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Checklist of the Indigenous Cerambycidae of the Hawaiian Islands D. J. Heffern - Electronic Version, 2002 A checklist and bibliography for the native Cerambycidae of the Hawaiian Islands is presented. The following format is used: Genus Author, year-page number (type species: Genus species Author) Generic synonym species Author, year-page number [ original genus, if different from current] specific synonym distribution Please send corrections, suggestions and comments to the author at the following email: titanusgiganteus@hotmail.com Subfamily Parandrinae Tribe Parandrini Parandra Latreille, 1804-134 (type species: Parandra laevis Latreille) puncticeps Sharp, 1878-202 Hawaii I., Kauai I., Maui I., Molokai I., Oahu I. Subfamily Prioninae Tribe Megopodini Megopis Audinet-Serville, 1832-161 (type species: Megopis mutica Audinet-Serville) reflexa (Karsch, 1881-7) [Aegosoma] Hawaii I., Kauai I., Lanai I., MauiI., Molokai I., Oahu I. Subfamily Cerambycinae Tribe Clytini Plagithmysus Motschulsky, 1845-41 (type species: Stenopterus pulverulentus Motschulsky) Callithmysus Sharp, 1896-238 (type species: Clytarlus microgaster Sharp) Neoclytarlus Bridwell, 1920-322 (type species: Clytarlus fragilis Sharp) Nesithmysus Perkins, 1920-343 (type species: Nesithmysus bridwelli Perkins) 1

Paraclytarlus Perkins, 1927-479 (type species: Plagithmysus (Paraclytarlus) timberlakei Perkins) Aeschrithmysus Perkins, 1929-261 (type species: Aeschrithmysus terryi Perkins) Peleithmysus Gressitt, 1972b-636 (type species: Nesithmysus forbesii Perkins) Plagithmysus (Plagithmysus) Motschulsky, 1845-41 aequalis Sharp, 1896-273 Kauai I. aestivus Sharp, 1896-272 Molokai I. albertisi Sharp, 1897-12 Oahu I. annectans (Sharp in Sharp & Perkins, 1900-104) [Clytarlus] Kauai I. attenuatus (Boisduval, 1835-485) [Clytus] Oahu I. bilineatus Sharp, 1896-243 Hawaii I. bishopi bishopi Sharp, 1896-242 Hawaii I. bishopi gracilis Sharp, 1896-243 Hawaii I. blackburni (Sharp in Blackburn & Sharp, 1885-195) [Clytarlus] Hawaii I. cheirodendri Gressitt & Davis, 1969-334 Maui I. collaris Sharp in Sharp & Perkins, 1900-107 Maui I. concolor concolor Sharp, 1896-241 Kauai I. concolor arachnipes Sharp, 1896-274 Kauai I. concolor munroi Sharp in Sharp & Perkins, 1900-112 Kauai I. cristatus (Sharp, 1878-207) [Clytarlus] Oahu I. cuneatus Sharp, 1896-241 Oahu I. sapindi Perkins, 1933-265 darwinianus Sharp, 1896-271 Hawaii I. davisi Swezey, 1946-621 Hawaii I. decorus Perkins, 1921-500 Hawaii I. diana Sharp in Sharp & Perkins, 1900-107 Kauai I. elegans Sharp in Perkins, Scott & Sharp, 1910-648 Hawaii I. eugeniae Gressitt & Davis, 1974-359 Maui I. finschi (Harold, 1880-166) [Clytarlus] Maui I. forbesianus Gressitt, 1972b-639 Kauai I. forbesii Perkins, 1927-471 not Nesithmysus forbesii Perkins, 1921-503 fractus Perkins in Perkins, Scott & Sharp, 1910-646 Molokai I. frater Perkins, 1921-500 Hawaii I. funebris Sharp, 1896-273 Maui I. giffardi Perkins, 1907-96 Hawaii I. greenwelli Gressitt & Davis, 1970-567 Hawaii I. hirtipes (Sharp in Sharp & Perkins, 1900-113) [Callithmysus] Oahu I. ignotus Perkins, 1916-249 Kauai I. ilicis Gressitt, 1972a-86 Molokai I. kawauae Gressitt & Davis, 1976b-241 Hawaii I. koae Gressitt & Davis, 1969-340 Maui I. koaiae Gressitt & Davis, 1969-335 Hawaii I. 2

