Nursehound. Scyliorhinus stellaris NE ATL MED SYT. Lateral View ( ) Ventral View ( ) COMMON NAMES

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Lateral View ( ) Ventral View ( ) APPEARANCE Two dorsal fins without spines. First dorsal fin larger than second, originates over pelvic fins. Second dorsal fin originates over or slightly behind anal fin. SYNONYMS Almost straight caudal fin with large ventral lobe. Squalus stellaris (Linnaeus, 1758), Scyllium catulus (Müller & Henle, 1838), Scyllium acanthonotum (Filippi and Verany, 1853), Scyliorhinus besnardi (Springer & Sadowsky, 1970). Nasal furrows do not reach mouth. 162cm maximum total length. Common to 130cm Creamy brown dorsally with numerous dark spots. Occasionally also white spots. White ventrally. The is known in the northeast Atlantic from southern Scandinavia and the British Isles to at least Morocco, including the Mediterranean Sea. Its presence in tropical west Africa from Senegal to Zaire is uncertain No Records and may be due to Occasional misidentifications Range of the West African Catshark, Scyliorhinus cervigoni (Compagno, 1984). Map base conforms with ICES grid squares. A large catshark which can reach up to 160cm in length, the can be found throughout the northeast Atlantic and Mediterranean. The head is moderately short and broad. On the underside there no nasoral grooves and labial furrows on the lower jaw only. The small anterior nasal flaps do not reach the mouth. The pectoral fins are relatively large. The first dorsal fin is set well back along the body, above the pelvic fins. The second dorsal fin is set above the anal fin. There are no dorsal fin spines. The caudal fin is long and almost straight with a well developed ventral lobe (Compagno, 1984). Colouration is pale brown dorsally, white ventrally. There is a pattern of numerous large and small dark/black spots and occasionally white spots on the back. It grows to a maximum size of 162cm, although it is more commonly found to a maximum of 130cm (Compagno, 1984). NT DISTRIBUTION SYT, Bull Huss, Greater Spotted Catshark, Greater Spotted Dogfish, Flake, Rigg, Grande Roussette (Fr), Alitán (Es). NE ATL MED COMMON NAMES

Supported by: SIMILAR SPECIES Scyliorhinus canicula, Small Spotted Catshark Galeus melastomus, Blackmouth Catshark Galeus atlanticus, Atlantic Sawtail Catshark Galeus murinus, Mouse Catshark, Scyliorhinus canicula, Smallspotted Catshark Galeus melastomus, Blackmouth Catshark Galeus atlanticus, Atlantic Sawtail Catshark Galeus murinus, Mouse Catshark (Not to scale)

TEETH There are 44 56 teeth in the upper jaw, including 0 2 symphysial teeth and 38 46 teeth in the lower jaw, including 2 4 symphysial teeth. The front teeth are single-cusped and erect, further back in the jaw they develop cusplets (Soldo et al., 2000). ECOLOGY AND BIOLOGY HABITAT The can be found to 125m but is most common around 20-63m. It seems to have a preference for rough/rocky ground or areas with good algal cover. It is locally abundant in some places and is regularly encountered by divers, though not as commonly as the Small Spotted Catshark, Scyliorhinus canicula (Compagno, 1984). EGGCASE 9cm in length (excluding horns). 3cm in width. Long tendrils from each corner (Compagno, 1984). Similar eggcase to the Small Spotted Catshark, Scyliorhinus canicula. DIET The feeds on a wide variety of prey but crustaceans dominate, particularly hermit crabs, swimming crabs, cancrid crabs and large shrimp. It also feeds on molluscs such as squid and octopi and a number of bony fish including mackerel, epigonids, dragonets, gurnards, flatfish, herring, and small codfish. It is also known to feed on the closely related Small Spotted Catshark, Scyliorhinus canicula (Compagno, 1984). REPRODUCTION Relatively little is known of the reproductive biology of the except that it is an oviparous species with an incubation period of between 9 and 11 months. Egg laying appears to occur during spring and summer in shallow water. It is thought that the Fal Estuary in Cornwall is an extensive and important laying area and nursery ground for the species due to the large number of eggcases and animals found there (Orton, 1926). From eggcase surveys, the Llŷn Peninsula in North Wales also appears to be important (Hood and Reeve, pers comm.). The eggs are large, around 9cm long and 3cm wide. Garstang (1894) notes that breeding around Plymouth occurs during November, December and January, although it is unclear if this refers to mating, laying or hatching (Garstang, 1894). The pups emerge measuring around 16cm in length (Compagno, 1984).

