Hamamelidaceae (& Altingiaceae)

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Hamamelidaceae (& Altingiaceae) Hamamelis study weekend IDS Arboretum Kalmthout, Belgium 1 & 2 February 2014

Hamamelidaceae (& Altingiaceae) vegetative key to species in cultivation

Morphologically very diverse genera. With number of species in this key: Altingia* [3] Corylopsis [10] Disanthus [1] Distylium [4] Exbucklandia [1] Fortunearia [1] Fothergilla [2] Hamamelis [6] Liquidambar* [5] Loropetalum [1] Parrotia [2] Parrotiopsis [1] Rhodoleia [1] Sinowilsonia [1] Sycoparrotia [1] Sycopsis [1] Trichocladus [2] (Altingiaceae*)

key to families Altingiaceae Hamamelidaceae

Altingiaceae Altingia gracilipes

Altingiaceae bruised lamina with sharp scent Altingia gracilipes

Altingiaceae bruised lamina with sharp scent leaf arrangement spirally Altingia gracilipes

Hamamelidaceae Loropetalum chinense var. rubrum

Hamamelidaceae bruised lamina scentless Loropetalum chinense var. rubrum

Hamamelidaceae bruised lamina scentless leaf arrangement two-ranked Loropetalum chinense var. rubrum

Altingiaceae bruised lamina with sharp scent leaf arrangement spirally Altingia gracilipes

key to groups 2 groups each with following diagnostic characters

GROUP A: LAMINA VENATION PINNATE (secondary veins scattered along midvein) Altingia poilanei

GROUP A: LAMINA VENATION PINNATE (secondary veins scattered along midvein) Altingia poilanei

GROUP B: LAMINA VENATION PALMATE (strong veins predominantly radiating from base) Liquidambar orientalis

GROUP B: LAMINA VENATION PALMATE (strong veins predominantly radiating from base) Liquidambar orientalis

Hamamelidaceae bruised lamina scentless leaf arrangement two-ranked Loropetalum chinense var. rubrum

key to groups 3 groups each with following diagnostic characters

GROUP A LAMINA VENATION PALMATE strong veins predominantly radiating from base

GROUP A LAMINA VENATION PALMATE strong veins predominantly radiating from base GROUP B LAMINA VENATION PINNATE most secondary veins curving + anastomosing before margin

GROUP A LAMINA VENATION PALMATE strong veins predominantly radiating from base GROUP B LAMINA VENATION PINNATE most secondary veins curving + anastomosing before margin GROUP C LAMINA VENATION PINNATE most secondary veins ending at margin

GROUP A LAMINA VENATION PALMATE strong veins predominantly radiating from base

GROUP A: LAMINA VENATION PALMATE Disanthus cercidifolius

GROUP A: LAMINA VENATION PALMATE Disanthus cercidifolius

GROUP A: LAMINA VENATION PALMATE Lamina unlobed, papery, apex obtuse to acute. Disanthus cercidifolius

GROUP A: LAMINA VENATION PALMATE Lamina unlobed, papery, apex obtuse to acute. Stipules free. Disanthus cercidifolius

GROUP A: LAMINA VENATION PALMATE Exbucklandia populnea

GROUP A: LAMINA VENATION PALMATE Lamina unlobed(rarely lobed), leathery, apex acuminate to caudate. Exbucklandia populnea

GROUP A: LAMINA VENATION PALMATE Lamina unlobed (rarely lobed), leathery, apex acuminate to caudate. Stipules united. Exbucklandia populnea

GROUP A: LAMINA VENATION PALMATE Lamina unlobed (rarely lobed), leathery, apex acuminate to caudate. Stipules united. Exbucklandia populnea

GROUP A LAMINA VENATION PALMATE strong veins predominantly radiating from base GROUP B LAMINA VENATION PINNATE most secondary veins curving + anastomosing before margin

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Distylium racemosum

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Distylium racemosum

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Trichocladus ellipticus

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Shoot, petiole and lamina pubescent with long-armed stellate hairs. Trichocladus ellipticus

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Shoot, petiole and lamina pubescent with long-armed stellate hairs. Trichocladus ellipticus

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Loropetalum sinense

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Shoot, petiole and lamina pubescent with long-armed stellate hairs. Loropetalum sinense

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Shoot, petiole and lamina pubescent with long-armed stellate hairs. Loropetalum sinense

