LYSMATA OLAVOI, A NEW SHRIMP OF THE FAMILY HIPPOLYTTDAE (DECAPODA, CARIDEA) FROM THE EASTERN ATLANTIC OCEAN*

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1 CRUSTACEA LIBRARY SMITHSONIAN INSTITUTION RETURN TO W-119 LYSMATA OLAVOI, A NEW SHRIMP OF THE FAMILY HIPPOLYTTDAE (DECAPODA, CARIDEA) FROM THE EASTERN ATLANTIC OCEAN* C.H.J.M. FRANSEN A DAI TIDE I kflf) FRANSEN, C.HJ.M Lysmt olvoi, new shrimp of the fmily AivyUlILLAVJvy Hippolytide (Decpod, Cride) from the estern Atlntic Ocen.- Arquipelgo. Life nd Erth Sciences 9: Angr do Herofsmo. ISSN A new species of Hippolytide, Lysmt olvoi, is described nd figured. The species ws found off Pico islnd, Azores nd Slvge islnds, Mdeir rchipelgo m depths between 135 nd 360 m. FRANSEN, C.HJ.M Lysmt olvoi, um nov especie d fmfli Hippolytide (Decpod, Cride) do Oceno Atlntico orientl.- Arquipelgo. Lite nd Erth Sciences 9: Angr do Herofsmo. ISSN Descreve-se e ilustr-se nov esp6cie de Hippolytide, Lysmt olvoi. Est esp6cie foi recolhid perto d cost d ilh o Pico, Azores, e ds Uns Selvgens, Arquipelgo d Mdeir, em profundiddes que vrirm entre 135 e 360 m. C.HJ.M. Frnsen, Ntionl Ntuurhistorisch Museum (Rijksmuseum vn Ntuurlijke Historie), Postbus 9517, 2300 RA Leiden, The Netherlnds. INTRODUCTION In October 1989 Helen R. Mrtins visited the Ntionl Ntuurhistorisch Museum (Rijksmuseum vn Ntuurlijke Historie) in Leiden nd showed Prof. L.B. Holthuis nd the uthor three femle specimens of n unknown Lysmt, collected in bited trps (see MARTINS & HARGREAVES 1991) set in deep wter off the Azores during tril fishery for shrimps crried out by the Secretrit of Agriculture nd Fisheries of the Regionl Government of the Azores with the M/V "Ptro Pedro" in An dditionl 21 specimens were cught during similr survey in 1989 with M/V "Perol do Fil". Two further specimens were obtined from The Nturl History Museum (British Museum (Nturl History)), lso from the M/V "Ptro Pedro" survey in In the CANCAPcollections of the Ntionl Ntuurhistorisch Museum three mle specimens of the sme species re present, they were found with deep wter trps off the Slvge islnds (Ilhs Selvgens). The CANCAP-project ws mrine biologicl reserch project crried out in the CANrin - CAPe Verdin region of the North * CANCAP-Contribution 96. Atlntic Ocen from 1976 to 1987 (VAN DER LAND 1987). All these specimens re described below s new species, Lysmt olvoi, nmed in honour of the lte Olvo Amrl, ssistnt t the Deprtment of Ocenogrphy nd Fisheries of the University of the Azores, who ws on bord the "Ptro Pedro" nd "P6rol do Fil" to tke subsmples of mteril for the University of the Azores. Rijksmuseum vn Ntuurlijke Historie is bbrevited s RMNH; postorbitl crpce length s pocl. Lysmt olvoi new species (Figs. 1-34) Mteril. - RMNH D 38484: M/V Ptro Pedro St. 8; Azores, Pico, Pont d Ilh, 38 25'00"N 27 59'10"W; depth 135 m; bited trps; 8.ii.l988; ovigerous femle, pocl mm (holotype). - BMNH Crust. 1989:935:1: M/V Ptro Pedro St. 8; Azores, Pico, Pont d Ilh, 38 25'00"N 27 59'10"W; depth 135 m; bited trps; 8.ii.l988; ovigerous femle, pocl mm (prtype). - RMNH D 38485: M/V Ptro Pedro St. 6; Azores, Pico, Ribeirs, 38 24'20"N 28 08'00"W; depth 360 m; bited trps; 6.ii.l988; 2 ovigerous 63

