NEW SPECIES OF CERAMBYCIDAE FROM THE ESTACION DE BIOLOGIA DE CHAMELA, JALISCO, MEXICO (COLEOPTERA)

NEW SPECIES OF CERAMBYCIDAE FROM THE ESTACION DE BIOLOGIA DE CHAMELA, JALISCO, MEXICO (COLEOPTERA) JoHN A. CHEMSAK * EoMUND F. GrESBERT **.,.University of California Berkeley, CA 90210 **9780 Drake Lane Tieverly Hills, CA 90210 USA Folia Entomológica Mexicana No. 69: 19-39 (1986) Recibido para publicación: 28 abril 1986. Aceptado para publicación: 29 agosto 1986. 19

20 FoLIA ENTOMOL. MEx. 69 ( 1986) ABSTRACT Eleven new species of Cerambycidae are described from the Estación de Biología de Charnela, Jalisco, México: Eburia hatsueae, Eburia perezi, Xeranoplium pubescens, Micropsyrassa doyeni, Aneflus bullocki; Megapsyrassa linsleyi, Megapsyrassa atkinsoni, Nemognathomimus michelbacheri, Oncideres albomarginata charnela, Aegomorphus chamelae, and Tetraopes ineditus. KEY WORDS: Taxonomy, Coleoptera, Cerambycidae, Mexico, Jalisco, Charnela. RESUMEN Se describen once especies nuevas de cerambícidos de la Estación de Biología de Charnela, Jalisco, México: Eburia hatsueae, Eburia perezi, X eranoplium pubescens, Micropsyrassa doyeni, Aneflus bullocki, Megapsyrassa linsleyi, Megapsyrassa atkinsoni, Nemognathomimus michelbacheri, Oncideres albomarginata charnela, Aegomorphus chamelae, y Tetraopes ineditus. PALABRAS CLAVE: Taxonomía, Coleoptera, México, Jalisco, Charnela. At the invitation of the officials of the Estación Biología de Charnela, Jalisco, México, a facility of the Universidad Nacional Autónoma de México, visits were made there in July, 1984 and 1985. The station is situated near the coast about 160 kilometers south of Puerto Vallarta. The area is densely covered with mixed thorn forest elements typical of the region. According to S. Bullock (pers. comm.) there are over 500 woody plants catalogued within the boundaries of the station. The purpose of this projects is to make known cerambycid fauna of the environs of the station with the ultimate goal of producing a checklist of this group. Also it is desirable to provide a comprehensive synoptic collection which will be of value in other studies. This paper is offered to make known sorne of the numerous undescribed species from the area to make the names available for the forthcoming checklist and to encourage more collecting. More collecting visits will be necessary at different times of the year to make the list as complete as possible. All primary typcs described here will be deposited in the collections of

CHEMSAK-GIESBERT. CERAMBYCIDAE FROM jalisco 21 the Instituto de Biología, Universidad Nacional Autónoma de México (IB UNAM). Paratypes are in the collections of the Essig Museum of Entomology, U niversity of California, Berkeley; Edmund F. Giesbert; and the Estación de Biología de Charnela. Eburia hatsueae, new species Male: Form modera te sized to large, subparallel; integument dar k reddishbrown, head and pronotum darker; eburneous fasciae whitish, basal pairs short, separated, post median pair narrowly separated, outside pair elongate, inside pair shorter; pubescen ce dense, short, appressed, areas around fasciae narrowly glabrous. Head moderate sized; from deeply impressed, medially with a transverse glabrous callus, vertex with an elevated callus between eyes; antennal tubercles prominent, obtuse at apices; pubescence dense, pale brownish, appressed; antennae more than 1-1/2 times longer than body, scape broad, cylindrical, deeply impressed dorsally at basal onehalf, coarsely punctate, rather sparsely pubescent, remaining segments moderately densely clothed with appressed pubescence, becoming finer and denser toward apex, basal segments with numerous, long, erect hairs beneath, hairs short and spare on segments seven & eight, thrid segment much longer than first, fourth and fifth subequal to third, eleventh elongate, slender, vaguely appendiculate at middle. Pronotum broader than long, sides with subacute tubercles and smaller calluses behind apex; disk with two elevated calluses before middle, a shallow elongate median callus and two flattened cal! uses toward sides near base; punctures large, deep, denser at middle around cal! uses, each puncture bearing a long, erect seta; pubescen ce dense, appressed interrupted by punctures; prosternum deeply impressed, coarsely punctate in impression, densely pubescent, intercoxal process abruptly declivous, dorsal apex tuberculate, coxal cavities wide open behind; meso- and metasternum minutely punctate, densely clothed with appressed pubescence with longer, paler, suberect hairs interspersed, scent glands distinct. Elytra a little more than 2-1/2 times as long as broad; eburneous fasciae lacking setae; each elytron bicostate between pairs of fasciae and slightly beyond toward apex; basal punctures rather fine, shallow, becoming obsolete toward apex; pubescen ce fine, appressed, erect setae absent; apices trunca te, bispinose. Legs large, slender, posterior femora not attaining elytral apices; middle and hind femora bispinose, spines short, inner slightly longer than outer, inner spines subequal to those of elytral apices; pubescence appressed, moderately dense. Abdomen minutely, densely punctate, densely clothed with

