored. When discal spots are present, they are small and usually yellowish. This species also tends to have the pronotum darker and less shining than the purplish elytra and has the pubescence of the ventral surface rather long and silvery. The elytral markings of A. plagiaticauda are reddish and usually extensively distributed as irregular spots over the entire surface. The pronotum of this species usually has the same coloration as the elytra and the pubescence of the ventral surface is short and dark. A. verecunda appears to be restricted to the mountainous areas of southwestern California whereas A. plagiaticauda is known from the northern Coast Range and the Sierra Nevada Range of California and mountainous areas of southcentral Oregon. Acmaeodera knullorum Barr, new species (Figure 5) Male: Medium-sized, robust, convex, shining blackish-brown with a brassy luster; elytra with a lateral and'a discal row of rather large, red and

12 156 William F. Burr yellow spots as follows: a small reddish spot behind humerus on third interstrial space, a tiny yellow spot on lateral margin at about basal fourth, a large, rounded, yellow sublateral spot that is internally reddish and located slightly in front of middle, a circular, sublateral reddish spot at about apical third and a smaller, circular, sublateral reddish spot behind apical fourth, each of the sublateral spots associated with a yellow discal spot which, except for the tiny basal spot, is large, transverse and slightly projected anteriorly, in most instances these spots are joined with the sublateral spots, the anterior discal spot is absent from the right elytron; upper surface moderately clothed with long, erect brownish hairs that are intermixed with fine silvery hairs, on elytra the silvery hairs are more pronounced, but somewhat depressed toward the sides and apices; ventral surface moderately clothed with fine silvery hairs that become denser on sides of meso- and metasternites. Head rather finely, densely punctured; vertex with a small, longitudinal median carina; front convex except for a slight median depression and a shallow, transverse depressed area above clypeus; clypeus finely, densely ounctured, roughened, front margin deeply and broadly arcuately emarginate; antenna feebly shining, extending nearly to base of pronotum, serrate from Mth segment, outer segments distinctly wider than long. Pronotum nearly twice as wide as long, slightly narrower than elytra, moderately convex except for a faintly indicated, shallow median depression at base and a faintly indicated, median longitudinal groove, basal pits distinct, without an associated anterior depression, transverse depression behind front margin distinct on either side of middle; sides nearly straight, widest at base, narrowing to front angles; lateral margins narrowly reflexed on anterior third, feebly reflexed posteriorly, visible from above only near front angles; front angles not produced, narrowly rounded; front margin slightly but broadly lobed at middle; hind margin slightly emarginate; disk very finely, sparsely punctured, punctures becoming coarser and more dense laterally. Elytra convex with a faintly indicated median depression at base; sides more or less vertical on posterior half, when viewed from above they are very slightly expanded immediately behind base and about at middle, then broadly, feebly arcuately narrowing to the conjointly, rather broadly rounded apices; front margin smooth at middle, not beaded; front angles in side view slightly produced ventrally, acute; lateral margins when viewed from the side feebly sinuate to basal fourth, slightly emarginate at about basal fourth, then very broadly and feebly sinuate to apices, coarsely and acutely serrate on apical half; stria1 punctures somewhat elongate, shallow and small near suture, becoming coarser, more circular and deeper at base and near sides, striae impressed behind middle; interstrial spaces subconvex, each with a row of tiny, shallow

13 New species of North American Acmaeodera 157 indistinct punctures, transversely ridged or roughened near lateral margins, third interstrial space slightly elevated, ninth interstrial space elevated on basal fourth, suture strongly elevated on apical third. Ventral surface coarsely, rather densely punctured at sides of thoracic sternites and first abdominal sternite, more sparsely punctured at middle, very finely, shallowly punctured on abdominal sternites two to five; front margin of prosternum slightly retracted, very slightly reflexed with a small rounded tooth on either side of middle and broadly, shallowly, arcuately emarginate at middle; last visible abdominal sternite with sides rather strongly reflexed on basal half, hind margin broadly rounded, disk convex with a very large, densely punctured subapical elevation that is broadly rounded and slightly undercut at apex. Length: 8.7 mm. Width: 3.3 mm. Holotype male (California Academy of Sciences) and 12 paratypes from Pinon Flat, San Jacinto Mountains,California, July 7, 1960, on Acacia blossoms (W. F. Barr).. Additional paratypes designated as follows: 30 specimens from Pinyon Flats, Highway 74, Riverside County, California, 4000 ft., June 15 and July 7 and 24, 1960, June 16, 1959, June 18, 1962 and July 8, 1964, on Eriogonum fasciculatum, Acacia greggii, Sphaeralcea ambigua blossoms, Purshia glandulosa and Quercus dumosa (G. H. Nelson); 5 specimens from above Palm Desert, Highway 74, Riverside County, California, 2500 ft., June 15, 1960, on Acacia greggii and June 22, 1964, on Quercus dumosa and Eriogonum fascicuiatum (G. H. Nelson); 20 specimens from 3 miles east of Banner, San Diego County, California, May 24, 1962 and June 20, 1960, on Quercus dumosa and Eriogonum fascicuiatum (G. H. Nelson); 2 specimens from Jacumba, San Diego County, California, July 2,1961, on Acacia greggii (G. H. Nelson); 2 specimens from 5 miles east of Jacumba, San Diego County, California, July 1960, on Acacia greggii (W. F. Barr). Paratypes in the collections of the California Academy of Sciences, G. H. Nelson and W. F. Barr. A. knullorum, because of its variable nature, is separated from the closely related A. holsteni White with some difficulty. In general, however, it can be recognized by the usually straight sided pronotum, by the presence of discal or lateral spots on the anterior half of the elytra, by the basally elevated ninth interstrial spaces of the elytra, by the occasional presence of a lateral spot on the pronotum and by the median longitudinal depression on the pronotum. Furthermore, this spcies seems to be a flower visitor and perhaps has several larval host plants as may be inferred from the collection data.

14 158 William F. Burr Adults of A. holsteni, on the other hand, do not visit flowers and the only known larval host of this species is Adenostoma fasciculatum. Variation in elytral markings, shape of the pronotum and in overall length has been noted in the paratypic series. The elytral spots may be considerably reduced in size and the anterior discal spots may be absent. The sides of the pronotum when viewed from above usually are straight but in some specimens they show an arcuate condition which is typical of A. holsteni, but in others they are very slightly emarginate. The median depression of the pronotum occasionally is evident only as a subflattened area. Finally, the paratypes have been found to vary from 6.5 to 11 mm. in length. This species is dedicated to Dorothy J. and Josef N. Knull of Ohio State University for their many contributions to taxonomy of North American beetles. Acmaeodera nevadica Barr, new species (Figure 6) Male: Medium-sized, moderately elongate; head, pronotum and ventral surface bronzy-black; pronotum with an elongate, sublateral yellow spot at base; elytra blue-black with lateral margins narrowly yellow on basal third and with irregular transverse yellow markings arranged as follows: a subbasal band extending from yellow lateral margin to disk where it branches obliquely anteriorly and posteriorly, a median irregular band extending from lateral margin obliquely towards posterior branch of subbasal marking, a small spot on lateral margin behind middle, a short oblique band at apical third, a short oblique band behind apical fourth, and a small subbasal spot; upper surface clothed with long, erect brownish hairs which are denser on head and pronotum, lateral areas of elytra and clypeus with a few suberect silvery hairs; ventral surface moderately clothed with suberect, fine silvery hairs. Head rather finely, densely punctured; vertex with a well-developed median longitudinal carina; front feebly depressed at middle and above clypeus; clypeus finely punctured, front margin rather deeply, broadly subtriangularly emarginate; antenna shining, extending to beyond middle of pronotum, serrate from fifth segment, outer segments distinctly wider than long; inner margin of eyes slightly elevated above head surface. Pronotum twice as wide as long, widest at base, slightly narrower than elytra, moderately convex except for a broad, shallow median depression in front of base, a faintly indicated median groove on anterior half, and a broad, shallow oblique depression in front of basal pits, transverse depression behind front margin distinct on either side of middle; sides feebly arcuate, narrowing towards front angles;

15 New species of North American Acmaeodera 159 lateral margins slightly reflexed on about anterior third, carinate posteriorly, visible from above only near front; front angles slightly produced, acute; front margin broadly and shallowly lobed at middle; hind margin very shallowly emarginate; surface moderately punctured at middle, punctures subequal to those on head, becoming coarser and denser toward sides. Elytra feebly convex except for a shallow but distinct median depression at base; sides nearly vertical on apical half, when viewed from above they gradually narrow from base to apical third and then more abruptly narrow to the conjointly, broadly rounded apices; front margin beaded; front angles in side view slightly produced ventrally, acute; lateral margins when viewed from the side feebly bisinuate on about basal fourth, then broadly arcuate to apices, coarsely, sparsely serrate on apical third; stria1 punctures deep and rather small, coarser toward sides, elongate behind middle where the striae become impressed; interstrial spaces flattened on disk, convex and distinctly punctured behind middle, third interstrial space elevated, ninth interstrial space elevated at base, suture strongly elevated on apical half. Ventral surface coarsely, densely punctured at sides of thoracic sternites and on basal half of first abdominal sternite, sparsely punctured medially, metacoxa more finely punctured, remainder of abdomen finely, densely punctured; front margin of prosternum distinctly retracted, narrowly reflexed on either side of middle, subtruncate; last visible abdominal sternite with sides very feebly arcuate, distinctly reflexed, hind margin nearly truncate, disk convex with a small, transverse subapical plate that is smooth and slightly undercut at apex. Length: 9.7 mm. Width: 3.3 mm. Holotype male (California Academy of Sciences) from Kyle Canyon, Clark County, Nevada, June 26,1966,011 Cowania flower (W. F. Barr); 37 paratypes with same collection data, except some specimens on Senecio flowers, collected by E. J. Allen, W. F. Barr, R. L. Penrose and R. L. Westcott and, from the same locality, 6 paratypes June 5, 1941 (E. C. Van Dyke, P. H. Timberlake) and 3 paratypes June 17, 1967, on Sphaeralcea (W. F. Barr). One paratype from Highland Range, above Mendha, Lincoln County, Nevada, June 25, 1966 (R. L. Penrose). Paratypes in the collections of the California Academy of Sciences,University of California at Riverside, University of Idaho, R. L. Westcott and W. F. Barr. This species appears to be a geographical replacement of the widely ranging and common Pacific Coast species A. liepburni LeConte. A. nevadica, which occurs under more arid conditions in certain areas of southern

16 160 William F. Burr Nevada, can be recognized by its darker coloration, the distinctive elytral markings and the small subapical plate of the last abdominal sternite. Acmaeodera vernalis Barr, new species (Figure 7) Male: Medium-sized, somewhat robust, feebly convex, shining, dark brown with a purplish-bronze luster; pronotum with a small lateral yell* wish spot in front of hind angles; elytra with pale yellow markings as follows: a marginal maculation extending from behind front angle to basal fourth which is internally expanded behind humerus, an irregularly transverse band extending from lateral margin to ninth interstrial space immediately in front of middle, a transverse band extending from lateral margin to seventh interstrial space at about apical third, an elongate spot on lateral margin behind apical fourth, a moderate-sized spot on eighth and ninth interstrial spaces immediately behind base and a small spot on eighth and a small spot on ninth interstrial spaces behind apical fourth; upper surface moderately clothed with rather long, erect brown hairs; ventral surface rather sparsely and inconspicuously clothed with suberect, fine white hairs. Head finely and densely punctured; vertex with a faintly indicated, longitudinal median carina; front transversely convex, broadly but shallowly concave on upper portion at middle, transversely depressed above clypeus; clypeus finely, densely punctured, front margin very broadly and shallowly, arcuately emarginate; antenna feebly shining, extending nearly to base of pronotum, serrate from fifth segment, outer segments nearly twice as wide as long. Pronotum twice as wide as long, slightly wider than elytra, moderately convex with a broad, shallow median depression at base, a faintlyindicated median groove in front of middle and a very shallow, broad depression in front of basal pits, transverse depression behind front margin moderately distinct on either side of middle; sides arcuately rounded, widest at about basal fourth; lateral margins narrowly reflexed except at basal fourth, visible from above on anterior half; front angles distinctly produced, subacute; front margin broadly lobed at middle; hind margin feebly emarginate; surface sparsely punctured at middle, punctures moderate in size becoming coarser and denser toward sides. Elytra feebly convex except for a small, shallow median depression at base; sides nearly vertical on apical half, when viewed from above they gradually narrow from base to middle except for a broad shallow constriction at about basal fourth, from middle they more abruptly and arcuately narrow to the conjointly narrowly rounded apices; front margin strongly beaded at middle; front angles in side view feebly produced ventrally, sub-