2 femles, pocl nd 13.4 mm (prtypes). - BMNH Crust. 1989:933:1: M/V Ptro Pedro St. 4; Azores, Pico, Ribeirs, 38 24'20"N 28 08'00"W; depth 126/144 m; bited trps (string 5(2)); 3.ii.l988; 1 ovigerous femle, pocl mm (prtype). - RMNH D 38487: M/V Per61 do Fil; Azores, Pico, Ribeirs; depth 160 m; 24.xi.1989; 3 mles, 8 femles of which 7 ovigerous, pocl mm (prtypes). - RMNH D 40156: M/V Perol do Fil; Azores, Pico, St. Cruz ds Ribeirs; depth 183 m; bited trps (string 6); 23.xi.1989; 1 mle nd 2 ovigerous femles, pocl mm (prtypes). - RMNH D 40157: M/V Per61 do Fil; Azores, Pico, Ribeirs; depth 190 m; bited trps (string 1); 23.xi.1989; 1 mle, pocl mm (prtype). - RMNH D 40155: M/V Perol do Fil; Azores, Pico, St. Cruz ds Ribeirs; depth 146 m; bited trps (string 5); 24.xi.1989; 3 mles nd 3 ovigerous femles, pocl mm (prtypes). - RMNH D 38486: St. CANCAP 4.V08; Slvge islnds, S of Selvgem Grnde; 30 07'N 15 53'W; depth 260 m; fish-trp, shrimp-trp, 4 plstic sniltrps; overnight 26/27.v.1980; 3 mles, pocl. 6.7, 9.2 mm (prtypes), nd 9.2 mm (llotype). Description of femle holotype. Rostrum (Figs. 1-3) reching s fr s second segment of ntennulr peduncle. Dorsl mrgin lmost stright with six evenly spced teeth, posterior three postorbitl. Ventrl mrgin with three prominent teeth. Lterl crin equidistnt from ech mrgin of rostrum, fusing with posterior mrgin of orbit. Antennl spine robust nd shrp. Anteroventrl mrgin with smll pterygostomin tooth. Tegument pitted, with smll scle-like hirs. Abdomen (Fig. 1) with pleur of first three somites rounded, tht of fourth blunt with tiny tooth, fih pleuron shrply cute. Sixth somite 1.5 times s long s fih. Telson 1.3 times s long s sixth bdominl somite. Two pirs of dorsl spines on telson (Fig. 4); nterior pir t 0.43, posterior t 0.68 of the telson length. Lterl mrgins cilited from level of posterior pir of dorsl spines to distl end. Posterior mrgin of telson (Fig. 5) with medin tooth nd two pirs of spines. Lterl pir of spines smll, mesil pir five times s long s lterl pir; two long sete present between two mesil spines; four short sete present on both sides of medin tooth of telson. Eyes with well developed corne; corne broder thn eyestlk nd bout s long. Between the eyes protrusion is present ('bee ocellir') nteriorly pointed nd with posterodorsl mrgin oblique, directed into concve posteroventrl mrgin of rostrl ventrl lmin (Fig. 34). Antennulr peduncle (Figs. 2, 3, 6) with stylocerite tpering to shrp tip, not reching end of bsl segment, inner mrgin stright, outer mrgin slightly concve. Ech ntennulr segment rmed with one to three short distl dorsl spines. Bsl segment longest, with smll mesil ventrl tooth ner midlength. Second segment bout hlf length of bsl segment. Third segment bout two thirds length of second segment. Antennulr flgell equl in length, c. 3.5 times the crpce length; le nd right dorso-lterl flgellum with 53 nd 55 rticles respectively, in thickened, setiferous bsl portion; distl setiferous rticle (Fig. 7) with vestigil ccessory flgellum. Scphocerite (Fig. 8) bout 4.5 times s long s wide; lterl mrgin concve; distl tooth shrp, overreching distl mrgin of lmin. Antennl peduncle short, not reching end of proximl fourth of scphocerite; bsl segment with outer lterl tooth ner bse of scphocerite. Mndibles simple, very unequl in form. Molr process of le mndible (Fig. 9) with three blunt lobes nd n re with short sete. Right mndible (Figs. 10, 11) with six blunt lterl lobes nd n re with short sete. First mxill (Fig. 12) with robust upper nd lower endite; plp unsegmented, bilobed. Second mxill (Fig. 13) with lower endite strongly reduced, upper endite divided into two lobes, upper better developed thn lower; plp well developed. Scphognthite with rounded posterior lobe. First mxilliped (Fig. 14) with endites of bsis nd cox seprted by notch; plp slender, three-segmented; criden lobe well developed; epipod lrge, bilobed. Second mxilliped (Fig. 15) with wide ultimte segment; penultimte segment oblong; 64