22 FoLIA ENTOMOL. MEx. 69 ( 1986) short, appressed pubescen ce, paler, suberect hairs fairly sparse; last sternite rounded at apex, lightly emarginate at middle. Lenght, 21.5-31 mm. Female: Form similar. Antennae a little longer than body, scape lightly impressed and finely punctate. Legs less robust. Abdomen with last sternite broadly rounded at apex, lightly notched medially. Lenght, 23.5-28 mm. Type material. H olotype mal e and 8 paratypes ( 4 males, 4 females) from Estación Biología de Charnela, Jalisco, México 15 July, 1982 (A. Pescador), 4 July, 1983 (S. Bullock), 22-27 July, 1984 (J. A. Chemsak, J. T. Doyen), 8-16 July, 1985 (Chemsak, H. Katsura, A. & M. Michelbacher), 10-20 July, 1985 (E. Giesbert), all taken at lights. Addition paratypes as follows: 1 female, 12 miles 5 Chivela, Oaxaca, Mexico, 18 August, 1959 (A. S. Menke, L. A. Stange); 1 male, 7 miles NE Juchitan, Oaxaca, 18 July, 1952 (E. E. Gilbert, C. D. Mac Neill); 1 male, Tehuantepec, Oaxaca, 11 June, 1964 (J. C. & D. Pallister); 1 male, 1 female, HWY 200, 51 km NE Ixtapa, Guerrero, 18-25 July, 1985 (.J. E. Wappes). Remarks. The outer post-median fasciae of the elytra are occasionally elongated and almost connect with the outer basal fasciae. The well developed eburneous surrounded by glabrous areas will readily separate this species from the others with pock-like punctures of the pronotum. A single male, probably assignable to this species, was collected at El Taladro, La Paz, Honduras, May 21, 1978, by E. Giesbert. This specimen differs from the Mexican material at and by the somewhat shorter postmedian eburneous fasciae and slightly darker integument. Etymology. This species 1s dedicated to Hatsue Katsura Chemsak for her assistance and inspiration. Eburia perezi, new species (Fig. 1) Female: From modera te sized, parallel; integument brownish testaceous, head and pronotum slightly reddish, appendages paler, elytral eburneous fasciae whitish, distinct, outside basal pair reduced or lacking, outside post median pair elongate, inside pair short, post median pairs subcontiguous, integument surrounding fasciae black, black lines narrowly extending behind post median pairs and up disk to basal pairs; pubescence fine, appressed, pale yellowish,

CHEMSAK-GIESBERT. CERAMBYCIDAE FROM jalisco 23 Fig. J. Eburia perezi Chemsak & Giesbert, f ema 1 e.

24 FoLIA ENTOMOL. MEx. 69 ( 1986) sericeus on elytra, long, erect hairs numerous. Head small, front deeply impressed transversely; median line deep, extending to margin of neck; palpi slightly unequal, apical segments barely expanded; antennal tubercles depressed, obtuse above; pubescence fine, dense, appressed; antennae about as long es elytra, sea pe slender; cylindrial, segment 3 to 10 vague! y expanded toward apices. Seape minutely punctate, moderately densely, finely pubescent, remaining segments finely, moderately densely clothed with very short, appressed pubescence, segments 2 to about 8 with long erect hairs beneath, these decreasing in number toward apex, thrid segment slightly longer than first, fourth and fifth subequal to first. Pronotum about as broad as long, sides with acute black median tubercles and two rounded, black glabrous calluses behind apex; disk with two black glabrous calluses slightly before middle and an elongate, shallow callus at middle behind dark calluses, area behind dark calluses impressed, coarsely, confluently pune tate; punctures at sides deep, scattered, each bearing a long erect seta; pubescence dense, pale, appressed; prosternum deeply impressed, deeply, sparsely punctate, finelly pubescent; intercoxal process abruptly declivous behind, coxal cavities open behind; meso- and metasternum finely punctate, dense! y clothed with appressed pubescence, long, suberect hairs numerous; scent glands distinct. Elytra less than 2-1/2 times as long as broad; eburneous fasciae with a few erect seta e; black vittae glabrous, sparsely punctate; sides above epipleura narrowly black; basal punctures shallow, moderately dense, becoming sparser toward apex, each bearing an erect seta; pubescence dense, short, appressed, lacking on fasciae and eburneous ridges; apices slightly obliquely trunca te, sutura! angles with a short spine. Legs slender; posterior femora not attaining elytral apices; middle and posterior femore with short interna! spines, spines larger than those of elytral apices. Abdomen sparsely, irregularly pune tate, dense! y pubescent, suberect hairs numerous; last sternite trunca te at apex. Length, 16-24 mm. Male: Form similar. Antennae a little longer than body. Abdomen with la~t sternite subtruncate at apex. Length, 16.5-19 mm. Type material. H olotype female and 14 paratypes ( 3 males, 11 females from Estación Biología de Charnela, Jalisco, México 8-16 July, 1985, at lights (J. A. Chemsak, H. Katsura, A. & M. Michelbacher), 4 July, 1983 (S. H. Bullock), 10-20 July, 1985 (E. Giesbert). Remarks. The black vittae of the elytra which connect the eburneous faciae make this species very distinctive. E. nigrovittata Bates differs by lacking the sharp lateral spines of the pronotum, possessing long dark calluses on the pronotum and by the prominent costae of the elytra.