17 New species of North American Acmaeodera 161 rectangular; lateral margins when viewed from the side broadly, shallowly sinuate on basal fourth, broadly but slightly lobed at basal fifth, broadly, shallowly arcuately emarginate at basal fourth, then broadly arcuate to apices, distinctly serrate from in front of middle; stria1 punctures rather coarse, deep and somewhat elongate near suture, becoming more circular and slightly larger toward sides, striae not impressed; interstrial spaces flattened on disk, subconvex elsewhere, somewhat transversely roughenend near apices, interstrial punctures indistinct, sparsely placed, third interstrial space distinctly elevated, seventh interstrial space feebly elevated, ninth interstrial space feebly elevated to about basal fourth, suture strongly elevated on apical third. Ventral surface coarsely, rather sparsely punctured at sides of thoracic sternites, becoming sparsely punctured towards middle, metacoxa more finely punctured, abdominal sternites finely, sparsely punctured; front margin of prosterum distinctly retracted, narrowly reflexed, very broadly and shallowinly bisinuate; last visible abdominal sternite with sides narrowly reflexed, sublinear, hind margin subtruncate, disk convex with a conspicuous, densely punctured, broad subapical plate that is distinctly undercut and subtruncate at apex. Length: 8.3 mm. Width: 3.1 mm. Holotype male (California Academy of Sciences) and 26 paratypes from Tahquitz Canyon, Palm Springs, Riverside County, California, April 21, 1964, on Encelia flowers (W. F. Barr). An additional 114 paratypes from type locality March 25, 1968 (R. L. Westcott), April 13, 1958, on Encelia farinosa (G. H. Nelson), April 22, 1963 (F. D. Parker and L. A. Stange) and April 16, Other paratypes designated as follows: 25 specimens from San Andreas Canyon, Palm Springs, Riverside County, California, March 2, 1934, March 8, 1956 and March 31, 1948, on Encelia farinosa (P. H. Timberlake), March 11 and 31, 1955 (W. R. M. Mason), April 21, 1951 (R. C. Bechtel and E. I. Schlinger), April 23, 1933, on Encelia farinosa (P. H. Timberlake), May 7, 1958, on Encelia farinosa (G. H. Nelson), and May 15, 1932, on Eriodictyon crassifolium (P. H. Timberlake); 70 specimens from Palm Springs, Riverside County, California, March 15, 1917, March 17 and 23, 1957 (R. L. Westcott), March 23, 1940 (B. E. White), March 26, 1926, March 29 (F. C. Clark), March 30, 1945, on Encelia farinosa (Cockerell), March 31, 1948, on Encelia farinosa (P. H. Timberlake), April 3, 4 and 5, 1965, April 11, 1945, on Encelia farinosa (Cockerell), April 12, 1958 (R. L. Westcott), April 21, 1951 (R. C. Bechtel, E. I. Schlinger), April 22, 1945 (J. A. Comstock), April 28, 1962 (G. H. Nelson). Paratypes in the collections of the California Academy of Sciences, Los Angeles County Museum, University of California at Davis and Riverside, Canada National Collection, H. F. Howden, G. H. Nelson, R. C. Bechtel, R. L. Westcott and W. F. Barr. a

18 162 William F. Barr This species occurs commonly throughout the Colorado Desert of southeastern California and adjacent areas. Spcimens not designated as paratypes have been seen from: Imperial County, California: Bertram Siding; Kane Springs. Riverside County, California: 1 mile south of Palm Desert; 2 miles south of Desert Center; Desert Center; Coyote Creek; Riverside; 4 miles south of Riverside; Gilman Hot Springs; White Water; Soboba Springs; Thousand Palms; Box Springs Mountains. San Diego County, California: Vallecito; Borrego ; Palm Canyon, Borrego, on Viguiera parishii: 3 miles west of Ocotillo; 2 miles west of the Narrows. Baja California: Arroyo Grande, 39 miles southwest of La Ventana. The dark brown coloration, the moderate sized discal punctures of the pronotum and the feebly elevated elytral interstrial spaces distinguishes this species from A. jlavosticta Horn. Acmaeodera depressa Barr, new species (Figure 8) Male: Medium-sized, elongate, subtriangular, black with a purplish- -bronze tinge; elytra with three rows of small yellow spots, first row on the lateral margin consisting of two spots, one very small at about basal fourth and the other slightly behind middle, second row on the third interstrial space and consisting of four spots, the first immediately behind humerus, the second immediately in front of middle, the third at about apical fourth, and the last, tiny in size, at about apical fifth, the third row touching the seventh interstrial space and consisting of four spots, the first in front of basal fifth, the second at about basal two-fifths, the third at about apical two-fifths and the fourth at about apical fifth; head and pronotum moderately clothed with rather long, erect brownish hairs, sparsely intermixed with white hairs, elytra sparsely clothed with erect brownish hairs and a few irregular and suberect, fine white hairs that become more densely placed and subrecumbent near lateral margins; ventral surface dark bronze with a purplish luster, moderately clothed with short to long, suberect silvery hairs. Head rather coarsely, densely punctured; vertex with a distinct longitudinal median carina; front transversely convex with a very shallow median depression on upper portion, area above clypeus feebly, transversely depressed; clypeus finely, densely punctured, somewhat roughened, front margin shallowly, broadly arcuately emarginate; antenna feebly shining, extending to about middle of pronotum, serrate from fifth segment, outer segments distinctly broader than long; inner margin of eyes slightly elevated above head surface. Pronotum approximately twice as wide as long, about equal in width to elytra,

19 New species of North American Acmaeodera 163 convex with a distinct median depression at base, a very feebly indicated longitudinal depression on disk and an indistinct shallow depression in front of the distinct lateral pits, transverse depression behind front margin feebly indicated on either side of middle; sides slightly expanded in front of base, broadly arcuately narrowing to front angles; lateral margins narrowly reflexed, visible from above except at extreme base; front angles produced, subacute; front margin broadly lobed at middle; hind margin broadly shallowly emarginate; surface somewhat shallowly and coarsely, sparsely punctured at middle, punctures becoming more coarse and denser laterally. Elytra subdepressed except for a transverse, shallow median depression at base; sides more or less vertical on posterior half, when viewed from above they are very slighty expanded immediately behind base and at about middle, then gradually arcuately converging to the conjointly, narrowly rounded apices; front margin distinctly beaded; front angles from side view subrectangular; lateral margins when viewed from the side feebly bisinuate to about basal fourth, then very broadly and shallowly, arcuately rounded to apices, coarsely and sparsely serrate from slightly in front of middle; stria1 punctures small, circular and deep near suture, becoming more coarse toward sides, striae feebly impressed behind apical third; interstrial spaces flattened, tiny interstrial punctures indistinct, third interstrial space slightly elevated, seventh interstrial space feebly elevated, ninth interstrial space distinctly elevated at extreme base, suture strongly elevated on apical half. Ventral surface rather coarsely and densely punctured on thoracic sternites and basal half of first abdominal sternite, metacoxa more finely punctured, very finely, sparsely punctured on apical half of first abdominal sternite and abdominal sternites two to four; front margin of prosternum distinctly retracted, slightly reflexed, very broadly and shallowly bisinuate; last visible abdominal sternite with sides feebly arcuate, strongly reflexed, hind margin broadly rounded, subtruncate, disk feebly convex, rather coarsely punctured, with a small, broadened, nearly smooth subrectangular subapical plate. Length: 9.1 mm. Width: 3.1 mm. Holotype male (California Academy of Sciences) and 4 paratypes from Morristown, Maricopa County, Arizona, April 15, 1964 (W. F. Barr). Five paratypes from 3 miles north of Florence, Pinal County, Arizona, April 17, 1964, on Sphaeralcea and Encelia (W. F. Barr, A. R. Gittins); 1 paratype from 13 miles northeast of Florence, Pinal County, Arizona, April 13, 1966; 4 paratypes from Florence Junction, Pinal County, Arizona, April 5, 1954 (T. R. Haig); 1 paratype from 15 miles east of Ehrenberg, Arizona, March 11, 1963, reared from Fouquieria splendens (N. OYConnell);

20 William F. Barr 1 paratype from 24 miles southeast of Wickieup, Yavapai County, Arizona, April 30, 1966, on Melampodium leucanthum (B. Osbeck, J. M. Davidson and M. A. Cazier). Paratypes in the collections of the University of California at Davis, Arizona State University, J. M. Davidson and W. F. Barr. At first glance this species may be taken for A. cuneata Fall. However, the consistancy of the small markings of the elytra which do not show a tendency toward the development of a pair of irregularly elongate subapical stripes, the dark coloration, the presence of white hairs on the elytra and the elongate rather than a cuneate form appear to be distinctive features for species recognition. Acmaeodera pletura Barr, new species (Figure 12) Male: Medium-sized, robust, moderately shining; black, head, pronotum and ventral surface with a bronzy luster, elytra with pale yellow spots and transverse markings arranged as follows: two small discal spots located at about basal forth and in front of middle, a narrow transverse marking extending from near lateral margin to behind umbone, a small spot on lateral margin at about basal fourth, a transverse marking extending from margin in front of middle posteriorly onto disk, a small spot on lateral margin at about apical third, a transverse marking at apical fourth extending anteriorly onto disk, a short transverse marking at about apical sixth extending onto disk, and a small apical spot, anterior discal spot and marking behind umbone joined on right elytron, subapical transverse marking broken on left elytron and joined with apical spot along margin; upper surface moderately clothed with rather long, erect blackish hairs except on area above clypeus which is clothed with shorter white hairs; ventral surface moderately clothed with rather long, fine white hairs. Head finely, densely punctured, punctures more sparsely placed on upper portion of front; vertex with a faintly indicated, longitudinal median carina; front convex except for a slight median depression and a shallow transversely depressed area above clypeus; clypeus punctured medially, roughened laterally, front margin broadly, shallowly, subtriangularly emarginate; antenna faintly shining, extending slightly beyond middle of pronotum, serrate from fifth segment, outer segments wider than long; inner margin of eyes not distinctly elevated above head surface. Pronotum twice as wide as long, widest behind middle, subequal in width to elytra, moderately convex except for a discal subflattening and a broad, shallow oblique depression in front of distinct basal pits; sides arcuate;

21 New species of North American Acmaeodera 165 lateral margins carinate throughout, slightly reflexed on about anterior third where they are visible from above; front angles slightly produced, subacute; front margin distinctly lobed at middle; hind mqigin subtruncate; surface rather finely and sparsely punctured at middle, becoming much more coarsely and densely punctured towards sides; transverse depression behind front margin indistinct on either side of middle. Elytra feebly convex with a transverse median depression at base; sides nearly vertical on apical half, when viewed from above subequal in width behind the base and at middle, slightly expanded immediately behind base, very broadly, shallowly constricted at about basal third, then very broadly, shallowly, arcuately narrowing to the conjointly, broadly rounded apices; front angle in side view slightly produced ventrally, subrectangular; lateral margin when viewed from the side very feebly bisinuate on about basal fourth, then broadly sinuate to apices, coarsely serrate on about apical third; stria1 punctures small and deep, striae deeply impressed behind middle; interstrial spaces subflattened, irregularly roughened or scabrous especially on anterior half and near sides on apical half, interstrial spaces indistinct, small, subequal to stria1 punctures, third interstrial space slightly elevated from behind umbone to near apical fourth, suture elevated from about basal fourth to apices, more strongly elevated apically. Ventral surface coarsely, densely punctured at sides of thoracic sternites and on basal half of first abdominal sternite, sparsely punctured medially, metacoxa more finely punctured, remainder of abdomen finely sparsely punctured; front margin of prosternum distinctly retracted, narrowly reflexed, subtruncate; abdominal sternites 2, 3 and 4 with hind angles slightly prolonged, subacute; last visible abdominal sternite with margins distinctly reflexed, sides very feebly arcuately narro- wing to the broadly rounded hind margin, disk moderately convex, subapica1 elevation absent. Length: 7.8 mm. Width: 2.9 mm. Holotype male (California Academy of Sciences) and 1 paratype from Lee Canyon, Clark County, Nevada, June 27, 1966, on Sphaeralcea (W. F. Barr); 2 paratypes from same locality, July 1, 1966, on Fallugia paradoxa (R. L. Westcott). Additional paratypes from Lower Lee Canyon, Clark County, Nevada, June 17, 1967, on Fallugia paradoxa (W. F. Barr) 2 specimens; Kyle Canyon, Clark County, Nevada, July 6, 1967, on blossoms of Fallugia paradoxa (W. F. Barr) 1 specimen; north rim, Grand Canyon National Park, Arizona, June 15, 1930 (C. Kelley) 1 specimen; south rim of Grand Canyon, Arizona, July 8, 1967, on blossom of Cowania stansburiana (W. F. Barr), 1 specimen. Paratypes in the collections of the California Academy of Sciences, R. L. Westcott and W. F. Barr.