3 epipod with podobrnch. Third mxilliped (Fig. 16) long nd slender reching beyond first pereiopod; ultimte segment (Fig. 17) with nine spines in distl third; penultimte segment 0.6 times s long s ultimte segment; bsl segment 2.5 times s long s penultimte segment; exopod short, bout hlf length of bsl segment; epipod well developed. Brnchil formul s in Tble 1. Tble 1. Brnchil formul in Lysmt olvoi rx.1 mx.2 mx.3 per.l per.2 per.3 per.4 per.5 pleurobrnchs rthrobrnchs - 2 podobrnchs - 1 epipods exopods Four nterior pereiopods with welldeveloped epipods. First pereiopods (Fig. 18) reching beyond scphocerite by hlf propodus length; fingers hlf s long s plm, cutting edges (Fig. 19) entire; crpus slightly longer thn chel, oblong, distl prt with setiferous clening orgn (Fig. 20); merus lmost 1.2 times crpus length; ischium short with row of short spinules on posterior mrgin. Second pereiopods (Fig. 21) subequl in shpe nd size, merus of right reching to end of third mxilliped, merus of le reching to distl end of ntennulr peduncle; chel minute, simple; crpus s long s merus + ischium, le composed of 34 rticles nd right of 35; merus longer thn ischium, le composed of 18 rticles nd right of 22; ischium of le pereiopod with five rticles nd of right with three rticles in distl prt, both with row of curved flttened spinules on posterior mrgin; bsis with three curved flttened spinules on posterior mrgin. Third pereiopod (Fig. 22) overreching scphocerite by length of dctylus, propodus nd hlf crpus; dctylus (Fig. 23) terminting in spine-like tooth continuous with extensor mrgin nd stouter tooth on flexor side, with four movble spines on flexor mrgin; propodus 6.5 times s long s dctylus, rmed with 14 slender spinules on flexor mrgin; crpus 0.9 times propodus length, rmed with nine spinules on flexor mrgin; merus 1.75 times crpus length, rmed with seven stout spines on ventrl nd lterl mrgins of le nd eleven on right pereiopod; ischium short, 0.25 times merus length, without rmture. Fourth pereiopod (Figs. 24,25) overreching scphocerite by dctylus + propodus, similr in shpe to third pereiopod; merus with eight stout spines on ventrl nd lterl mrgin of le nd seven of right pereiopod. Fih pereiopod (Figs. 26, 27) overreching scphocerite by dctylus + one-hlf of propodus, similr in shpe to third nd fourth pereiopods; merus rmed with one ventrl distl stout spine; flexor mrgin of propodus distlly setiferous, sete in oblique rows. Distl end of lterl mrgin of outer brnch of uropod rmed with two fixed teeth flnking single longer movble spine. Mny eggs, 1 x 0.5 mm. Colour. The following colour notes were tken from colour slide of mteril of St. Cruz ds Ribeirs, Pico, Azores. Eyes blck. Thorcic ppendges bright red. Abdomen red with eight longitudinl white stripes, lterlmost two pirs of stripes extending hlfwy onto the yellowish brown crpce, becoming indistinct nteriorly. Posterior prts of bdominl tergites deeper shde of red thn nterior prts, forming red trnsversely bnded pttern superimposed upon the longitudinl pttern. Pleopods with red endopods nd exopods, other prts white. Telson nd uropods red. Eggs green. Description of mle llotype. Generl ppernce s of femle, slightly more slender (Fig. 28). Pleur of fourth bdominl segment with more distinct posterior tooth. Endopod of first pleopod (Fig. 29) tpering to slender end-piece bering cluster of coupling pds. Appendix msculin (Fig. 30) on endopod of second pleopod s long s ppendix intern, rmed with four distl sete. Remrks. The specimens studied show minor vrition (Tble 2). In prticulr the denttion of the rostrum (Figs. 1, 2, 28, 31-33), the spintion of the mbultory pereiopods, nd the number of rticles composing the crpus, merus nd ischium of the second pereiopods. The number of rticles in the setiferous prt of the dorso-lterl flgellum of the ntennule seems 65