CHEMSAK-GIESBERT. CERAMBYCIDAE FROM JALISCO 25 Etymology. We are pleased to dedícate this species to Luis Alfredo Pérez J., Jefe of Estación Biología de Charnela. Xeranoplium pubescens, new species Male: Form small; integument pale reddish brown, antennae and femora orange-testaceous, head and pronotum darker reddish brown; pubescence dense, grayish, appressed. Head with dense pubescence obscuring punctures, median!in e deep; severa! long, erect hairs present around mouthparts; palpi with apical segments expanded; antennal tubercles shallow; antennae extending two segments beyond elytra, third segment longer than fourth, fifth longer than third, basal segments sparsely ciliate beneath, segments densely clothed with very short, appressed pubescence. Pronotum longer than broad, sides sinuate, shallowly impressed behind middle; disk with an elongate, glabrous, median callus an a small round one on each side anterior to median one; punctures minute, sides with large puntures sparsely interspersed; pubescence dense, appressed, sides with a few, long, erect setae; prosternum impressed before coxae, impressed areas coarsely punctate, pubescence dense; meso- and metasternum micro-punctate, densely pubescent, median line of metasternum glabrous. Scutellum densely pubescent. Elytra slightly over 2-1/2 times longer than broad; moderately coarsely, separately punctae at base, punctures becoming smaller towar apex; pubescence dense, short, appressed, with short, suberect hairs numerous; apices rounded. Legs modera te; femora minute! y punctate, densely pubescent; tibiae and tarsi dark brownish. Abdomen minutely punctate, dense! y pubescent; last sternite rounded at apex. Lenght, 8-11 mm. Female: Form similar. Antennae slightly longer than body. Prosternum barely impressed before coxae. Abdomen with last sternite broadly rounded at apex. Length, 10-13 mm. Type material. Holotype ma!e and 9 paratypes (3 males, 6 females) from Estación Biología de Charnela, Jalisco, México 6-8 July, 1985, at lights (J. Chemsak, H. Katsura, H. & M. Michelbacher), 28 July to 2 August, 1984 (J. Chemsak, S. T. Doyen), 10-20 July, 1985 (E. Giesbert). An additional male paratype from 7 miles NE Barra de Navidad, Jalisco, México, 25 July 1983 R. L. Westcott). Remarks. This species 1s related to X. flavofemoratum Chemsak & Linsley.

26 FoLIA ENTOMOL. MEx. 69 ( 1986) The thicker, denser pubescence, pale antennae and paler integument will separate X. pubescens. Etymology. The name is derived from the dense pubescence of the body. Micropsyrassa doyeni, new species (Fig. 2) Male: Form small; integument shining, color reddish, elytra with a broad testaceous median band, black behind pale band and with narrow angulate, obligue black fasciae at anterior margins of median pale band. Head with vertex coarsely reticulate punctate; long, erect hairs sparse on front and beneath; palpi with apical segments very slightly expanded; antennae extending about 3-1/2 segments beyond elytra, segments three to eight dorsal! y carinate, third segment shorter than fourth, fifth longer than fourth, spine of third segment at lcast 1/2 as long as fourth segment, fourth segment prominently spined. fifth with a distinct short spine, basal segments shining moderately densely clothed with short, appressed pubescence, erect hairs long, numerous. Pronotum longer than broad; disk coarsely reticulate-punctate except for central smooth area behind middle; pubescence at base fine, whitish, appressed, long, erect hairs numerous; prosternum coarsely punctate on posterior one-half; meso- and metasternum sparsely punctate. Elytra about 2-1/2 times as long as broad; basal punctures vaguely asperate, well separated, punctures behind smaller and sparser, each puncture bearing a long, erect seta, short, subdepressed hairs present over apical one-half; apices trunca te, very shallowly emarginate. Legs sparsely clothed with long, erect hairs and short subdepresser hairs. Abdomen sparsely punctate and pubescent; last sternite broadly, shal lowly rounded at apex. Lenght, 7-9 mm. Female: Form similar. Antennae slightly longer than body. Abdomen with last sternite subtruncate at apex. Length, 5-12 mm. Type material. Holotype male, allotype and 36 paratypes ( 15 males, 21 females) from Estación Biología de Charnela, Jalisco, México 20-27 July, 1984, at lights (J. A. Chemsak, J. T. Doyen), 8-16 July, 1985 (J. A. Chemsak, H. Katsura, A. & M. Michelbacher), 10-20 July, 1985 (E. Giesbert). Remarks. This species is very similar in appearance to M. stellata Martins & Chemsak. They can be readily separated by the obligue black bands anterior