22 166 William F. Barr This robust species appears to be related to A. retifer LeConte. The black coloration, elytral markings and absence of a subapical plate on the last abdominal sternite of A. pletura will distinguish these species. From A. bishopiana Fall which it superficially resembles, A. pletura can be separated by the above features and also by the absence of white hairs on the elytra. Acmaeodera sphaeralceae Barr, new species (Figure 13) Male: Medium-sized, rather robust, subconvex, black with a distinct bronze luster; elytra with irregular yellow markings over entire surface that are somewhat sinuous and longitudinally oriented; head and pronotum densely clothed with long, erect brown hairs that are sparsely intermixed with white hairs, elytra moderately clothed with a mixture of long, suberect brown and white hairs that become subrecumbent along sides; ventral surface moderately clothed with inconspicuous, rather short, suberect white hairs that become more dense and longer at sides of thoracic sternites. Head coarsely and shallowly, rather densely punctured; vertex with a longitudinal median carina, feebly depressed on either side of middle; front with a median longitudinal depression and a deep transverse depression above clypeus; clypeus finely, densely punctured, front margin broadly and deeply, triangularly emarginate; antenna shining, extending to slightly beyond middle of pronotum, serrate from fifth segment which is slightly smaller than following segments, outer segments slightly wider than long; inner margin of eyes slightly elevated above surface of head. Pronotum approximately one and three-fourths times wider than long, feebly widened at about basal third, subequal in width to elytra, moderately convex with a small, shallow median depression at base and a distinct median groove on disk, with a very broad and shallow oblique depression extending from the distinct basal pits, transverse depression behind front margin distinct on either side of middle; sides shallowly, arcuately rounded; lateral margins narrowly reflexed on about anterior half, carinate posteriorly, visible from above on anterior third; front angles slightly produced, narrowly rounded; front margin broadly and shallowly lobed at middle; hind margin subtruncate; surface coarsely, deeply and sparsely punctured at middle, punctures becoming very coarse and dense toward sides. Elytra slightly convex except for a shallow median depression behind the strongly beaded front margin, subequal in width behind base and at middle; sides more or less vertical on posterior half, when viewed from above they are very slightly and broadly constricted at about basal third and then gradually arcuately converging from middle to the

23 New species of North American Acmaeodera 167 conjointly broadly rounded apices; front angles in side view subrectangular; lateral margins when viewed from the side feebly bisinuate on basal fourth then broadly, shallowly arcuate to apices, coarsely serrate on apical half; strial punctures moderate in size, deep and somewhat elongate on disk, becoming slightly larger at base and toward sides, striae slightly impressed on apical third; interstrial spaces flattened near suture, convex elsewhere, roughened at base, subserrate and transversely roughened near sides, interstrial punctures shallow and indistinct, third interstrial space slightly elevated, suture moderately elevated behind middle. Ventral surface coarsely, densely punctured at sides of thoracic sternites, more finely punctured on metacoxa and basal half of first abdominal sternite, sparsely punctured medially, finely, sparsely punctured on apical half of first abdominal sternite and sternites two to five; front margin of prosternum strongly retracted, subtruncate, distinctly reflexed; last visible abdominal sternite with sides distinctly reflexed, hind margin subtruncate at middle, disk convex with a small, transverse, smooth subapical elevation that is undercut at apex. Length: 8.1 mm. Width: 2.8 mm. Holotype male (California Academy of Sciences) and 20 paratypes from Molino Basin, Santa Catalina Mountains, Arizona, April 8, 1960, on Sphaeralcea laxa (K. Roever). Seven paratypes from Catalina Mountains, Arizona, April 15, 1948 (R. S. Beal) and 2 paratypes from 14 miles south of Holbrook, Arizona, June 1, 1962, on Sphaeralcea (G. H. Nelson). Paratypes in the collections of the California Academy of Sciences, Northern Arizona University, G. H. Nelson and W. F. Barr. The elytral markings of A. sphaeralceae are similar to those of some specimens of A. retifer LeConte, whereas in the general form and structure this species resembles A. bishopiana Fall. In any case the longitudinal canna on the vertex and the median longitudinal depression of the pronotum will serve to separate A. sphaeralceae from these species. Acmaeodera ornatoides Barr, new species (Figure 9) Male: Large-sized, robust, black; head and pronotum with a bronzy luster; elytra steel blue except bronzy on basal fourth, with small, irregular yellow spots scattered along sides from behind umbone to behind apical fourth and on posterior half of disk; head and pronotum densely pubescent, elytra moderately clothed with a mixture of erect brown and white hairs;

24 William F. Burr ventral surface moderately but inconspicuously clothed with suberect, short white hairs. Head finely, densely punctured, densely clothed with a mixture of rather long, fine, erect white and brown hairs which become entirely white on clypeus; vertex with a longitudinal carina that extends onto front; front convex except for a shallow median depression and a transverse depression above clypeus; clypeus blue-black, finely punctured laterally, front margin broadly and shallowly, triangularly emarginate; antenna extending to about basal third of pronotum, serrate from fifth segment, outer segments slightly wider than long. Pronotum slightly less than twice as wide as long, widest at base, slightly wider than elytra, convex except for a conspicuous median depression at base, a shallower and narrower median longitudinal depression on disk and a broad oblique sublateral depression extending to anterior third from the conspicuous basal pits, transverse depression behind front margin evident on either side of middle; sides broadly rounded, narrowing to front angles; lateral margins narrowly reflexed except at extreme base, visible from above only at front angles; front angles feebly produced, narrowly rounded; front margin broadly lobed at middle; hind margin broadly emarginate; surface coarsely, densely punctured at middle, becoming somewhat more coarse and dense toward sides, uniformly and densely clothed with rather long, erect brown and a few white hairs especially near base and sides. Elytra subconvex, flattened along suture and shallowly depressed at base; sides nearly vertical on apical half, when viewed from above they are feebly expanded behind base and near middle and then rather strongly, arcuately narrowed to the conjointly and broadly rounded apices; front margin slightly beaded; front angles in side view slightly produced ventrally, narrowly rounded; lateral margins when viewed from the side very feebly bisinuate on basal fourth, rather deeply emarginate at basal fourth, then very broadly arcuate to apices, coarsely serrate on apical half; stria1 punctures coarse and deep on disk becoming more coarse toward base and sides, striae feebly impressed behind middle; interstrial spaces subconvex, distinctly roughened or subserrate apically and laterally, interstrial punctures distinct, third and seventh interstrial spaces elevated throughout, ninth interstrial space elevated on about basal third, suture moderately elevated behind middle. Ventral surface coarsely, rather densely punctured at sides of thoracic sternites and on basal half of first abdominal sternite, more sparsely punctured medially, metacoxa finely, densely punctured, apical half of first abdominal sternite and sternites two to five rather finely and sparsely punctured; front margin of prosternum retracted, very feebly bisinuate, subtruncate, narrowly but distinctly reflexed; last visible abdominal sternite with sides conspicuously reflexed, hind margin broadly nearly semicircularly

25 New species of North American Acmaeodera 169 rounded, disk convex with a broad, conspicuous subapical plate that is nearly smooth, subtruncate and deeply undercut at apex.. Length: 11.6 mm. Width: 4.3 mm. Holotype male (American Museum of Natural History) and 69 paratypes from Weser, Goliad County, Texas, May 11, 1952 (M. Cazier, W. Gertsch and R. Schrammel). Additional paratypes designated as follows: 37 specimens from Garner State Park, Uvalde County, Texas, June 22-24, 1961 (R. L. Westcott); 2 specimens from Reagan Wells, Uvalde County, Texas, May 11, J. and J. N. Knull); 8 specimens from 10 miles north of Kerrville, Texas, May 22, 1953 (R. H. Beamer); 10 specimens from Austin, Texas; 3 specimens from Cedarville, Dallas County, Texas, April 20, 1935 and May 2, 1934; 10 specimens from 15 miles northwest of San Antonio, Texas, June 18, 1940 (B. Struck and D. Orchard); 4 specimens from Salado Creek, Fort Sam Houston, Bexar County, Texas, May 7, 1952 (M. Wasbauer) and May 7, 1956 (S. E. Knapp); 13 specimens from Harper, Texas, May 6, 1954, on Opuntia and Callirhoa (L. D. Beamer). Paratypes in the collections of the American Museum of Natural History, California Academy of Sciences, California Department of Agriculture, Canada National Collection, Cornell University, University of Idaho, Washington State University, F. M. Beer, G. H. Nelson, R. L. Westcott and W. F. Barr. Individual specimens not designated as paratypes have been seen from the following localities: Oklahoma: Wichita National Forest. Texas : Harris County; College Station; Taylor; Brownwood; Fedor; Concan; Sonora; Giddings; and Palmetto State Park, Gonzales. A. ornatoides resembles A. ornata (Fabricius) and can be found with that species in many collections. The two species, however, are distinctive in anatomical features and distributional occurrence. Compared with A. ornata, A. ornatoides is bluish rather than black, the pronotum is always immaculate, the sides of the pronotum are expanded behind the middle rather than in front of the base, the elytra are in part costate, the post humeral yellow spot usually is not elongate or curved and the elytral pubescence is longer and paler in coloration. In general distributional occurrence, A. ornatoides is known from the south central United States and A. ornata from the southeastern United States.