4 Tble 2. Vrition in chrcters in Lysmt olvoi specimens. - ^ K; s; 00 S2 I & g Registrtion * 5 <. <3 c,1s.. 3 Q I 1 s: R st? I * I <u ^ -SP. A c "5 I rs ^ S. -2? 5 S 6 & 5. X3., -2 <3 fe.. -e tic 3 I S 5 W <>5 S S 5 5 ^ 5 S v. W. xs R <5..2 S I 1 1..O QJ k o -52 t! t! S3 <u. xs K; o c 5 v! si * ^.. s» I * S o yn t\j s 3 VJ t! «RMNH D m RMNH D m RMNH D m RMNH D fov RMNH D fov RMNH D fov RMNH D fov RMNH D f RMNH D fov 3 17.S RMNH D fov RMNH D fov BM1989:933:1 12 fov BM1989:935:1 13 fov RMNH D fov RMNH D fov RMNH D fov RMNH D m RMNH D m RMNH D m * m=mle, f=femle, fov= ovigerous femle. 66

5 correlted with the size of the specimen. The second pereiopods re lwys subequl in size nd no length dominnce for the le or right pereiopod ws pprent. Length nd number of rticles of the crpus of the second pereiopod is not correlted with size. The pleur of the fih bdominl segment in femles (especilly ovigerous femles) hve very indistinct posterior tooth, in mles this tooth is more prominent. Systemtic position. Ten species of Lysmt re known from the Atlntic: L. nchisteus Chce, 1972; L. grbhmi (Gordon, 1935); L. intermedi (Kingsley, 1878); L. moorei (Rthbun, 1901); L. nilit Dohrn & Holthuis, 1950; L. rthbune Chce, 1970; L. seticudt (Risso, 1816); L. stenolepis Crosnier & Forest, 1973; L. uncicornis Holthuis & Murin, 1952; L. wurdemnni (Gibbes, 1850). L. nchisteus is known from Puerto Rico to Grnd, western Atlntic in depths less thn 3 meters. It differs from L. olvoi in the scphocerite not overreching the ntennulr peduncle; the presence of only one postorbitl dorsl rostrl tooth; the presence of only one ventrl rostrl tooth; the bsence of pterygostomin tooth; the pleur of the fourth bdominl pleur being posteriorly rounded. L. olvoi ws compred with mteril of L. nchisteus from Curso, Crriben Se (RMNH D 30376) nd the type-description (CHACE 1972). L. grbhmi is known from tropicl nd subtropicl regions in the western, centrl nd estern Atlntic in depths to bout 60 m. It differs from L. olvoi by the scphocerite not overreching the ntennulr peduncle; the rostrum nd ntennulr peduncle being longer in reltion to the crpce length; in colourtion: L. grbhmi hving two bright red dorsolterl longitudinl bnds seprted by white dorsl bnd. L. olvoi ws compred with specimens of L. grbhmi from the Cnry Islnds (RMNH D 40154) nd the type-description by GORDON (1935). L. intermedi Kingsley, 1878 is shllow wter species known from depths to 22 m, originlly described from the Dry Tortugs, Florid, nd figured by Kingsley in The species is recorded in the western Atlntic from Bermud to Brzil, in the centrl Atlntic from Ascension (MANNING & CHACE 1990: 23) nd in the estern Atlntic from the Azores (RATH- BUN 1901: 116); in the estern Pcific it ws recorded from the Gulf of Cliforni to Peru. It differs from L. olvoi in hving the ccessory brnch of the dorso-lterl ntennulr flgellum well developed; the stylocerite reching nerly to, or beyond the distl mrgin of the bsl segment of the ntennulr peduncle. L. olvoi ws compred with mteril of L. intermedi from Arub, Cribben (RMNH D 4767) nd the description nd figures given by SLVERTSEN (1933). RATHBUN (1901:116) recorded L. intermedi from Pim By, Fyl, Azores (Lewis Dexter coll.). The lrge femle specimen ws reexmined by MANNING & CHACE (1990: 23) nd found to be correctly identified. L. moorei (Rthbun, 1901) is known from the tropicl western, centrl nd estern Atlntic occurring in tide-pools nd the littorl zone. L. moorei differs from L. olvoi in hving only one postorbitl dorsl rostrl tooth; in the bsence of pterygostomin tooth; in the shorter nd more robust ntennulr peduncle; in hving the ccessory brnch of the dorso-lterl ntennulr flgellum well developed. L. olvoi ws compred with mteril of L. moorei from Arub, Cribben (RMNH D 4768) nd the description nd figures given by RATHBUN (1901). L. nilit Dohrn & Holthuis, 1950 is known from the estern Atlntic (Cnry Islnds) nd the Mediterrnen, occurring in shllow wter. It differs from L. olvoi by the stylocerite overreching the first segment of the ntennulr peduncle; by the ccessory brnch of the dorsolterl ntennulr flgellum being well developed; by the more robust mbultory pereiopods; by the colourpttern composed of brod drk ornge bnds cross the body. L. olvoi ws compred with the holotype nd prtypes of L. nilit from the western prt of the By of Npels, Mediterrnen (RMNH D 7258) nd the description nd figures of DOHRN & HOLTHUIS (1950). L. rthbune Chce, 1970 is known from estern Florid to Yuctn, western Atlntic, in depths between 37 nd 119 m. It differs from L. 67