Supported by: COMMERCIAL IMPORTANCE The is mostly taken as bycatch but is also targeted throughout its range by bottom trawls, gill nets, bottom set longlines, handlines, fixed bottom nets and occasionally pelagic trawls. It is utilised fresh or dried salted for human consumption and can be processed into fishmeal (Gibson et al., 2006). THREATS, CONSERVATION, LEGISLATION Around the British Isles the can be locally abundant, particularly around the coasts of Pembrokeshire, Anglesey, the Llŷn Peninsula and Cardigan Bay. However, it may be at risk of localised depletion. It is often fished with bottom trawls, gill nets, bottom set longlines, handlines, fixed bottom nets and even pelagic trawls. Limited data is available on exploitation and abundance trends, although declines have been recorded in the northwest Mediterranean (Gibson et al., 2006). While little is known of the biology of the, it is a late maturing species with low fecundity. Combined with a limited level of interconnectivity between island populations, the species has a low potential level of recovery. It lives entirely within the depth range heavily exploited by near shore fisheries and as such, is likely to have seen significant population declines (Gibson et al., 2006). IUCN RED LIST ASSESSMENT Near Threatened (2008). HANDLING AND THORN ARRANGEMENT Handle with care. Sharp teeth. Abrasive skin.

REFERENCES COMPAGNO, L. J. V. 1984. Sharks of the World: An Annotated and Illustrated Catalogue of Shark Species Known to Date. Volume 4, Part 1. Hexanchiformes to Lamniformes. FAO. Rome, Italy. ELLIS, J., SERENA, F., MANCUSI, C., HAKA, F., MOREY, G., GUALLART, J., SCHEMBRI, T. 2006.. In: IUCN 2010. IUCN Red List of Threatened Species. Version 2010.1. www.iucnredlist.org GARSTANG, W. 1894. Faunistic Notes at Plymouth 1893-4: With Observations on the Breeding Seasons of Marine Animals, and on the Periodic Changes of the Floating Fauna. Marine Biol. Assoc. (N. s.) Ill, pp. 210-235. GIBSON, C., VALENTI, S. V., FOWLER, S. L., FORDHAM, S. V. 2006. The Conservation Status of Northeast Atlantic Chondrichthyans; Report of the IUCN Shark Specialist Group Northeast Atlantic Regional Red List Workshop. VIII + 76pp. IUCN SSC Shark Specialist Group. ORTON, J. H. A Breeding Ground of the (Scyliorhinus stellaris) in the Fal Estuary. Nature, Vol. 118: 732. SOLDO, A., DULČIĆ, J., CETINIĆ, P. 2000. Contribution to the study of the morphology of the teeth of the Scyliorhinus stellaris (Chondrichthyes: Scyliorhinidae). Scientia Marina. 64 (3): 355-356. Text: Richard Hurst. Illustrations: Marc Dando. Citation Shark Trust; 2010. An Illustrated Compendium of Sharks, Skates, Rays and Chimaera. Chapter 1: The British Isles and Northeast Atlantic. Part 2: Sharks. Any ammendments or corrections, please contact: The Shark Trust 4 Creykes Court, The Millfields Plymouth, Devon PL1 3JB Tel: 01752 672008/672020 Email: enquiries@sharktrust.org For more ID materials visit www.sharktrust.org/id. Registered Company No. 3396164. Registered Charity No. 1064185