26 'UIW E'P :YIP!M '"W 8.01 :y18ua7 -papun01 Xlpeo~q xade 'su~8~em Ielalel 8uole aae3uo3 LIVU~JS!~ 'qs~p no xaauo:, aqulals len!mopqe ajq!s!a lsel falpppu jo apys layl!a uo paqol Xlqaaj X~aa 'ale3unl)qns 'pal3el)a~ urnulalsold jo u!8leur luolj f slpy Lla~l!s auy '13alaqns 'I.IO~S ~IIM pay2013 dlasleds 'a~g 01 OM) salrulals ~euyu~opqe uo Jany 'sapls sp~e~ol lasnap 'palnl3nnd dl~olleqs 'Llasleo3 a3vj~ns fv~~ua/i.lnoy8nolyl xa~uo:, Lp3ugsyp a3eds IeyJlslalu! p~yq 'Llmoy~alsod paleaala 61qaaj puo K1leseq palmala dl13n!)sg1 a3eds IeplsJalu! ylnahas 'xade leau lda3xa pauauep 'salni3und 13u!ls!p Ilems jo MOJ e yl!~ samds Itzplslalu! f X1je3~de passaldm! aogs 'dlle~ale~ las~eo:, 'alnlns loau ~~eurs Jargel 'daap put! le1n31y3 saznpund je!qs fs~!eq n ~olq y~ep 3313 '8uoj q p paylop ~ dlas~eds aqms f pqyl 1e3!de no alollas d~asleds 'dlasleoci 'sa3ido 01 alen3le d~peolq uayl put! pllyl [sseq lnoqe le aleu!8leura d~~olleys 'laaugqns 'apls ayl molj pama!a uaym sar8lem Ielalel fleln8ue~alqns Mapi aprs u! sal8ue luoy fsacydo papunol Llpoolq dpu!o[uo3 ayl 01 aqaauo3 Alalen3le Lay1 alaym plyyl 1e3!de lnoqe 01 Ial~e~edqns uayl put! aseq puryaq papuedxa 6py8!1s ale day1 aaoqe moy pahap uaym 'le3yna 6p3nys!p IOU sapls fyllnoj pseq puyq alnlns palehala ayl loj lda3xa passaldapqns VJIL~~.sl!ey u~olq y~ep l3aja In01 ql!~ paqlol3 Ljas-leds 'sap!s plemol paqd dlasuap alom 8uymo3aq sampund '~s!p uo pa~npnnd L~asleds put dlasleoa laylel a3gms f alppym le paqoi Llqaaj lnq d~peolq uy8~em luo~j f 'pa3npo~d Lpy8!js sal8ue 1~013 fqnoy8nolyl aaoqe molj alqysp 'aseq amaqxa le ldacixa paxagal dp3ugs;rp sur8lem ~e~apl f plyyl pseq lnoqe le 8uyy31ou 1y8yls e loj lda3xa sapue lno~j 01 uoyuod lsap!m moq ~U!MOJJ~U dlalen3le saprs f aippym jo apys layira no 13uysyp u!%leur luog puyaq uo!ssa~dap as1ahsue.u f slyd leseq urog Llanbqqo %u!paa)xa uo!ssaldap 13ngs1p e pue uo!ssaldap uerpam [eu!pnl!8uo~ '~olloys o loj da~xa xaauo3 'aseq jo luog u! X[ale!pamm! lsap!m 'eq6la ueyl Japy '8nol se ap!m se a3yml ueyl alom d~lqb!js uinlouord.yl%ua1 lyayl 01 ~enbaqns sluam8as wno jo qlpr~ '1uau18as yljy 1.~013 ale~~as 'mnlouold jo a1ppim Inoqe 01 Buypualxa sunalue f uo!leu!8~ema jo aprs layira no pavoo) dpeln8uey.q 'Llpeolq 'alsuy8-1eura dpeln8uep~ Lpeau 'L~Mo~~E~s pne d~peolq u18lem lnolj 'pauaq8no~ Lpeln8a~l! snaddla falpprm le ladaap ')3u!ls!p uorssa~dap 1suua)ue-lap aslahsuell 'uo!ssa.rdap ue!pam ~olpys Xlaa 'peo~q e loj lda3xa xaauo:, luog feu!~a3 uerpaur pnypnl!%nol 'padolaaap X~qaaj e ~IIM xapar\ fsqeq UMOJ~ ~JBP l3a~a '8~01 yl!~ paylola Llasuap 'paml3nnd d~asuap 'Xlaug pva~ qs!p uo slods IIems lnoj jo MOJ ~ep8a~~y ue put! saprs le slods ~ollad IIeurs xrs jo MOI leinball! ue ~JIM e~ll1a 'y3elq 'lsnqol 'paz!s-a8~e~:alvm (0 I a~n%d) sapads ~ a 'lma u sualla alapoaam3v

27 New species of North American Acmaeodera 171 Holotype male (California Academy of Sciences) from Kyle Canyon, Clark County, Nevada, June 26, 1966 on flowers of Cowania stansburiana. Twenty-one paratypes from same locality collected by E. J. Allen, R. L. Penrose, R. L..Westcott and W. F. Barr, with one specimen on wild rose flower. Additional paratypes labeled Kyle Canyon, Charleston Mountains, Clark County, Nevada, June 27, 1969 (J. W. Tilden), 3 specimens; 7200 ft., July 1, 1969 (L. I. Pechuman) 1 specimen; about 7000 ft., July 25, 1932, 1 specimen; and Clark County, Nevada, July 18, 1929, 2 specimens. Paratypes in the collections of the American Museum of Natural History, California Academy of Sciences, Oregon State University, University of Idaho, G. H. Nelson, R. L. Westcott and W. F. Barr. This localized spcies closely resembles large dark specimens of A. knowltoni Barr, especially those from localities in the southern portion of the range of that species. Perhaps A. errans will eventually prove to be only subspecifically distinctive. However, for the present its recognition as a valid species is based on the coarsely punctured black abdomen, the concave nature of the last abdominal sternite which is broadly rounded in both sexes and on the consistently immaculate humeral angle of the elytra. Acmaeodera condita Barr, new species (Figure 11) Male: Medium-sized, robust, black; elytra with several pairs of large transverse and small lateral yellow spots. Head rather finely, densely punctured, densely clothed with long, erect blackish hairs; vertex with a faintly indicated median longitudinal carina; front convex except for a subflattened area at middle; clypeus distinctly punctate, front margin broadly and deeply, arcuately emarginate, broadly lobed on either side of emargination; antenna extending to about middle of pronotum, serrate from fifth segment, width of outer segments subequal to their length. Pronotum slightly more than twice as wide as long, slightly wider than elytra, widest at about basal fourth, convex except for a distinct depression in front of scutellum and a shallow, indistinct depression extending obliquely from basal pits, transverse depression behind front margin distinct on either side of middle; sides arcuately narrowing from widest portion to front angles; lateral margins narrowly reflexed except at extreme base, visible from above throughout; front angles slightly produced, acute; front margin broadly lobed at middle; surface rather coarsely and sparsely punctured on disk, punctures becoming more densely placed toward sides, densely clothed with

28 172 William I;. Barr long, erect black hairs. Elytra subdepressed except for the elevated suture behind basal fourth; sides not distinctly vertical, when viewed from above they are slightly expanded behind base and at middle where they broadly, arcuately converge to the conjointly, narrowly rounded apices; front angles in side view subrectangular; lateral margins when viewed from the side nearly straight to basal fifth and then broadly and shallowly arcuate to apices, coarsely, sparsely serrate from behind middle; surface rather sparsely clothed with long, erect black hairs; stria1 punctures circular and deep, rather small near suture, coarser and somewhat elongate laterally, striae impressed apically; interstrial spaces with an indistinct row of small punctures, flattened except near apex, seventh interstrial space not distinctly elevated, third interstrial space slightly convex; markings arranged as follows: a pair of irregular somewhat elongate subbasal spots near lateral margin and on disk, a median transverse band extending from lateral margins to near suture the hind margin of which is slightly prolonged near sides, a small spot on lateral margin behind middle, a pair of spots at apical third on margin and on disk, a small slightly transverse lateral spot in front of apex and a tiny indistinct spot at apex. Ventral surface finely punctured, denser towards sides, moderately clothed with short, suberect, fine silvery hairs; front margin of prosternum retracted, subtruncate, very feebly lobed on either side of middle; last visible abdominal sternite convex on disk, depressed along lateral margins, subapical plate distinct, rather narrow and smooth, slightly undercut at apex, hind margin broadly subtruncate. Length: 9.7 mm. Width: 3.5 mm. Holotype male (California Academy of Sciences) from Kyle Canyon, Clark County, Nevada, June 26, 1966, on flowers of Rosa sp. (W. F. Barr). Paratypes designated as follows: 112 specimens from type locality collected by E. J. Allen, W. F. Barr, D. S. Horning, Jr., R. L. Penrose and R. L. Westcott from the flowers of Rosa sp. and Cowania stansburiana; 112 specimens from type locality, May 23, 1940 (W. Reeves, M. Cazier and P. C. Ting) and June 17, 1967 (W. F. Barr), June 27 and July 15, 1969 (J. W. Tilden), July 1, 1969 (L. I. Pechuman), July 25, 1942 (E. C. Van Dyke, H. E. Scullen); 5 specimens from Lee Canyon, Charleston Mountains, Clark County, Nevada, July 21, 1952 (M. Cazier, W. Gertsch and R. Schrammel) and May 29, 1950 (C. D. MacNeill); 1 specimen from Mt. Charleston, Nevada, ,000 ft., July 6, 1944 (D. J. Zinn); 2 specimens from Charleston Mountain Park, Nevada, 9000 ft., June 21,1940 (R. M. Bohart); 2 specimes from Clark County, Nevada, July 18, 1929 (C. C. Searl); 1 specimen from Governors Canyon, Nevada, 5000 ft., June 25, 1936 (McAbee). Paratypes in the collections of

29 New species of North American Acmaeodera 173 the American Museum of Natural History, California Academy of Sciences, Oregon State University, University of Idaho, University of California at Riverside, G. H. Nelson, R. L. Westcott and W. F. Barr. This species which has been recorded in the literature as A. inyoensis Cazier is distinct from that species by the presence of faintly indicated carinae on the elytra and by the reflexed lateral margins of the pronotum. A. condita is more closely related to A. knowltoni Barr and A. errans Barr. It can be separated from these species by a combination of features that include the transversely arranged elytral markings, the black abdomen and the distinct subapical plate on the last abdominal sternite. Acmaeodera alpha Barr, new species (Figure 14) Male: Medium-sized, moderately robust, black; each elytron with four, pale-yellow, transverse fasciae extending from lateral margin to near suture, first fascia immediately behind umbone, constricted at middle, second fascia slightly in front of middle, gradually expanded from lateral margin, abruptly narrowed on disk, third fascia at about apical fourth, very broad and broken at lateral margin, narrowing before disk, slightly expanded on disk, fourth fascia in front of apex, small, constricted near lateral margin; upper surface rather densely clothed with moderately long, erect black hairs except for lateral areas of elytra which have shorter, suberect black hairs; ventral surface moderately clothed with short, suberect black hairs. Head densely punctured, punctures small and deep; vertex without a median carina; front feebly convex except for transverse depression above clypeus; clypeus roughened laterally, front margin broadly, subtriangularly emarginate; antenna shining, extending to about middle of pronotum, serrate from fifth segment, outer segments slightly wider than long. Pronotum slightly more than twice as wide as long, wider than elytra, widest at basal third, convex except for a broad, shallow median depression in front of base and a broad shallow depression extending obliquely from the distinct basal pits, transverse depression behind front margin distinct on either side of middle; sides strongly arcuate; lateral margins very narrowly reflexed on anterior half where they are visible from above; front angles feebly produced, narrowly rounded; front margin broadly lobed at middle; hind margin subtruncate; surface more coarsely, densely punctured than head, somewhat reticulate, becoming more densely roughened laterally. Elytra flattened except for a feebly elevated area between first and second fasciae and a slight basal depression; sides vertical on apical half, when viewed from above they are slightly expanded behind base and then gradually, broadly sinuately narrowing to

30 174 William F. Burr the conjointly, broadly rounded apices; front margin beaded; front angles in side view slightly produced ventrally, acute; lateral margins when viewed from the side broadly and feebly sinuate on basal fourth, shallowly emarginate behind first fasciae, and then broadly arcuate to apices, coarsely serrate on apical third; stria1 punctures coarse and deep, rather uniform except irregular and dense at base and very coarse on discal black areas between third and seventh interstrial spaces, striae impressed behind middle; interstrial spaces flattened, irregularly roughened at base, transversely wrinkled near sides, interstrial punctures relatively large, third interstrial space slightly elevated, seventh interstrial space faintly elevated on basal half, ninth interstria1 space faintly elevated at base, suture moderately elevated on apical half. Ventral surface corasely, densely punctured on thoracic sternites, more finely punctured on metacoxa, coarsely, sparsely punctured on basal half of first abdominal sternite, more finely punctured on apical half, finely, densely punctured on remaining abdominal sternites, but punctures very shallow on sternites three to five; front margin of prosternum retracted not reflexed, feebly bisinuate; last visible abdominal sternite with sides broadly reflexed, hind margin nearly semicircularly rounded, disk convex, microreticulate on apical half, with a rudiment of a tiny subapical plate. Length: 10.0mm. Width: 3.8mm. Holotype male (California Academy of Sciences) and 11 paratypes from Second Lake, Big Pine Creek Canyon, Inyo County, California, 10,040 ft., July 1, 1960 (W. F. and M. E. Barr). Paratypes designated as follows: 4 specimens from Upper Big Pine Creek Canyon, Inyo County, California, ft., July 1, 1960 (W. F. and M. E. Barr); 7 specimens from Glacier Lodge, Inyo County, California, August 4, 7 and 8, 1942 (G. P. Mackenzie); 2 specimens from Lone Pine Creek, Inyo County, California, 8250 ft., July 7, 1961 (D. R. Miller); 1 specimen from Whitney Portal, Inyo County, California, July 3, 1953 (H. Nakakihara); 2 specimens from White Mountains, Mono County, California, 10,000 ft., June 26, R. Miller); 1 specimen from 1 mile south of Tom's Place, Mono County, California, August 8, 1962 (A. S. Menke); 1 specimen from Rock Creek, Tom's Place, Mono County, California, June 22, Paratypes in the collections of the American Museum of Natural History, California Academy of Sciences, University of California at Davis and Riverside and W. F. Barr. This high elevation species can be recognized most readily by its black to blue-black base coloration and by the presence of black hairs on the body, especially on the ventral surface. Its affinities are with A. idahoensis Barr.