6 olvoi by the rostrum overreching the ntennulr peduncle; the pterygostomin tooth being usully bsent; the stylocerite being less thn hlf the length of the bsl segment of the ntennulr peduncle; the pleur of the fourth bdominl segment being postero-lterlly rounded. L. olvoi ws compred with the type-description of L. rthbune by CHACE (1970). L. seticudt is known from the Mediterrnen nd the estern Atlntic from Morocco to the British Isles, occurring in depths to bout 15 m. It differs from L. olvoi in the stylocerite overreching the bsl segment of the ntennulr peduncle; in hving the ccessory brnch of the dorso-lterl ntennulr flgellum well developed; in the scphocerite being twice s long s the ntennulr peduncle; in the colourpttern composed of longitudinl red stripes on the body. L. olvoi ws compred with mteril from the By of Nples, Mediterrnen (RMNH D 14274). In 1973 CROSNIER & FOREST described L. stenolepis from deep wter off So Tigo, Cpe Verde Islnds. One mle (pocl 4.5 mm) ws cught t depth between 275 nd 150 m by the Tlismn in The specimen differs from L. olvoi in the denttion of the rostrum. In L. stenolepis the rostrum is lmost without ventrl lmin nd bers only one smll ventrl tooth. In the mteril of L. olvoi mles nd femles of different sizes hve three to five ventrl rostrl teeth. The number of fused segments of the ntennule dorso-lterl flgellum number 17 in L. stenolepis, much fewer thn in L. olvoi. This however could be correlted with body size. The pleur of the fourth bdominl segment in L. stenolepis hve more cute postero-lterl pex thn in L. olvoi. The specimen of L. stenolepis is incomplete, ll mbultory pereiopods re missing, nd it is of much smller size which mkes comprison with L. olvoi difficult. More mteril of L. stenolepis from the type-loclity hs to be collected to estblish the vribility of the vrious chrcters. L. uncicornis is known from the estern Atlntic from Morocco to Congo, occurring in shllow wters. L. uncicornis differs from L. olvoi in the bsence of the pterygostomin tooth; the bsence of the postero-lterl tooth on the fourth bdominl pleur; the broder scphocerite. L. olvoi ws compred with the holotype nd prtypes of L. uncicornis from the hrbour of Csblnc, Morocco (RMNH D 7812), nd the type-description of HOLTHUIS & MAURIN (1952). L. wurdemnni (Gibbes, 1850) is known from the western Atlntic from Virgini to Estdo de So Pulo, Brzil, occurring in depths to 30 m. L. wurdemnni differs from L. olvoi in hving the pleur of the fourth bdominl segment postero-lterlly rounded; in the bsence of the pterygostomin tooth; in hving the mbultory pereiopods more robust. L. olvoi ws compred with mteril of L. wurdemnni from French Guyn (RMNH D 12643). Lysmt olvoi is deep wter species. The other Atlntic Lysmt, except L. stenolepis, seem littorl or sublittorl but hve not been found deeper thn bout 120 m. ACKNOWLEDGEMENTS The uthor thnks Helen Mrtins, Universidde dos Azores, for llowing me to study her mteril nd for the vluble comments on the mnuscript; Luis Monteiro, Universidde dos Azores, for providing colour photogrphs; Prof. Dr. L.B. Holthuis, Ntionl Ntuurhistorisch Museum, Leiden, nd Dr. R. W. Ingle, The Nturl History Museum, London, for kindly criticizing the mnuscript. REFERENCES CHACE, F.A A new shrimp of the genus Lysmt (Decpod, Hippolytide) from the western Atlntic. - Crustcen 19(1): 59-66, figs CHACE, F.A The shrimps of the Smithsonin-Bredin Cribben Expeditions with summry of the West Indin shllow-wter species (Crustce: Decpod: Ntnti). - Smithsonin Contributions to Zoology 98: i-x, 1-179, figs CROSNIER, A. & J. FOREST Les crevettes profondes de l'atlntique Orientl Tropicl. - Fune Tropicle 19:1-409, figs

