Curriculum Vitae MARK A. STEELE

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Mark A. Steele page 1 Curriculum Vitae MARK A. STEELE Department of Biology California State University 18111 Nordhoff Street Northridge, CA 91330-8303 Phone: (818) 677-4270 FAX: (818) 677-2034 e-mail: msteele@csun.edu EDUCATION B.A. Ph.D. 1989. University of California, San Diego. Major: Ecology, Behavior, and Evolution; Cum Laude 1995. University of California,. Biology ACADEMIC POSITIONS 2007 present Assistant Professor, Department of Biology, California State University, Northridge 2006 2007 Visiting Assistant Professor, Department of Biology, California State University, Northridge 2006 present Associate Research Biologist, Marine Science Institute, University of California, 2002 2006 Assistant Research Biologist, Marine Science Institute, University of California, 2003 present Adjunct Assistant Professor, Department of Natural Resources Science, University of Rhode Island 1998 present Faculty, East/West Marine Biology Program, Northeastern University 2002 2003 Adjunct Assistant Professor, Department of Biological Sciences, University of Rhode Island 1999 2001 Assistant Professor (Research), Department of Biological Sciences, University of Rhode Island 2000 Lecturer, Department of Organismic Biology, Ecology and Evolution, University of California, Los Angeles 1999 2000 Postdoctoral Associate, Department of Organismic Biology, Ecology and Evolution, University of California, Los Angeles, with Rick Vance 1995 1999 Postdoctoral Associate, Department of Organismic Biology, Ecology and Evolution, University of California, Los Angeles, with Graham Forrester 1991, 1995 Research Assistant, Department of Biological Sciences, University of California, 1990-1995 Teaching Assistant, Department of Biological Sciences, University of California, 1987-1989 Teaching Assistant, Biology Department, University of California, San Diego

Mark A. Steele page 2 TEACHING EXPERIENCE 2008 Graduate Seminar in Ecology (Department of Biology, California State University, Northridge) 2007-present Ichthyology (Department of Biology, California State University, Northridge) 2006-present Marine Ecology (Department of Biology, California State University, Northridge) 2006-present Ecology of Marine Fishes (Department of Biology, California State University, Northridge) 1998-present Biology of Fishes (East/West Marine Biology Program, Northeastern University) 2000 Ecology (Department of Organismic Biology, Ecology and Evolution, University of California, Los Angeles) 1990-1995 Ecology; Ecology Lab; General Biology (Teaching Assistant, Department of Biological Sciences, University of California, ) 1987-1989 Introductory Biology; Human Nutrition (Teaching Assistant, Biology Department, University of California, San Diego) AWARDS, FELLOWSHIPS, SCHOLARSHIPS, AND HONORS 1997 Mildred Mathias Award for best natural science proposal submitted to the UC MEXUS program 1995 UCSB General Affiliates Graduate Dissertation Fellowship 1994 Regent's Fellowship, University of California, 1994 International Women's Fishing Association Scholarship 1994 Graduate Student Fee Fellowship, University of California, 1993 International Women's Fishing Association Scholarship 1993 Graduate Student Fee Fellowship, University of California, 1992 International Women's Fishing Association Scholarship 1992 Graduate Student Fee Fellowship, University of California, 1991 Honorable Mention, National Science Foundation Graduate Fellowship 1990 Regent's Fellowship, University of California, 1988 President's Undergraduate Fellowship, University of California, San Diego GRANTS AND CONTRACTS 2008-2009 California Coastal Commission. San Onofre Nuclear Generating Station Mitigation Project Monitoring Program: Measuring fecundity, growth, and food-chain support of kelp forest fishes on the SONGS-mitigation artificial reef and reference reefs. $175,670. (PI with L. Allen) 2007 California Coastal Commission. San Onofre Nuclear Generating Station Mitigation Project Monitoring Program: Measuring fecundity of five species of kelp forest fishes. $38,920. (PI with L. Allen) 2007-2008 California Department of Fish and Game. Pilot study to monitor the occurrence of invasive species occupying rip-rap habitat in Los Angeles Harbor, California. $47,813. (co-pi with L. Allen) 2006-2007 California Department of Fish and Game. Nearshore gill net sampling program for white seabass (age I-IV). $95,663. (co-pi with 2 others)