31 New species of North American Acmaeodera 175 Acmaeodera nelsoni Barr, new species (Figure 15) Male: Medium-sized, elongate, dark brown with a bronzy tinge except on elytra which has a more purplish tinge; elytra with a tiny elongate yellow spot on lateral margin at basal fourth, with a large transverse yellow spot in front of middle that extends from lateral margin to fifth interstrial space, with a large transverse yellow spot at about apical three-fourths that extends from lateral margin to eighth interstrial space and with a small rounded subapical yellow spot on lateral margin in front of apex; dorsal surface rather densely clothed with long, erect dark brown hairs; ventral surface moderately clothed with short to long, suberect silvery hairs. Head rather coarsely, densely punctured; vertex with a longitudinal median carina; front convex except for a.shallow but distinct depression at middle and a transverse depression above clypeus; clypeus densely punctured, front margin rather deeply and broadly subtriangularly emarginate; antenna feebly shining, extending to about base of pronotum, serrate from fifth segment, width of outer segments subequal to their length. Pronotum twice as wide as long, about equal in width to elytra; subconvex except for feebly indicated depressions at middle in front of base, at middle near front margin, and in front of basal pits, transverse depression behind front margin not evident; sides slightly expanded at basal fourth, broadly arcuately narrowing to front angles; lateral margins narrowly reflexed, visible from above except at extreme base; front angles produced, broadly acute; front margin broadly but feebly lobed at middle; hind margin subtruncate; surface rather coarsely and sparsely punctured medially, punctures becoming very coarse and more dense laterally. Elytra subdepressed except for a broad, shallow median depression at base and a slight elevation midway between suture and each humerus; sides more or less vertical on posterior half, when viewed from above they are very slightly expanded immediately behind base, broadly but feebly constricted at about basal third, slightly expanded at middle, and then gradually converging to the conjointly, broadly rounded apices; front margin slightly beaded; front angles from side view slightly produced ventrally, subacute; lateral margins when viewed from the side broadly bisinuate to basal fourth and then very broadly and feebly sinuate to apices, very coarsely and sharply serrate on apical half; stria1 punctures rather coarse, somewhat elongate and deep on disk, becoming coarser and irregular at base and coarser and denser laterally, striae impressed behind apical third; interstrial spaces subflattened, tiny interstrial punctures indistinct on disk, conspicuous on apical half, third, seventh and ninth interstrial spaces not elevated, suture strongly ele-

32 176 William F. Barr vated on apical half. Ventral surface coarsely, very densely punctured at sides of thoracic sternites and first abdominal sternite, more sparsely punctured medially, slightly less coarsely punctured on abdominal sternites two to five; front margin of prosternum distinctly retacted, narrowly reflexed, very broadly, shallowly bisinuate; last visible sternite nearly semicircularly rounded at apex, sides distinctly reflexed, disk feebly convex with a faint indication of a nearly smooth, broadly rounded subapical plate. Length: 8.4 mm. Width: 2.8 mm. Holotype male (California Academy of Sciences) and 23 paratypes from 3 miles south of Camp Angeles, San Bernardino Mountains, California, 5000 ft., June 6, 1958, on Ceanothus crassifolius (G. H. Nelson). Additional paratypes as follows: 16 specimens from the type locality June 1, 1959 and June 6, 8, 15 and 20, 1958 (G. H. Nelson); 11 specimens from Tanbark Flat, Los Angeles County, California, June 12 and 20, 1950, on Lotus (J. W. MacSwain), June 21 and 22, 1950, on Ceanothus (H. F. Robinson), June 22, 1950 (J. C. Hall), June 23, 1950 (W. 0. Marshall, K. G. Whitesell), and June 28 and July 3, 1950 (A. T. McClay); 15 specimens from Stone Canyon Res., Santa Monica Mountains, Los Angeles County, California, June 4, 1957 (R. L. Westcott); 11 specimens from Mt. Diablo, Contra Costa County, California, June 6, 1954 (J. R. Helfer), June 9, 1952, June 10-14, 1949 and June 10, 1952 (F. X. Williams), June 14, 1933, on Ceanothus, June 25, 1919, and August 12, 1939 (W. F. Barr); 4 specimens from Beverly Glen, Los Angeles County, California, May 24, 1958 (D. S. Verity); 37 specimens from Sunset Valley, Santa Barbara County, California, June 28, 1940 (W. F. Barr), July 3 and 4, 1939, on Cercocarpus betuloides, Adenostoma fasciculatum, Quercus dumosa, and Photinia arbutifolia (W. F. Barr, E. C. Van Dyke, B. E. White); 11 specimens from California Hot Springs, Tulare County, California, June 3-5, 1939 and July 7, 1925; 10 specimens from 18 miles south of Zenia, Trinity County, California, June 23, 1969, on Ceanothus cuneatus (W. F. Barr). Paratypes in the collections of the American Muesum of Natural History, California Academy of Sciences, Los Angeles County Museum, Philadelphia Academy of Sciences, University of California at Berkeley and Davis, University of Idaho, R. C. Bechtel, G. H. Nelson, F. H. Parker, R. L. Westwtt and W. F. Barr. Additional specimens not designated as paratypes have been examined from the following California localities: Mt. Wilson, Los Angeles County; Beaumont, Keen Camp, Herkey Creek, Idyllwild and San Jacinto Mountains above Hemet, Riverside County; 8 miles west of Anberry, Fresno County; Ben Lomond; Carpenteria; Mariposa; Santa Lucia Mountains; Sequoia

33 New species of North American Acmaeodera 177 National Park; Atascodero; Hullville, Lake County; Havilah; Placerville; and Pope Valley, Napa County. For many years this species has been known in the literature and in most collections as A. dohrni Horn. That name however, applies to another species and thus a new name is necessary for the species here under consideration. A. nelsoni belongs to the California chapparal inhabiting complex of species that includes A. mariposa Horn, A. dohrni Horn, A. prorsa Fall, A. simulata Van Dyke and A. nautica Van Dyke. It is recognized principally by the dark brown body usually with orange colored elytral markings, by the faintly indicated subapical plate of the last abdominal sternite and by the broadly arcuate lateral margins of the pronotum. This species is named after Dr. Gayle H. Nelson who is well-known for his taxonomic studies and field work on North American Buprestidae. Acmaeodera acanthicola Barr, new species (Figure 16) Male: Small-sized, subcylindrical, blue-black, shining; elytra dark blue with a large, median red spot extending from lateral margin to about sixth interstrial space; upper and lower surfaces sparsely clothed with very fine, short silvery hairs that become denser on lateral areas of meso- and metathorax and abdomen. Head rather finely and densely punctured, punctures becoming less dense on vertex; vertex with a short, indistinct longitudinal median carina; front concave above clypeus, subflattened elsewhere; clypeus finely, densely punctured, front margin deeply, subtriangularly emarginate; antenna black, shining, extending to about middle of pronotum, serrate from fifth segment, outer segments slightly wider than long. Pronotum approximately one and one-half times wider than long, widest at base, slightly narrower than elytra; convex, subflattened at middle in front of base, basal pits distinct, not associated with an oblique depression, transverse depression behind front margin indistinct; sides sinuately narrowing from base to front angles; lateral margins indicated by a narrow carina, not visible from above; front angles not produced, obtuse; front margin broadly lobed at middle; hind margin broadly, arcuately emarginate; surface less coarsely and less densely punctured than head, punctures becoming coarser and more dense laterally. Elytra convex except for a small depression on either side of suture at base; sides when viewed from above feebly constricted at about basal third, broadly, arcuately converging to the conjointly, rather narmwly

34 178 William F. Burr rounded apices; front angles in side view produced obliquely to the front, narrowly acute; lateral margins when viewed from the side very broadly and shallowly emarginate at about basal fourth, then broadly arcuate to apices, apical fourth indistinctly, coarsely and sparsely serrate; stria1 punctures small, deep, somewhat elongate on disk becoming much coarser and circular toward sides, striae deeply impressed behind apical third; interstrial spaces flattened on disk, slightly convex apically and laterally where they are transversely roughened, interstrial punctures distinct, third interstrial space and suture very slightly elevated. Ventral surface coarsely, densely punctured on sides of thoracic sternites and first abdominal sternite, finely, densely punctured on abdominal sternites two to five, sparsely punctured medially; front margin of prosternum not retracted, narrowly reflexed, subtruncate with a slight lobe-like swelling on either side of middle; last visible abdominal sternite with hind margin broadly rounded, sides reflexed, disk convex, with an irregularly transverse, densely punctured, subapical swelling. Length: 6.2mm. Width : 2.lmm. Holotype male (California Academy of Sciences) and 13 paratypes from 2 miles east of Portal, Cochise County, Arizona, August 18, 1957, on Acacia constricts (M. A. Cazier). Two paratypes from type locality August 26, 1957 (W. F. Barr); 6 paratypes from 1 mile east of Portal, Cochise County, Arizona, August 18, 1957 and August 19, 1960, on Acacia vernicola (J. A. Chemsak, B. J. Rannels); 6 paratypes from 2-3 miles southeast of Portal, Cochise County, Arizona, July 13 and August 1, 1959 (M. Statham); 24 paratypes from Porta1,Chiricahua Mountains, Arizona, August 2, 1958 (J. M. Marston), August 2-5, 1965 (B. K. Dozier); 2 paratypes from Southwest Research Station, 5 miles west of Portal, Cochise County, Arizona, 5400 ft., August 12, 1956 (E. Ordway); 34 paratypes from Globe, Arizona, July 6, 1938, July 24,1960, on palo verde, August 2, 1933, August 6, 1939, August 9 and 27 and September 6, 1942, and September 9, 1941 (F. H. Parker); 18 paratypes from Tucson, Arizona, July 13-15, August 8-16 and (Wickham), July 16 and 20 and August 16, 1940 (D. J. and J. N. Knull), August 13, 1966 (R. L. Westcott) and September 4, 1928 (F. W. Nunenmacher); 209 paratypes from Sabino Canyon, Santa Catalina Mountains, Arizona, August 5, 1946, on palo verde (F. H. Parker), August 14, 1950 (R. S. Beal) and August 24, 25 and 26, 1959 (G. H. Nelson); 8 paratypes from Santa Catalina Mountains, Arizona, August 13, 1940; 67 paratypes from Lower Madera Canyon, Santa Rita Mountains, Arizona, August 23, 1959, on Acacia (G. H. Nelson), and October 29, 1970 (J. W. and B. A. Tilden); and 14 paratypes from 39 miles north of Hermosillo, Sonora, July 28, 1957, on Palo Verde (R. L. Westcott).