7 GORDON, On new or imperfectly known species of Crustce Mcrur. - Journl of the Linnen Society, Zoology 39: , Figs HOLTHUIS, L.B. & C. MAURIN Note sur Lysmt uncicornis nov. spec, et sur des utres espfcces intdressntes de crustc6s d6cpodes mcroures de l cote tlntique du Mroc. - Proceedings of the Koninklijke Nederlndse Akdemie vn Wetenschppen, ser. C, 55(2): , figs. 1,2. KINGSLEY, J.S Notes on the North Americn Cride in the museum of the Pebody Acdemy of Science t Slem, Mss. - Proceedings of the Acdemy of Nturl Sciences of Phildelphi, 1878: KINGSLEY, J.S Crcinologicl Notes; Number V. - Bulletin of the Essex Institute 14: , pis LAND, J. VAN DER Report of the CAN- CAP-project for mrine biologicl reserch in the Cnrin - Cpe Verden region of the North Atlntic Ocen ( ). Prt I. List of sttions. - Zoologische Verhndelingen, Leiden 243:1-94, figs. 1,2. MANNING, R.B. & F.A. CHACE Decpod nd Stomtopod Crustce from Ascencion Islnd, South Atlntic Ocen. - Smithsonin Contributions to Zoology 503: i-v, 1-91, figs MARTINS, H.R. & P.M. HARGREAVES Shrimps of the fmilies Pndlide nd Hippolytide (Crustce: Decpod) cught in the benthic trps off the Azores.- Arquipilgo. Life nd Erth Sciences 9: RATHBUN, M.J The Brchyur nd Mcrur of Porto Rico. - Bulletin of the United Sttes Fish Commission, 1900, 20(2): 1-127, figs Accepted 4 My Fig. 1. Lysmt olvoi new species, hbitus femle holotype, pocl mm. Scle 8 mm. 69

8 Figs Lysmt olvoi new species, femle holotype, pocl mm. 2, crpce nd nterior ppendges, lterl spect; 3, nterior ppendges, dorsl spect; 4, telson nd uropods, dorsl spect; 5, distl prt telson, dorsl spect; 6, le ntennulr peduncle, ventrl spect; 7, distl end of setiferous prt le ntennulr dorsolterl flgellum; 8, ntennl peduncle nd scphocerite, ventrl spect. (Scle: 2-4,6,8 = 4 mm; 5,7 = 1 mm). 70

9 Figs Lysmt olvoi new species, femle holotype, pocl mm. 9, le mndible; 10, right mndible, ventrl spect; 11, right mndible, dorsl spect; 12, first mxill; 13, second mxill; 14, first mxilliped; 15, second mxilliped; 16, third mxilliped; 17, first mxilliped, distl prt. (Scle 9-15 = 2 mm; 16 = 4 mm; 17 = 1 mm). 71

10 Figs Lysmt olvoi new species, femle holotype, pocl mm. 18, first le pereiopod; 19, first le pereiopod, chel; 20, first pereiopod, setiferous clening orgn on distl prt crpus; 21, second le pereiopod; 22, third le pereiopod; 23, third le pereiopod, dctylus; 24, fourth le pereiopod; 25, fourth le pereiopod, dctylus; 26, fih le pereiopod; 27, fih le pereiopod, dctylus. (Scle: 18,21,22,24,26 = 4 mm; 19,20,23,25,27 = 1 mm). 72

11 Figs Lysmt olvoi new species , mle llotype, pocl. 9.2 mm. 28, hbitus; 29, endopod of first right pleopod; 30, ppendix msculin nd ppendix intern of second right pleopod; 31, prtype mle, pocl. 6.7 mm, rostrum, lterl spect; 32, prtype mle, pocl. 9.2 mm, rostrum, lterl spect; 33, prtype femle, pocl mm, rostrum, lterl spect; 34, holotype femle, pocl mm, nterior prt, lterl spect - 'bee ocellire', le eye nd le ntennul dissected. (Scle: 28 = 8 mm; 29, 30 = 1 mm; = 2 mm).. 73

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