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Shoot, petiole and lamina glabrous or pubescent with scale-like to short-armed stellate hairs. Rhodoleia championii

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Lamina LS +/- glaucous green, margin darker than surface. Rhodoleia championii

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN leaf arrangement spirally Rhodoleia championii

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN leaf arrangement spirally Rhodoleia championii

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Rhodoleia championii sensu lato lamina rather variable

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Sycopsis sinensis

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Lamina LS green, margin translucent (10 LENS). Sycopsis sinensis

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Lamina LS green, margin translucent (10 LENS). Sycopsis sinensis

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Lamina LS green, margin translucent (10 LENS). Lamina apex predominantly gradually acuminate to (sub-)caudate. Sycopsis sinensis

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Lamina LS green, margin translucent (10 LENS). Distylium myricoides

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Lamina LS green, margin translucent (10 LENS). Distylium myricoides

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Lamina LS green, margin translucent (10 LENS). Apex obtuse to (abruptly) acute or abruptly acuminate. Distylium myricoides

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Distylium racemosum

GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Distylium racemosum

GROUP A LAMINA VENATION PALMATE strong veins predominantly radiating from base GROUP B LAMINA VENATION PINNATE most secondary veins curving + anastomosing before margin GROUP C LAMINA VENATION PINNATE most secondary veins ending at margin

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Hamamelis mollis

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Hamamelis mollis

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Fortunearia sinensis

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lamina margin with >25 teeth or vein endings/side. Fortunearia sinensis

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lamina margin with >25 teeth or vein endings/side. Lamina base rounded to cuneate. Fortunearia sinensis

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Sinowilsonia henryi

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lamina margin with >25 teeth or vein endings/side. Sinowilsonia henryi

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lamina margin with >25 teeth or vein endings/side. Lamina base cordate to obliquely cordate. Sinowilsonia henryi

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Parrotia persica

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lamina margin with 25 teeth or vein endings/side. Parrotia persica

GROUP C: LAMINA VENATION PINNATE Lamina margin with 25 teeth or vein endings/side. MOST SECONDARY VEINS ENDING AT MARGIN Corylopsis sinensis

Teeth bristly at apex. GROUP C: LAMINA VENATION PINNATE Lamina margin with 25 teeth or vein endings/side. MOST SECONDARY VEINS ENDING AT MARGIN Corylopsis sinensis

Teeth bristly at apex. GROUP C: LAMINA VENATION PINNATE Lamina margin with 25 teeth or vein endings/side. MOST SECONDARY VEINS ENDING AT MARGIN Corylopsis sinensis

Teeth bristly at apex. GROUP C: LAMINA VENATION PINNATE Lamina margin with 25 teeth or vein endings/side. Lamina base cordate. MOST SECONDARY VEINS ENDING AT MARGIN Corylopsis sinensis

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Hamamelis mollis

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lamina LS with lowermost secondary vein completely surrounded by leaf tissue. Hamamelis mollis

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Fothergilla major

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lowermost secondary vein OFTEN marginal for at least basal 2-10 mm. Fothergilla major

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lowermost secondary vein OFTEN marginal for at least basal 2-10 mm. Fothergilla major

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Hamamelis japonica

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lowermost secondary vein OFTEN marginal for at least basal 2-10 mm. Hamamelis japonica

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lowermost secondary vein OFTEN marginal for at least basal 2-10 mm. Hamamelis japonica

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Parrotia persica

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lamina LS secondary vein axils with tiny membranous domatia. Parrotia persica

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Parrotiopsis jacquemontiana

GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lamina often suborbicular. Parrotiopsis jacquemontiana

JDL vegetative identification keys jan.de.langhe@telenet.be www.arboretumwespelaar.be

JDL vegetative identification keys jan.de.langhe@telenet.be www.arboretumwespelaar.be

thanks to IDS Wolfgang Bopp Tom Hudson Peter Brownless Jaime Morin Koen Camelbeke Jacky Pousse Tom Clark Abraham Rammeloo Francisco Garin Paul Reader Paul Goetghebeur Rolf Zumbrunn John Grimshaw Arboretum Kalmthout Arboretum Wespelaar Ghent University Botanical Garden Hillier Gardens and Arboretum Iturraran Botanical Garden Polly Hill Arboretum Royal Botanic Gardens Edinburgh Royal Botanic Gardens Kew