koebelei (Perkins, 1908-210) [Callithmysus] Oahu I. kohalae Perkins, 1927-473 Hawaii I. kraussi Gressitt & Davis, 1969-339 Hawaii I. kuhnsi Perkins, 1916-248 Oahu I. lamarckianus Sharp in Sharp & Perkins, 1900-110 Hawaii I. lanaiensis Sharp, 1896-244 Lanai I. laui Gressitt, 1972b-645 Maui I. longicollis Perkins, 1927-474 Maui I. mauiensis Gressitt, 1972b-644 Maui I. microgaster microgaster (Sharp, 1879-103) [Clytarlus] Oahu I. microgaster hirtipes (Sharp in Sharp & Perkins, 1900-113) [Callithmysus] Oahu I. molokaiensis Sharp, 1927-475 Molokai I. muiri Perkins, 1927-476 Oahu I. newelli Sharp, 1896-240 Maui I. nicotianae Gressitt & Davis, 1969-338 Maui I. nihoae Perkins, 1926-53 Nihoa I. paludis Perkins, 1927-472 Kauai I. perkinsi Sharp, 1896-244 Hawaii I. permundus Sharp in Sharp & Perkins, 1900-105 Kauai I. perrottetiae Gressitt & Davis, 1971-71 Kauai I. pipturi Gressitt, 1972b-639 Maui I. pittospori Gressitt, 1972a-85 Lanai I. platydesmae Perkins, 1920-345 Hawaii I. polystictus Perkins, 1933-266 Kauai I. pulverulentus (Motschulsky, 1845-41) [Stenopterus] Oahu I. robustus Sharp, 1878-206 [Clytarlus] pulvillatus (Karsch, 1881-9) [Clytarlus] Maui I. rebeccae Gressitt, 1972b-643 Maui I. rubi Perkins, 1931-415 Maui I. sharpianus Perkins, 1927-475 Kauai I. simillimus Perkins, 1931-415 Maui I. simplicollis Sharp in Perkins, Scott & Sharp, 1910-648 Hawaii I. solitarius Sharp, 1896-241 Oahu I. speculifer Sharp, 1896-272 Maui I. sugawai Gressitt & Davis, 1971-72 Kauai I. sulphurescens Sharp, 1896-271 Hawaii I. swezeyi Perkins, 1920-344 Hawaii I. ukulele Gressitt, 1972a-87 Maui I. urerae Gressitt & Davis, 1976a-238 Maui I. varians Sharp, 1896-245 Hawaii I. vicinus Sharp, 1896-243 Hawaii I. vitticollis vitticollis Sharp, 1896-240 Hawaii I. vitticollis longulus Sharp, 1896-240 Hawaii I. 3

Plagithmysus (Aeschrithmysus) Perkins, 1929-261 dubautianus (Gressitt & Davis, 1969-332) [Aeschrithmysus] Maui I. swezeyanus Gressitt & Davis, 1971-71 Maui I. swezeyi (Perkins, 1929-262) [Aeschrithmysus] not Plagithmysus swezeyi Perkins, 1920-344 terryi (Perkins, 1929-261) [Aeschrithmysus] Maui I. yoshimotoi Gressitt & Davis, 1969-343 Maui I Plagithmysus (Neoclytarlus) Bridwell, 1920-322 abnormis (Sharp in Sharp & Perkins, 1900-102) [Clytarlus] Hawaii I. acaciae Gressitt & Davis, 1969-351 Hawaii I. akoko Gressitt & Davis, 1976a-239 Kauai I. atricolor (Perkins, 1933-266) [Neoclytarlus] Hawaii I. bidensae Gressitt, 1972a-89 Molokai chenopodii (Perkins, 1938-59) [Neoclytarlus] Oahu I. claviger (Sharp in Sharp & Perkins, 1900-101) [Clytarlus] Hawaii I. debilis (Sharp in Sharp & Perkins, 1900-99) [Clytarlus] Hawaii I. decurrensae Gressitt & Davis, 1969-347 Hawaii I. dodonaeae (Swezey, 1946-621) [Neoclytarlus] Hawaii I. dodonaeavorus Gressitt, 1972a-90 Molokai dubautiae dubautiae Gressitt & Davis, 1969-352 Hawaii I. dubautiae arboreae Gressitt & Davis, 1969-354 Hawaii I. euphorbiae (Bridwell, 1920-323) [Neoclytarlus] Oahu I. filipes filipes (Sharp in Blackburn & Sharp, 1885-196) [Clytarlus] Hawaii I. filipes sophorae Gressitt & Davis, 1969-350 Hawaii I. fragilis (Sharp, 1881-534) [Clytarlus] Oahu I. fugitivus Perkins, 1927-483 Maui I. geranii (Perkins, 1931-417) [Neoclytarlus] Maui I. hardyi Gressitt, 1972a-87 Maui I. hoikuahiwi Gressitt & Davis, 1974-362 Kauai I. immundus Sharp in Perkins, Scott & Sharp, 1910-646 Hawaii I. indecens (Perkins, 1920-346) [Clytarlus] Oahu I. jacobii Gressitt & Davis, 1974-361 Hawaii I. kainaluensis (Perkins, 1931-418) [Neoclytarlus] Molokai I. longipes (Sharp in Sharp & Perkins, 1900-103) [Clytarlus] Kauai I. lookii lookii (Swezey, 1947-101) [Neoclytarlus] Hawaii I. lookii keanakolui Gressitt & Davis, 1969-344 Hawaii I. lookii ukae Gressitt & Davis, 1969-344 Hawaii I. medeirosi Samuelson & Davis, 1986-125 Maui I. mediocris (Sharp in Sharp & Perkins, 1900-99) [Clytarlus] Maui I. metrosideri Gressitt & Davis, 1969-346 Kauai I. mezoneuri (Swezey, 1946-623) [Neoclytarlus] Hawaii I. 4

modestus (Sharp, 1879-104) [Clytarlus] Maui I. montgomeryi Gressitt & Davis, 1971-73 Hawaii I. nodifer (Sharp in Sharp & Perkins, 1900-102) [Clytarlus] Hawaii I. obscurus (Sharp in Sharp & Perkins, 1900-100) [Clytarlus] Kauai I. pennatus (Sharp, 1881-532) [Clytarlus] Maui I. pulchrior (Perkins, 1927-482) [Neoclytarlus] Hawaii I. railliardiae (Perkins, 1931-416) [Neoclytarlus] Maui I. rusticus Gressitt & Davis, 1969-348 Hawaii I. smilacis (Perkins, 1927-484) [Neoclytarlus] Maui I. smilacivorus Gressitt, 1972a-88 Lanai I. superstes (Zimmerman, 1940-479) [Neoclytarlus] Oahu I. ultimus (Sharp in Perkins, Scott & Sharp, 1910-645) [Clytarlus] Oahu I. usingeri Gressitt & Davis, 1969-345 Oahu I. wattleae Gressitt & Davis, 1969-355 Maui I. Plagithmysus (Nesithmysus) Perkins, 1920-343 bridwelli (Perkins, 1920-343) [Nesithmysus] Oahu I. peleanus Gressitt & Davis, 1971-70 Molokai I. swezeyellus Gressitt, 1972a-84 Maui I. swezeyi (Perkins, 1927-485) [Nesithmysus] not Plagithmysus swezeyi Perkins, 1920-344 sylvai Gressitt, 1972b-637 Maui I. Plagithmysus (Paraclytarlus) Perkins, 1927-479 annectans (Sharp in Sharp & Perkins, 1900-104) [Clytarlus] Kauai I. laticollis (Sharp in Sharp & Perkins, 1900-101) [Clytarlus] Maui I. fugitivus (Perkins, 1927-483) [Neoclytarlus] pipturicola Perkins, 1927-481 Maui I. podagricus Perkins, 1927-482 Hawaii I. timberlakei Perkins, 1927-480 Oahu I. Plagithmysus (Peleithmysus) Gressitt, 1972b-636 alani Gressitt, 1972b-636 Maui I. forbesi (Perkins, 1921-503) [Nesithmysus] Maui I. haasi (Perkins, 1921-504) [Nesithmysus] Oahu I. peleae (Gressitt & Davis,1971-67) [Nesithmysus] Molokai I 5

Bibliography for Original Descriptions of Indigenous Hawaiian Cerambycidae Blackburn, T. and D. Sharp. 1885. Memoirs on the Coleoptera of the Hawaiian Islands. Transactions of the Royal Dublin Society. 2(3):119-290, 2 pls. Boisduval, J. B. A. 1835. Voyage de découvertes de l Astrolabe. Exécuté par ordre du Roi, pendant les années 1826-1827-1828-1829, sous le commandement de M. J. Dumont D Urville. Faune entomologique de l Océan Pacifique, avec l illustration des insectes nouveaux recueillis pendant le voyage, deuxième partie, coléoptères et autres ordres. 