35 New species of North American Acmaeodera 179 Paratypes in the collections of the American Museum of Natural History, California Academy of Sciences, University of California at Berkeley, Davis and Riverside, F. M. Beer, B. K. Dozier, G. H. Nelson, F. H. Parker, B. A. Tilden, R. L. Westcott and W. F. Barr. This is another species that has been known for many years by another name, which in this case is A. stigmata Horn. The incorrect application of this name results from the fact that Horn had two species making up his type series. His lectotype represents a Lower California species and his other type material is what is described herein as A. acanthicola. This species appears to be related to A. oaxacae Fisher and to A. stigmata. From the former, it is separated by the more conspicuously punctured pronotum, by the absence of lateral pits on the third and fourth abdominal sternites and by usually having an apical thickening of the last abdominal sternite. From the latter, it differs by being more slender, by having blue rather than blackish elytra and by the more weakly developed teeth on the front margin of the prosternum. Acmaeodera davidsoni Barr, new species (Figure 17) Male: Small-sized, subcylindrical, black, shining; elytra with a large, rounded, median red spot extending from lateral margin to fourth intrstrial space; head and pronotum moderately clothed with rather short silvery hairs, elytra sparsely clothed with very fine, suberect silvery hairs; ventral surface sparsely clothed with very fine, short silvery hairs that become coarser and very dense on lateral areas of meso- and metathorax and abdomen. Head rather coarsely and sparsely punctured; vertex without an apparent median carina; front convex except for a small transverse depression above clypeus and a median, subflattened area on upper portion; clypeus finely, densely punctured, front margin very deeply, arcuately emarginate; antenna black, shining, extending to about middle of pronotum, serrate from fifth segment, outer segments distinctly wider than long. Pronotum approximately one and one-half times wider than long, widest at base, slightly narrower than elytra, convex except for a very small faintly indicated shallow, median depression at base, basal pits distinct, not associated with an oblique depression, transverse depression behind front margin faintly indicated; sides broadly and shallowly, arcuately narrowing from base. to front angles; lateral margins indicated by a small carina, not visible from above; front angles not produced, subrentangular; front margin broadly lobed at middle; hind margin

36 ,180 William F. Burr broadly, arcuately emarginate; disk finely, sparsely punctured, punctures becoming very coarse and more densely placed laterally. Elytra convex except for a faint median depression at base; sides when viewed from above subparallel to near middle except very slightly and broadly constricted at about basal third, broadly, arcuately converging to the conjointly, rather broadly rounded apices; front margin beaded at middle; front angles in side view slightly produced obliquely to the front, narrowly acute; lateral margins when viewed from the side broadly and shallowly emarginate at about basal third, then very broadly and shallowly arcuate to apices, apical fourth indistinctly coarsely and sparsely serrate; stria1 punctures small and shallow on disk, becoming coarser and deeper toward base and sides and indistinct apically, striae deeply impressed bahind apical third; interstrial spaces flattened on disk, slightly convex apically and laterally, interstrial punctures subequal to stria1 punctures on disk, becoming slightly smaller laterally and apically; third interstrial space slightly elevated, suture not elevated. Ventral surface coarsely, rather densely punctured on lateral areas of thoracic sternites and first abdominal sternite, more sparsely punctured at middle, very finely punctured on remaining abdominal sternites; front margin of prosternum very slightly retracted, not reflexed except at extreme sides, shallowly emarginate with a distinct rounded tooth on either side of middle; last visible abdominal sternite with hind margin broadly rounded, sides slightly reflexed, disk convex with an irregularly transverse, feebly elevated, densely punctured subapical ridge. Length: 6.7 mm. Width: 2.4 mm. Holotype male (California Academy of Sciences) from Granite Gap, Peloncillo Mountains, Hidalgo County, New Mexico, July 10, 1965, on Acacia constricta (J. H. Davidson, J. M. Davidson and M. A. Cazier). Forty paratypes from type locality June 14, 16 and 21, 1964 and June 20 and 29, 1965, on Acacia constricta and Mimosa biuncifera (J. H. Davidson, J. M. Davidson and M. A. Cazier); 18 paratypes from 15 miles north of Rodeo, New Mexico, June 15, 1965 and July 10, 1969, on Acacia constricta and Mimosa sp. (G. H. Nelson); 3 paratypes from 3 miles north of Presidio, Texas, July 20, 1966 on Acacia greggii (C. L. Cole); 4 paratypes from 3.5 miles northwest of Presidio, Texas, July 31, 1969 (D. E. Foster, L. S. Hawkins and R. L. Penrose); 1 paratype from 20 miles north of Presidio, Texas, July 31, 1969, on Acacia greggii (G. H. Nelson); and 3 paratypes from from 6 miles east of Panther Junction, Big Bend National Park, Texas, July 4-6, 1961, on Acacia (R. L. Westcott). Paratypes in the collections of Arizona State University, the University of Idaho, J. M. Davidson, G. H. Nelson, S. G. Wellso, R. L. Westcott and W. F. Barr.

37 New species of North American Acmaeodera 181 The black body with the pair of red spots at the middle of the elytra, the punctures of the interstrial spaces being nearly subequal to the stria1 punctures at the middle and the finely, sparsely punctured pronotal disk differentiate this species from its allies. In form it approximates A. acanthicola and in colaration it resembles A. stigmata. This species is named after one of the indefatigable collectors of the type series, J. M. Davidson, formerly of Arizona State University. Acmaeodera clypeata Barr, new species (Figure 18) Male: Small-sized, subcylindrical, shining, purplish-black; each elytron with a lateral and a discal row of very small yellow spots, spots of lateral row located at about basal fourth, in front of middle, behind middle, at about apical fourth and at about apical eighth, those of discal row located in front of basal fourth, at about basal third, at about apical third, at about apical fourth, and at apex, in addition a small spot is present at inner margin of umbone, the two apical spots of lateral row joined on left elytron; head moderately clothed with suberect, recurved white hairs, a few of which are branched above clypeus; pronotum sparsely clothed with strongly recurved, fine white hairs that become coarser and branched and more densely placed laterally; elytra moderately clothed with subrecumbent, short, fine white hairs that become doubly branched at sides; ventral surface moderately clothed with subrecumbent short digitate scales that become more elongate and more densely placed at sides of metathorax and first three abdominal sternites and which are replaced medially by short bowed hairs. Head coarsely, rather densely, shallowly punctured; vertex with a faintly indicated, short, median longitudinal carina; front broadly, deeply, longitudinally depressed; clypeus indistinctly punctured, roughened, front margin very deeply, nearly semicircularly emarginate; antenna heavy, extending slightly beyond middle of pronotum, serrate from fifth segment, outer segments wider than long. Pronotum slightly more than one and two-thirds times wider than long, widest at middle, subequal in width to elytra, convex except for disk feebly longitudinally depressed and antescutellar area broadly shallowly depressed, basal pits distinct, transverse depression behind front margin distinctly indicated on either side of middle; sides strongly arcuate; lateral margins very slightly reflexed, carinate throughout, visible from above on about anterior third; front angles feebly produced, narrowly rounded; front margin broadly lobed at middle; hind margin broadly, shallowly emarginate; surface coarsely, rather densely, deeply punctured on disk, punctures

38 182 William F. Burr becoming nearly confluent toward sides. Elytra moderately convex; sides when viewed from above very slightly wider behind base than at middle, broadly, feebly constricted at about basal fourth, subparallel medially, broadly, evenly arcuate from behind middle to the conjointly, broadly rounded apices; front angle in side view strongly produced ventrally and anteriorly, acute; lateral margin when viewed from the side shallowly emarginate at about basal fifth, then broadly, shallowly sinuate to apices, coarsely serrate on about apical half; stria1 punctures moderate in size, deep and moderately placed on disk, coarser and more densely placed laterally, striae strongly impressed behind middle; interstrial spaces subflattened on disk, roughened and with many small but distinct teeth laterally, interstrial punctures rather small and shallow on disk, indistinct laterally, third interstrial space faintly indicated from behind umbone to near apex, suture slightly elevated behind middle. Ventral surface rather sparsely and shallowly punctured, punctures moderate in size except at base of first abdominal sternite where they are very coarse near sides; front margin of prosternum distinctly retracted, feebly reflexed near sides, with a blunt, rather broad tooth on either side of middle; abdominal sternites two, three and four with hind angles acute, sternites three and four with lateral margins distinctly reflexed and strongly depressed at base, last visible abdominal sternite with margins narrowly reflexed, lateral margins strongly narrowing, very feebly arcuate, moderately depressed at base, hind margin broadly rounded, disk convex with a broad, densely punctured subapical elevation. Length: 7.0 mm. Width: 2.6 mm. Holotype male (California Academy of Sciences) and 62 paratypes from 3 miles west of Tehuantepec, Oaxaca, Mexico, July 9, 1965 (G. H. and D. E. Nelson); 1 paratype from Barranca Hondo, 19 miles south of Uruapan, Michoacan, 3000 ft., July 5, 1970 (E. M. Fisher); and 1 paratype from 9 miles east of Capirio, Michoacan, 800 ft., July 6, 1970 (E. M. Fisher, P. Sullivan). Paratypes in the collections of G. H. Nelson, D. S. Verity, R. L. Westcott and W. F. Barr. This species, a member of the pinalorum group, is unlike any others. It can be easily recognized by the semicircularly emarginate front margin of the clypeus, the reduced elytral markings and the finely toothed lateral interstrial spaces. In general form and elytral pubescence this species resembles A. pinalorum Knull.

39 New species of North American Acmaeodera 183 Acmaeodera adusta Barr, new species (Figure 19) Male: Small-sized, subcylindrical, shining, dark coppery in color; each elytron with a metallic bluish tinge at base and a lateral and a discal row of medium-sized yellow spots that are somewhat transverse behind middle, spots of lateral row located at front angle, at basal fourth, in front of middle, behind middle, at about apical fourth, and at about apical eighth, those of discal row located at base, at about basal fourth, in front of middle, behind middle, at about apical fourth, and at apex; head moderately clothed with subrecumbent, stout white hairs; pronotum sparsely clothed with subrecumbent white hairs which become more numerous and two-branched near front angles; elytra rather sparsely clothed with subrecumbent, short white hairs; ventral surface moderately clothed with subrecumbent, elongate digitate scales that are more sparsely placed medially and replaced by dense, elongate compound hairs along sides of metathorax and by short bowed hairs at middle of apical abdominal sternites. Head coarsely, densely, rather shallowly punctured; vertex with a well-developed median longitudinal carina that extends onto frons between upper portions of eyes; front broadly, longitudinally depressed at middle and transversely depressed above clypeus; clypeus indistinctly punctured, shining, front margin shallowly, broadly, arcuately emarginate; antenna heavy, extending to slightly beyond middle of pronotum, serrate from fifth segment, outer segments wider than long. Pronotum nearly one and three-fourths times wider than long, widest at middle, subequal in width to base of elytra, convex, subflattened in front of scutellar area, basal pits distinct, transverse depression behind front margin feebly indicated on either side of middle; sides rather strongly arcuate; lateral margins not reflexed, carinate on apical half where they are visible from above; front angles feebly produced, rather narrowly rounded; front margin strongly and broadly lobed at middle; hind margin very shallowly emarginate; surface densely, deeply punctured at middle, punctures more dense and shallower toward sides. Elytra strongly convex; sides when viewed from above slightly wider behind base than at middle, broadly but feebly constricted at about basal fourth, subparallel medially, broadly evenly arcuate from behind the middle to the conjointly, broadly rounded apices; front angles in side view strongly produced ventrally and anteriorly, acute; lateral margin when viewed from the side rather deeply, arcuately emarginate at about basal fifth, then broadly, shallowly sinuate to apices, coarsely serrate on about apical half; stria1 punctures coarse, deep and densely placed on basal half and near sides, smaller on apical half near suture, striae strongly impres-