Mark A. Steele page 3 2002-2007 National Science Foundation (OCE 0222087). A test for shelter limitation of reef fish populations at large spatial scales: an integrated empirical and theoretical approach. $549,167. (co-pi; G. Forrester PI) 2004-2006 Caribbean Marine Research Center, National Undersea Research Program, National Oceanic and Atmospheric Administration (CMRC-03-NRME-01-04A). Density dependence in exploited marine fish: an experimental test and theoretical exploration of the implications for fisheries enhancement via marine protected areas. $57,983. (PI) 2004-2005 California Coastal Commission. San Onofre Nuclear Generating Station Mitigation Project Monitoring Program. $2,388,957. (co-pi with 3 others) 2002-2004 Caribbean Marine Research Center, National Undersea Research Program, National Oceanic and Atmospheric Administration (CMRC-01-NRME-01-02A). Source-sink dynamics, density dependence, and the efficacy of marine protected areas. $59,028. (PI) 2001-2002 Caribbean Marine Research Center, National Undersea Research Program, National Oceanic and Atmospheric Administration (CMRC-01-PRGF-01-01A). Population dynamics of coral reef fishes at multiple scales: an integrated empirical and theoretical approach. $7,475. (PI) 1997-2001 National Science Foundation (OCE 96-18011). Local population dynamics of temperate and tropical reef fishes at multiple scales. $403,000. (co-author; G. Forrester PI) 1997-1998 UC MEXUS Grant. Assessment of artisanal fisheries of the east coast of Baja California Sur, Mexico. $12,500. (co-author; G. Forrester PI) 1995 University of California Travel Grant 1993 Sigma Xi Grant-in Aid of Research 1993 Lerner-Gray Fund for Marine Research 1992 Sigma Xi Grant-in Aid of Research 1992 Lerner-Gray Fund for Marine Research PROFESSIONAL MEMBERSHIPS American Academy of Underwater Sciences American Society of Ichthyologists and Herpetologists California Estuarine Research Society Ecological Society of America Estuarine Research Federation Sigma Xi Southern California Academy of Sciences Western Society of Naturalists PUBLICATIONS Samhouri, J. F., R. R. Vance, G. E. Forrester, and M. A. Steele. 2009. Musical chairs: densitydependent mortality caused by competition for unguarded refuges. Oecologia 160:257 265. Paddack, M. J., J. D. Reynolds, C. Aguilar, R. S. Appeldoorn, J. Beets, E. W. Burkett, P. M. Chittaro, K. Clarke, R. Esteves, A. C. Fonseca, G. E. Forrester, A. M. Friedlander, J. García- Sais, G. González-Sansón, L. K. B. Jordan, D. McClellan, M. W. Miller, P. P. Molloy, P. J. Mumby, I. Nagelkerken, M. Nemeth, R. Navas-Camacho, J. Pitt, N. V. C. Polunin, M. C. Reyes-Nivia, D. R. Robertson, A. Rodríguez Ramírez, E. Salas, S. R. Smith, R. E. Spieler,

Mark A. Steele page 4 M. A. Steele, I. D. Williams, C. Wormald, A. R. Watkinson, and I. M. Côté. 2009. Recent region-wide declines in Caribbean reef fish abundance. Current Biology 19:590-595. Samhouri, J. F., M. A. Steele, and G. E. Forrester. 2009. Intercohort competition drives selective mortality and shelter-mediated density dependence in a coral reef fish. Ecology 90:1009 1020. Stier, A. C., M. A. Steele, and A. J. Brooks. 2009. Coral reef fishes use the crown-of-thorns seastar as habitat. Coral Reefs 28:227. Forrester, G. E., M. A. Steele, J. F. Samhouri, B. Evans, and R. R. Vance. 2008. Spatial density dependence scales up but does not produce temporal density dependence in a coral reef fish. Ecology 89:2980-2985. Forrester, G. E., M. A. Steele, J. F. Samhouri, and R. R. Vance. 2008. Settling reef fish discriminate among habitats at large, but not small, spatial scales. Limnology and Oceanography 53:1956-1962. Wormald, C. L. and M. A. Steele. 2008. Testing assumptions of mark-recapture theory in the coral reef fish Lutjanus apodus. Journal of Fish Biology 73:498 509. Williams, J. P., L. G. Allen, M. A. Steele, and D. J. Pondella. 2007. El Niño periods increase growth of juvenile white seabass (Atractoscion nobilis) in the Southern California Bight. Marine Biology 152:193 200. Steele, M. A., S. C. Schroeter, and H. M. Page. 2007. Preliminary investigation of the effects of purse seine size on estimates of density and species richness of estuarine fishes. Estuaries and Coasts 30:344 347. Steele, M. A., S. C. Schroeter, and H. M. Page. 2006. Experimental evaluation of biases associated with sampling estuarine fishes with seines. Estuaries and Coasts 29:1172-1184. Steele, M. A., S. C. Schroeter, R. C. Carpenter, and D. J. Kushner. 2006. Top-down vs. bottom-up effects in kelp forests. Science 313:1738. Steele, M. A., S. C. Schroeter, and H. M. Page. 2006. Sampling characteristics and biases of enclosure traps for sampling fishes in estuaries. Estuaries and Coasts 29:630-638. Forrester, G. E., B. Evans, M. A. Steele, and R. R. Vance. 2006. Assessing the magnitude of intra- and interspecific competition in two coral-reef fishes using a response surface experiment. Oecologia 148:632 640. Steele, M. A. and T. W. Anderson. 2006. Predation. In: L. G. Allen, M. H. Horn, D. J. Pondella II (eds) Ecology of California Marine Fishes, U.C. Press, p. 428-448. Steele, M. A. and G. E. Forrester. 2005. Small-scale field experiments accurately scale up to predict density dependence in reef fish populations at large scales. Proceedings of the National Academy of Sciences 102:13513-13516. Forrester, G. E. and M. A. Steele. 2004. Predators, prey refuges, and the spatial scaling of density- dependent prey mortality. Ecology 85:1332-1342. Swearer, S. E., G. E. Forrester, M. A. Steele, A. J. Brooks, and D. W. Lea. 2003. Spatiotemporal and interspecific variation in otolith trace-elemental fingerprints in a temperate estuarine fish assemblage. Estuarine, Coastal and Shelf Science 56:1111-1123. Forrester, G. E., B. I. Fredericks, D. Gerdeman, B. Evans, M. A. Steele, K. Zayed, L. E. Schweitzer, I. H. Suffet, R. R. Vance, R.F. Ambrose. 2003. Growth of estuarine fish is associated with the combined concentration of sediment contaminants and shows no adaptation of acclimation to past conditions. Marine Environmental Research 56:423-442.