716 pp. Paris. Bridwell, J. C. 1920. A new lowland plagithmysine cerambycid from Oahu with notes on its habits (Coleoptera). Proceedings of the Hawaiian Entomological Society. 4(2):314-323. Gressitt, J. L. 1972a. New plagithmysines from Molokai, Lanai, and Maui (Col.: Cerambyc.). Pacific Insects. 14(1):83-92, 7 fig. Gressitt, J. L. 1972b. New plagithmysines from West Maui (Col.: Cerambyc.). Pacific Insects. 14(4):635-645, 7 fig. Gressitt, J. L. and C. J. Davis. 1969. Studies in the plagithmysines, endemic Hawaiian Cerambycidae (Coleopt.). Proceedings of the Hawaiian Entomological Society. 20(2):331-393. fig. 18. Gressitt, J. L. and C. J. Davis. 1970. A new plagithmysine beetle from sandalwood (Col. Cerambycidae). Proceedings of the Hawaiian Entomological Society. 20(3):567-569, 1 fig. Gressitt, J. L. and C. J. Davis. 1971. New plagithmysines from Kauai, Molokai and Hawaii (Coleopt.: Cerambycidae). Proceedings of the Hawaiian Entomological Society. 21(1):67-77, 5 fig. Gressitt, J. L. and C. J. Davis. 1974. New plagithmysines from Kauai, Maui and Hawaii (Col.: Cerambycidae). Proceedings of the Hawaiian Entomological Society. 21(3):359-363. Gressitt, J.L. & C.J. Davis. 1976a. New plagithmysines from Kauai and West Maui (Coleoptera: Cerambycidae). Proceedings of the Hawaiian Entomological Society. Proceedings of the Hawaiian Entomological Society. 22(2):237-239. Gressitt, J.L. & C.J. Davis. 1976b. A new plagithmysine from Ilex on Hawaii (Coleoptera: Cerambycidae). Proceedings of the Hawaiian Entomological Society. 22(2):241-243. Harold, E. von. 1880. Einige neue Coleopteren. Mitteilungen der Münchener Entomologischen Vereins. 4:148-171. Karsch, F. 1881. Zur Käferfauna der Sandwich-, Marshall- und Gilberts-Inseln. Berliner Entomologisches Zeitschrift. 25:1-14, pl. 1. Maehler, K. L. 1947. Notes and exhibitions. Plagithmysus newelli Sharp. Proceedings of the Hawaiian Entomological Society. 13(2):203. Motschulsky, V. 1845. Remarques sur la collection de coléoptères Russes da Victor Motschulsky. Bulletin de la Société Impériale des Naturalistes de Moscou. 18(1):1-85. Perkins, R.C.L. 1907. Insects at Kilauea, Hawaii. Proceedings of the Hawaiian Entomological Society. 1(3):89-99. Perkins, R. C. L. 1908. Callithmysus koebelei sp. nov. Proceedings of the Hawaiian Entomological Society. (5):210. 6

Perkins, R. C. L. 1916. Some new Hawaiian Coleoptera. Proceedings of the Hawaiian Entomological Society. 3(3): 247-251. Perkins, R. C. L. 1920. Some new Hawaiian Coleoptera. Proceedings of the Hawaiian Entomological Society. 4(2):314-359. Perkins, R. C. L. 1921. Notes on Hawaiian Plagithmysides and Anobiides (Col.) with descriptions of new species. Proceedings of the Hawaiian Entomological Society. 4(3):493-506. Perkins, R.C.L. 1926. Coleoptera, weevils. Pp. 53-66. in E.H. Bryan, Jr. & Collaborators. Insects of Hawaii, Johnston Island and Wake Island. Bulletin of the Bernice P. Bishop Museum. 31:94 pp. Perkins, R. C. L. 1927. Notes on Hawaiian Coleoptera (Curculionidae, Proterhinidae and Cerambycidae) and descriptions of new species. Proceedings of the Hawaiian Entomological Society. 