40 184 William F. Barr sed behind middle; interstrial spaces subconvex, transversely roughened or scabrous apically near sides, interstrial punctures rather large, distinct, third intrstrial space slightly elevated from behind umbone to near middle, ninth interstrial space feebly elevated immediately behind base, suture elevated behind middle. Ventral surface coarsely and densely punctured, punctures finer and sparsely placed medially; front margin of prosternum distinctly retracted, slightly reflexed near sides, with a conspicuous, broad median lobe that is strongly toothed laterally and broadly, arcuately emarginate between the teeth; last visible abdominal sternite with margins very narrowly reflexed, hind margin broad, feebly rounded, disk slightly convex with a very faint indication of a densely punctured transverse subapical elevation. Length: 7.2 mm. Width: 2.8 mm. Holotype male (California Academy of Sciences) and 17 paratypes from 9 miles south of Cuatros Caminos, Michoacan, Mexico, July 20, 1966 (D. S. Verity, G. Walters) collected from Acacia sp. Paratypes in the collections of G. H. Nelson, D. S. Verity, G. Walters and W. F. Barr. This southern Mexico species closely resembles the widely distributed A. pinalorum Knull. However, A. adusta is much more coppery in coloration with the elytral markings larger, the elytral pubescense shorter and more sparsely placed and the disk of the pronotum more coarsely punctured than that species. Acmaeodera lauta Barr, new species (Figure 20) Male: Small-sized, subcylindrical, feebly shining, dark brown with a conspicuous coppery luster except at base of elytra where a bluish luster is evident; each elytron with a lateral and a discal row of medium-sized irregular yellow spots, spots of lateral row located at front angle, behind umbone, in front of middle, immediately behind middle, at about apical fourth, and at about apical sixth, those of discal row located at base, at basal Bth, in in front of middle, behind middle, and at about apical fourth, in addition, a small spot is present at inner margin of right umbone, three apical spots of lateral row faintly reddish internally; head and lateral areas of pronotum moderately clothed with recumbent scale-like white hairs, elytra more sparsely clothed along sides and on apical half with elongate appressed scales that are intermixed with short, suberect white hairs; ventral surface moderately clothed with subrecumbent, robust digitate scales that become denser and more elongate toward sides of metathorax and which are replaced medially

41 New species of North American Acmaeodera 185 on apical abdominal sternites by short bowed hairs. Head coarsely, densely punctured; vertex with a very short, median longitudinal carina; front strongly depressed at middle and transversely depressed above clypeus; clypeus irregularly punctured, roughened, front margin shallowly and broadly arcuately emarginate; antenna heavy, extending slightly beyond middle of pronotum, serrate from fifth segment, outer segments distinctly wider than long. Pronotum nearly one and three-fourths times wider than long, widest at middle, slightly narrower than elytra, strongly convex with disk narrowly, longitudinally depressed anteriorly and broadly, shallowly depressed posteriorly, basal pits distinct, transverse depression behind the front margin evident on either side of middle; sides strongly arcuate; lateral margins not reflexed, but visible from above on apical half; front angles feebly produced, narrowly rounded; front margin strongly lobed at middle; hind margin very broadly and shallowly, subtriangularly emarginate; surface coarsely, deeply, rather sparsely punctured at middle, punctures more coarse and very dense toward sides. Elytra strongly convex; sides when viewed from above broadest behind base, broadly, feebly constricted at about basal fourth, subparallel medially, broadly, evenly arcuate from behind middle to the conjointly, broadly rounded apices; front angle in side view strongly produced ventrally and anteriorly, acute; lateral margins when viewed from the side deeply emarginate at about basal fifth, then very broadly, shallowly sinuate to apices, coarsely serrate on apical third; stria1 punctures coarse, deep and densely placed, smaller and more linear near suture and apically where the striae are strongly impressed; interstrial spaces subconvex, somewhat irregular because of dense stria1 punctures except for spaces nine, ten and eleven, third interstrial space slightly elevated from behind umbone to near apex, suture feebly elevated, interstrial spaces near sides roughened or scabrous apically, interstrial punctures rather large, distinct. Ventral surface coarsely, rather densely punctured, punctures slightly smaller and shallower on abdominal sternites two to five; front margin of prosternum distinctly retracted, not evidently reflexed laterally, with a conspicuous, broad median lobe that is strongly toothed laterally and arcuately emarginate between the teeth; last visible abdominal sternite only with the broadly rounded hind margin reflexed, disk convex without a subapical elevation. Length: 7.7 mm. Width: 2.9 mm. Holotype male (California Academy of Sciences) and 55 paratypes from 3 miles west of Tehuantepec, Oaxaca, August 2, 1965, on dead limbs (G. H. Nelson). Additional paratypes from the type locality include: 245 specimens July 9, 10, 19 and 20, 1965, on Acacia pennatula and on dead

42 186 William F. Barr limbs (G. H. Nelson) and 42 specimens reared during the period June 28-September 10, 1966 from Loncharpus sp. that had been collected in August Other paratypes designated as follows: 2 specimens from 8 miles west of Tehuantepec, Oaxaca, August 16, 1963 (A. Hardy); 1 specimen from S. Gerommo, Oaxaca, August 6, 1925 (E. G. Smyth); 1 specimen from Xalitla, Guerrero, June 4, 1946 (J. and D. Pallister); 1 specimen from Canyon del Zopilote, 24 miles north of Chilpancingo, Guerrero, July 11, 1970 (E. Fisher and P. Sullivan); 24 specimens from 9 miles east of Capirio, Michoacan, 800 ft., July 6, 1970 (E. Fisher and P. Sullivan); 1 specimen from 12 miles southwest of Tiquicheo, Michoacan, 2000 ft., July 9, L970 (E. Fisher and P. Sullivan); 1 specimen from 10 miles west of Colima, Colima, August 1, 1954 (W. Gertsch and Bradts); and 1 specimen from 8 miles north of Tecoman, Colima, July 25, 1966 (D. S. Verity). Paratypes in the collections of the American Museum of Natural History, Los Angeles County Museum, G. H. Nelson, D. S. Verity, R. L. Westcott and W. F. Barr. In form and structure, expecially in the convexity of the dorsal surface, the angulate sides of the pronotum, the discal depression of the pronotum and the punctation of the pronotum and elytra, A. lauta closely resembles A. adusta Barr. These species are distinctive however, in their coloration and in the nature of their elytral pubescense. Because of the presence of recumbent scales on the dorsal surface A. lauta may be confused with A. mimicata Knull and A. fattigi Knull of central Arizona and southeastern California. These two species, however, have a darker body color, they are not so coarsely sculptured, and they do not have the disk of the pronotum depressed and the sides of the pronotum angulate. Acmaeodera westcotti Barr, new species (Figure 21) Male: Small-sized, subcylindrical, shining, brown with a distinct brassy luster; each elytron with a sublateral and a discal row of small yellow spots, sublateral row in part associated with second interstrial space, spots located in front of basal fourth, in front of middle, behind middle and at apical fourth, spots of discal row located in front of basal fourth, at middle, at apical third, and behind apical fourth; head sparsely clothed with stout recumbent white hairs that become finer on vertex and coarser and more scalelike above clypeus, pronotum sparsely clothed with fine, recumbent white hairs that become coarser toward sides and stout and scale-like along lateral margins, hairs directed away from a small median area in front of base,

43 New species of North American Acmaeodera 187 elytra sparsely clothed with short, stout broad hairs that are curved posteriorly at the tip, the hairs tending to be pointed on basal half of disk and flattened and somewhat scale-like along lateral margins; ventral surface densely clothed with broad, appressed, finely branched scales that are intermixed along sides of thoracic sternites with long, recumbent, multi-branched white hairs, surface obscured. Head coarsely and rather densely, very shallowly punctured; vertex with a longitudinal smooth area at middle, not carinate; front feebly convex except for a distinct transverse depression above clypeus; clypeus finely, densely punctured, front margin very shallowly, broadly, arcuately emarginate; antenna blackish, extending to slightly beyond middle of pronotum, serrate from Uth segment, outer segments subequal in width and length. Pronotum nearly one and one-third times wider than long, widest at middle, narrower than elytra, convex, without any depressed areas, basal pits distinct; sides evenly arcuate; lateral margins not reflexed, faintly carinate near front angles, not visible from above; front angles not produced, subrectangular; front margin shallowly lobed at middle; hind margin very feebly emarginate; surface rather coarsely, sparsely and shallowly punctured at middle, punctures becoming coarser, deeper and more densely placed toward sides. Elytra convex; sides when viewed from above broadest at about middle, distinctly expanded immediately behind base, broadly and distinctly constricted at about basal fourth, then broadly, evenly arcuate to the conjointly, broadly rounded apices; front margin beaded at middle; front angles in side view strongly produced ventrally and slightly produced anteriorly, acute; lateral margins when viewed from the side feebly sinuate on about basal sixth, rather deeply emarginate at about basal fifth, then broadly arcuate to apices, serrate on about apical fourth; stria1 punctures small and deep on disk, becoming smaller and shallower apically and coarser and denser toward base and sides, striae deeply impressed behind middle; interstrial spaces slightly convex, becoming transversely roughened toward sides, interstrial punctures distinct, third interstrial space slightly elevated on basal half, suture not elevated except between basal fourth and middle. Ventral surface with punctures nearly obscured by pubescence, however, the thoracic sternites appear to be coarsely, densely punctured and the abdominal sternites finely, densely punctured; front margin of prosternum hot retracted, slightly separated from front margin of pronotum, not reflexed, subtruncate with a small rounded tooth om either side of middle; last visible abdominal sternite with hind margin broadly, nearly semicircularly rounded, sides not distinctly reflexed, disk subflattened with a very faint indication of a subapical elevation. Length: 6.5 mm. Width: 2.9 mm.

44 188 William F. Barr Holotype male (California Academy of Sciences) and 39 paratypes from 15 miles east of Calexico, Imperial County, California, June 6, 1961, on Prosopis chilensis (G. H. Nelson and H. F. Howden). Additional paratypes as follows : 1 13 specimens from Winterhaven, Imperial County, California, June 10, 1961 and July 1, 1962, on S. Verity), June 20, 1965, on Prosopis (G. Walters) and June 22, 1963, on mesquite (R. L. Westcott); 1 specimen from 8 miles east of Holtville, Imperial County, California, July 4, 1959, on Verity); 28 specimens from Santa Rosa Mountains, Highway 74, Riverside County, California, 2000 ft., June 12, 1960, on mesquite (R. L. Westcott) and June 17, 1961, on Prosopis chilensis and Acacia greggii (D. S. Verity); 15 specimens from above Palm Desert, Highway 74, Riverside County, California, June 21 and 28, 1961, on Prosopis chilensis; 10 specimens from 1 mile north of Mecca, Riverside County, California, June 21 and 25, 1959, on Atriplex lentiformis (G. H. Nelson); 21 specimens from 2 miles north of Mecca, Riverside County, California, June 10, 1961, on dead twigs of Prosopis chilensis (D. S. Verity); 1 specimen from Thermal, Riverside County, California, June 16, 1963 (D. S. Verity); 1 specimen from 1 mile south of Thermal, Riverside County, California, June 30, 1960, on Prosopis pubescens (G. H. Nelson); 1 specimen from Indio, Riverside County, California, June 23, 1948 (A. T. McClay); 1 specimen from Moronga Valley, San Bernardino County, California, July 4, 1958, on Prosopis (G. H. Nelson); 1 specimen from Borego Valley, San Diego County, California, June 8, 1940, on Acacia; 3 specimens from 57 miles south of Mexicali, Baja California, June 3, 1966 (D. S. Verity); 1 specimen from 3 miles east of El Progresso, Baja California, June 8, 1966, on dead branch of Prosopis (D. S. Verity); and 1 specimen from El Mayor, Baja California, June 15, 1952 (M. Cazier, W. Gertsch and R. Schrammel). Paratypes in the collections of the American Museum of Natural History, Los Angeles County Museum, the University of California at Davis and Riverside, G. H. Nelson, D. S. Verity, G. Walters, R. L. Westcott and W. F. Barr. A. westcotti shows a distributional pattern that restricts it to the Colorado Desert area of southern California and northern Baja California. This species can be separated from A. hassayampae Knull to which it is most closely related and from the other species of the guttifera group by a combination of characters that include a brown to dark brown body coloration with a shining brassy luster, two rows of spots on each elytron, a shallowly punctured pronotal disk, and the presence of pointed or parallel sided hairs on the elytra. Frequently, the spots near the middle of each elytron are