Mark A. Steele page 5 Steele, M. A. and G. E. Forrester. 2002. Variation in the relative importance of sublethal effects of predators and competitors on growth of a temperate reef fish. Marine Ecology Progress Series 237:233-245. Steele, M. A., J. C. Malone, A. M. Findlay, M. H. Carr, and G. E. Forrester. 2002. A simple method for estimating larval supply in reef fishes and a preliminary test of limitation of population size by larval supply in the kelp bass, Paralabrax clathratus. Marine Ecology Progress Series 235:195-203. Steele, M. A. and G. E. Forrester. 2002. Early post-settlement predation on three reef fishes: effects on spatial patterns of recruitment. Ecology 83: 1076-1091. Forrester, G. E., R. R. Vance, and M. A. Steele. 2002. Simulating large-scale population dynamics using small-scale data. In: P. F. Sale (ed) Coral reef fishes: dynamics and diversity in a complex ecosystem. Academic Press, San Diego, p 275-301. Forrester, G. E. and M. A. Steele. 2000. Variation in the presence and cause of densitydependent mortality in three species of reef fishes. Ecology 81:2416-2427. Malone, J. C., G. E. Forrester, and M. A. Steele. 1999. Effects of subcutaneous microtags on the growth, survival, and vulnerability to predation of small reef fishes. Journal of Experimental Marine Biology and Ecology 237:243-253. Steele, M. A. 1999. Effects of shelter and predators on reef fishes. Journal of Experimental Marine Biology and Ecology 233:65-79. Steele, M. A., G. E. Forrester, and G. R. Almany. 1998. Influences of predators and conspecifics on recruitment of a tropical and a temperate reef fish. Marine Ecology Progress Series 172:115-125. Steele, M. A. 1998. The relative importance of predation and competition in two reef fishes. Oecologia 115:222-232. Steele, M. A. 1997. Population regulation by post-settlement mortality in two temperate reef fishes. Oecologia 112:64-74. Steele, M. A. 1997. The relative importance of processes affecting recruitment of two temperate reef fishes. Ecology 78:129-145. Steele, M. A. 1996. Effects of predators on reef fishes: separating cage artifacts from effects of predation. Journal of Experimental Marine Biology and Ecology 198:249-267. Steele, M. A. 1995. The contributions of predation, competition, and recruitment to population regulation of two temperate reef fishes. Ph.D. Dissertation. University of California, Santa Barbara, California, USA. Solicited Book Reviews Steele, M. A. 1998. Life and Death of Coral Reefs. The Quarterly Review of Biology 73:367 Papers Presented at Scientific Meetings 60 total: 30 first author: 28 talks; 2 posters 30 co-author: 27 talks; 3 posters Invited Seminar Presentations 19 first author

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