6(3):465-488. Perkins, R. C. L. 1929. On a new genus, with two new species, of Hawaiian Cerambycidae (Col.). Proceedings of the Hawaiian Entomological Society. 7(2):261-262. Perkins, R. C. L. 1931. New species of Hawaiian Cerambycidae (Col.). Proceedings of the Hawaiian Entomological Society. 7(3):415-418. Perkins, R. C. L. 1933. New Hawaiian Coleoptera, with notes on some previously known species. Proceedings of the Hawaiian Entomological Society. 8(2):265-272. Perkins, R. C. L. 1938. Another new cerambycid in Hawaii. Proceedings of the Hawaiian Entomological Society. 10(1):59. Perkins, R. C. L., H. Scott and D. Sharp. 1910. Coleoptera. V. Coleoptera (various). Fauna Hawaiiensis 3(6): 581-644. Samuelson, G. A. and C. J. Davis. 1986. A new Euphorbia-associated plagithmysine from Maui (Coleoptera: Cerambycidae). Proceedings of the Hawaiian Entomological Society. 27:125-128. Sharp, D. 1878. On some longicorn Coleoptera from the Hawaiian Islands. Transactions of the Entomological Society of London. 1878:201-210. Sharp, D. 1879. On some Coleoptera from the Hawaiian Islands. Transactions of the Entomological Society of London. 1879:77-105. Sharp, D. 1881. On some new Coleoptera from the Hawaiian Islands. Transactions of the Entomological Society of London. 1881:507-534. Sharp, D. 1896. On Plagithmysus: A Hawaiian genus of longicorn Coleoptera. Entomologist's Monthly Magazine. 32:237-245; 271-274. Sharp, D. 1897. On Plagithmysus: A Hawaiian genus of longicorn Coleoptera. - Supplement. Entomologist's Monthly Magazine. 33: 12. Sharp, D. and R. C. L. Perkins. 1900. Coleoptera Phytophaga. Fauna Hawaiiensis. 2(1):91-116, pl. 6. Swezey, O. H. 1946. Some new Cerambycidae from the Island of Hawaii (Coleoptera). Proceedings of the Hawaiian Entomological Society. 12(3):621-623. Swezey, O. H. 1947. A new Neoclytarlus from Chenopodium oahuense (Coleoptera: Cerambycidae). Proceedings of the Hawaiian Entomological Society. 13(1):101-102. Zimmerman, E. C. 1940. A new lowland Neoclytarlus from Oahu (Coleoptera: Cerambycidae). Proceedings of the Hawaiian Entomological Society. 10(3):479-481. 7

Useful Additional Bibliography Davis, C. J. 1953. New host and insect records from the Island of Hawaii. Proceedings of the Hawaiian Entomological Society. 15(1):135-158. Duffy, E. A. 1953. The immature stages of Hawaiian Cerambycidae with a key to larvae. Proceedings of the Hawaiian Entomological Society. 15(1):135-158. Fullaway, D.T. & W.M. Giffard. 1919. Notes on collection of Hawaiian insects on island of Maui. Proceedings of the Hawaiian Entomological Society. 4:50-52. Gressitt, J. L. and C. J. Davis. 1972. Seasonal occurrence of the Hawaiian Cerambycidae (Col.). Proceedings of the Hawaiian Entomological Society. 21(2):213-221. Southwood, T. R. E. 1960. The abundance of the Hawaiian trees and the numbers of their associated insect species. Proceedings of the Hawaiian Entomological Society. 17(2):293-303. Swezey, O. H. 1932. The host trees of the endemic Cerambycidae in Hawaii. Proceedings of the Hawaiian Entomological Society. 8(1):127-140. Swezey, O. H. 1954. Forest entomology in Hawaii. Bernice P. Bishop Museum Special Publications. 44:1-266. ill. 8