45 New species of North American Acmaeodera 189 joined transversely, but they are never joined longitudinally. Also, a subbasal spot is often present at the inner margin of the humeral umbone. This species is dedicated to Richard L. Westcott of the Oregon Department of Agriculture who has generously provided me with much of the material used in this study. Acmaeodera paravaripilis Barr, new species (Figure 22) Male: Small-sized, subcylindrical, feebly shining, black; head, pronoturn and ventral surface with a strong cupreous luster, each elytron with a sublateral and a discal row of small yellow spots, sublateral row in part associated with second interstrial space, spots located behind umbone, in front of middle, behind middle, at apical fourth and at about apical sixth, spots of discal row located inside umbone, at basal fourth, at about basal third, in front of apical third and behind apical fourth; head and pronotum clothed with moderately long, recumbent white hairs that are replaced by closely appressed digitate scales along sides of pronotum; elytra clothed with short, suberect white hairs that become stouter and parallel-sided behind middle and stouter and apically forked along sides; ventral surface densely clothed with short, recumbent digitate scales that become longer but more sparsely placed and subrecumbent along sides of thoracic sternites. Head coarsely and rather densely, shallowly punctured; vertex not carinate; front convex except for a distinct transverse depression above clypeus; clypeus finely, densely punctured, front margin very shallowly, broadly, arcuately emarginate; antenna black, extending to slightly beyond middle of pronotum, serrate from fifth segment, outer segments distinctly wider than long. Pronotum one and one-half times wider than long, widest in front of middle, slightly wider than elytra, convex, without depressed areas, basal pits indistinct; sides evenly arcuate; lateral margins not reflexed or carinate, not visible from above; front angles very feebly produced, narrowly rounded; front margin very shallowly lobed at middle; hind margin feebly emarginate; surface coarsely, densely and deeply punctured at middle, punctures becoming coarser and more densely placed toward sides. EIytra convex; sides when viewed from above subequal in width at base and middle, broadly, shallowly constricted at about basal fourth, broadly, evenly arcuate from middle to the conjointly, broadly rounded apices; front angles in side view strongly produced ventrally and slightly produced anteriorly, acute; lateral margins when viewed from the side rather deeply emarginate at about basal sixth, then very broadly and very shallowly arcuate to apices, serrate on

46 190 William F. Burr about apical sixth; stria1 punctures moderate in size and deep along suture, becoming slightly larger laterally, larger and more irregular at base and smaller toward apices, striae distinctly impressed behind apical third; interstrial spaces subconvex, slightly roughened at sides, interstrial punctures distinct, third interstrial space slightly elevated between umbone and middle, suture slightly elevated behind middle. Ventral surface coarsely, rather densely punctured on thoracic sternites and basal half of first abdominal sternite, more finely, densely punctured elsewhere; front margin of prosternum not retracted, separate from front angles of pronotum, distinctly reflexed at sides, broadly, arcuately emarginate with a small rounded tooth on either side of middle; last visible abdominal sternite with sides not reflexed, hind margin feebly rounded, narrowly reflexed, disk slightly convex without a subapical plate or elevation, but with a subapical concentration of small, densely placed punctures. Length: 6.8 mm. Width: 2.1 mm. Holotype male (California Academy of Sciences) and 4 paratypes from Globe, Arizona, June 9, 1938 (F. H. Parker). Additional paratypes from type locality, 2 specimens, June 15, 1959 and June 26, 1950 (F. H. Parker). Paratypes from other localities in Arizona, are: 19 specimens from Icehouse Canyon, near Globe, Pinal Mountains, June 3, 1962 (D. S. Verity) and June 16, 1961 (R. L. Westcott); 9 specimens from Pinal Mountaisn, June 16, 1935 (F. H. Parker) and July 3, 1941 (W. F. Barr); 2 specimens from Sabino Canyon, Tucson, May 16, 1953 (A. and H. Dietrich); 4 specimens from Tucson, May 17, 1953 (A. and H. Dietrich) and June 23, 1939 (D. J. and J. N. Knull); 16 specimens from Madera Canyon, Santa Rita Mountains, June 15, 1961, June 19, 1959 and July 19-23, 1957, on mesquite (R. L. Westcott); 1 specimen from Nogales, May 26, 1950, on Mimosa (F. H. Parker); and 3 specimens from Organ Pipe National Monument, Pima County, June 10, 1952 (M. Cazier, W. Gertsch and R. Schrammel). Paratypes in the collections of the American Museum of Natural History, California Academy of Sciences, University of California at Riverside, Cornell University, F. M. Beer, D. S. Verity, R. L. Westcott and W. F. Barr. The coarsely, densely punctured pronotum and the two rows of spots on each elytron will separate this species from all others in the guttifera group except A. varipilis Van Dyke from southern Baja California. A. paravaripilis is distinguished from that species by having short, suberect pointed hairs on the elytral disk, by having the front of the prosternum flattened and by lacking a subapical elevation on the last abdominal sternite.

47 New species of North American Acmaeodera 191 Acmaeodera vaga Barr, new species (Figure 23) Male: Small-sized, subcylindrical, feebly shining, black with a bluish luster; each elytron with a discal row of yellow spots located at about middle, at about apical fourth and at about apical sixth, median spot moderate in size, others progressively smaller; head moderately clothed with stout, recumbent white hairs that become more dense above clypeus; pronotum sparsely clothed with short, recumbent, pointed white hairs on disk and moderately clothed with appressed digitate scales near sides of front margin and along lateral margins; elytra moderately clothed with short, blunt suberect and subrecumbent hairs, the hairs are parallel-sided on basal half of disk and spatulate on apical half, hairs on basal half near lateral margins tending to be in the form of compact recumbent digitate scales; ventral surface rather densely clothed with broad, appressed finely branched scales that become more elongate and hair-like along sides of metasternum and abdominal sternites, surface not obscured. Head coarsely and rather densely, very shallowly punctured, punctures becoming finer near eyes and on vertex; vertex with a faint indication of a short, median longitudinal carina; front convex on upper portion, subflattened on lower portion and with a distinct transverse depression above clypeus; clypeus finely, sparsely punctured, somewhat roughened, front margin rather deeply, broadly, arcuately emarginate; antenna black, extending to slightly beyond middle of pronotum, serrate from fifth segment, outer segments subequal in width and length. Pronotum about one and one-half times wider than long, widest at middle, narrower than elytra, convex except narrowly subflattened on disk; basal pits distinct; sides evenly arcuate; lateral margins not reflexed, feebly carinate throughout, not visible from above; front angles feebly produced, subrectangular; front margin broadly arcuate; hind margin broadly emarginate; surface rather coarsely and densely punctured on anterior half of disk, more finely, sparsely and shallowly punctured on posterior half of disk, very coarsely, densely punctured laterally. Elytra convex; sides when viewed from above subequal in width behind base and at middle, distinctly expanded immediately behind base, broadly constricted at about basal fourth, subparallel medially and then broadly arcuate to the conjointly, very broadly rounded apices; front angle in side view slightly produced ventrally and anteriorly, acute; lateral margin when viewed from the side very broadly, shallowly arcuate to apices, serrate on about apical fourth; stria1 punctures small and deep along suture and on apical half of disk, becoming coarser and more densely placed basally and laterally, striae deeply impressed behind

48 192 William F. Barr middle; interstrial spaces subflattened on basal half of disk, subconvex elsewhere, becoming transversely roughened near sides, interstrial punctures small and indistinct, third interstrial space elevated on basal half; suture distinctly elevated behind basal fourth. Ventral surface rather coarsely, densely punctured except on abdominal sternites two to five which are finely, sparsely and shallowly punctured; front margin of prosternum very slightly retracted, separated from front margin of pronotum, narrowly reflexed laterally, shallowly emarginate with a large rounded tooth on either side of middle; last abdominal sternite with lateral margins nearly straight, oblique, hind margin semicircularly rounded, narrowly reflexed, disk feebly convex with a faint indication of a transverse subapical elevation. Length: 6.7 mm. Width: 2.0 mm. Holotype male (California Academy of Sciences) and 3 paratypes from 41 miles north of San Luis Potosi, San Luis Potosi, June 26, 1965, on Prosopis chilensis (G. H. Nelson). Paratypes in the collections of G. H. Nelson and W. F. Barr. The blue-black coloration and subrecumbent stout hairs of the elytra and its relatively large size for a member of the guttifera group make this species easily recognizable. Its affinities are with A. hulli Knull and A. chisosensis Knull. Acmaeodera peninsularis Barr, new species (Figure 24) Female: Small-sized subcylindrical, robust, blackish with a strong brassy luster, moderately shiny; each elytron with a sublateral row of small yellow and red spots and a discal row of small yellow spots, the sublateral row located on third interstrial space with spots situated behind umbone in front of basal fourth, at about middle, in front of apical third, at about apical fourth, and in front of apex, the two apical spots elongate and reddish, the three anterior spots yellow, spots of the discal row situated in front of basal fourth, behind basal third, in front of apical third, and at about apical fourth, the third spot of this row more or less transverse, broken on the left elytron; upper surface moderately clothed with small appressed scales that are longer than wide, widened apically and which are sparsely intermixed with very fine, short, nearly inconspicuous recumbent hairs on head and pronotum and very short, stout suberect hairs on elytra; ventral surface densely clothed with elongate, appressed branched scales that obscure

49 New species of North American Acmaeodera 193 surface except in areas where they have been rubbed off, abdominal sternites three to five densely clothed with long, suberect and recurved white hairs. Head rather coarsely and densely punctured; vertex without a longitudinal carina; front feebly concave at middle, more deeply depressed above clypeus; clypeus finely, densely punctured, front margin very deeply, arcuately emarginate; antenna extending to slightly beyond pronotum, serrate from fifth segment, outer segments slightly wider than long. Pronotum approximately one and three-fourths times wider than long, widest slightly behind middle, narrower than elytra, strongly convex, feebly depressed medially at base and on disk and in front of the indistinct basal pits; transverse depression behind front margin not evident; sides strongly, evenly arcuate; lateral margins not reflexed, carinate throughout, not visible from above; front angles not produced, narrowly rounded; front margin strongly lobed at middle; hind margin very feebly and broadly emarginate; surface rather coarsely, densely and deeply punctured at middle, punctures becoming more densely placed toward sides. Elytra convex; sides when viewed from above slightly wider behind base than at middle, distinctly expanded immediately behind base, broadly constricted at about basal fourth and then broadly and evenly arcuate to the conjointly, broadly rounded apices; front angles in side view strongly produced ventrally and slightly produced anteriorly, acute; lateral margins when viewed from the side rather deeply emarginate at about basal fifth then broadly, shallowly arcuate to apices, rather finely serrate on apical fourth; stria1 punctures moderate in size, deep and somewhat elongate, uniform throughout, striae deeply impressed behind middle; interstria1 spaces appearing alternately subconvex and subflattened, transversely wrinkled and roughened, interstrial punctures distinct, numerous, appearing in two rows on the tenth and eleventh interstrial spaces, third interstrial space strongly elevated throughout, fifth interstrial space slightly elevated apically, seventh and ninth interstrial spaces slightly elevated throughout, suture feebly elevated behind middle. Ventral surface very coarsely, densely punctured on sides of thoracic sternites and basal half of first abdominal sternite, more sparsely punctured medially, finely, densely and shallowly punctured on apical half of first abdominal sternite and on second and third abdominal sternites, microreticulate, slightly roughened and finely punctured on fourth and fifth abdominal sternites; front margin of.prosternum distinctly retracted, slightly separated from front margin of pronotum, not reflexed, broadly emarginate with a broad rounded tooth on either side of middle, prosternum transversely ridged medially immediately behind front margin; last visible abdominal sternite with hind margin broadly but very feebly rounded, sides not reflexed, disk feebly convex with a very broad,