CHEMSAK-GIESBERT. CERAMBYCIDAE FROM jalisco 27 Fig. 2. Micropsyrassa doyeni Chemsak & Giesbert, maje.

28 FoLIA ENTOMOL. MEx. 69 ( 1986) to the pale fasciae in A1. doyeni. The median pale fasciae of M. stellata are only about 1/2 as broad as in M. doyeni. Also the third segment of the antennae is longer than the fourth in M. stellata, shorter in M. doyeni. The rounded oblique pale fasciae of the elytra, medially broadened sides of the pronotum, and shorter antennal spines will separate M. bimaculata (Bates) from M. doyeni. Etymology. It is a pleasure to dedicate this species to J. T. Doyen for his efforts in collecting Cerambycidae Aneflus (Aneflus) bullocki, new species Male: Form large, robust, somewhat compressed; integument dark reddishbrown, antennae piceous; pubescen ce dense, short, pale, subdepressed. Head rather fine! y, confluently punctate; antennal tubercles depressed, apices produced, blunt; pubescence modera te! y dense, depressed; antennae stout, longer than body, segments 3 to 8 spinose at apices, segments 3 to 5 dentate externally at apices, segments from fifth carinate dorsally, basal segments shining, moderately densely clothed with pale pubescence, outer segments opaque, minutely pubescent, scape stout, cylindrical, longer than third segment, fourth shorter than third, fifth equal to scape, eleventh longer than tenth, strongly appendiculate. Pronotum broader than long, sides irregularly rounded; apex narrowly impressed at sides, impression becoming very broad toward middle of disk, base narrowly impressed; disk with four obtuse tubercles around middle, middle vague! y, longitudinal! y callused; punctures coarse, confluent, rugose; pubescent moderate, short, subdepressed; prosternum deeply impressed, shallowly, rugoseli punctate over basal two-thirds, apical one-third subglabros, finely, transversely plicate, pubescence rather sparse, coxal cavities open behind; mesosternum subopaque, sparsely punctate and pubescent, surface micropunctate; metasternum minutely densely punctata except middle, dense! y clothed with appressed pubescence, middle glabros. Scutellum densely yellowish pubescen t. Elytra Iess than three times as long as broad; each elytron tricostate, humeral pair short; basal punctures rather fine, confluent, punctures becoming finer and contiguous toward apex; pubescen ce dense, short, subdepressed, vaguely, longitudinally interrupted along costate, a few erect hairs arranged on costae; apices obliquely subtruncate, angles with short spines. Legs large, slender; femora densely, shallowly punctate, moderately densely pubescent. Abdomen finely, densely punctate at sides, moderately densely pubescent, middle sparsely punctate and pubescent; apex of last sternite rounded. Lenght, 38-41 mm.

CHEMSAK-G!ESBERT. CERAMBYCIDAE FROM JALISCO 29 Type material. H olotype mal e and 5 mal e paratypes from Charnela, Jalisco, México, 12 November 1983 (S. H. Bullock), 14 November, 1983 (Bullock), 23 November, 1983 (Bullock), 15 September, 1983 (Bullock), 15 September, 1983 (Bullock), all collected at lights. Remarks. This is one of the largest and most distinctive species in the genus. The large size, bispinose third antena! segment, and rather dense subdepressed pubescence of the elytra which is vaguely interrupted by longitudinal lines along the costae will separate bullocki from other known Aneflus. In the key to the Mexican species of Aneflus (Cremsak & Linsley, 1963),A. bullocki will come out in the couple with A. sonoranus Casey and A. levettei (Casey). The combination of characters listed above will readily separate A. bullocki from those species. If the character of longitudinal, glabrous lines is not utilized ( couplet 5), A. bullocki will key out with A. calvatus Horn. The much denser, coarser pubescence, larger size, and prominently callused pronota! disk of A. bullocki will distinguish th two. Etymology. We are pleased to dedícate this species to S. H. Bullock. Megapsyrassa linsleyi, new species (Fig. 3) Male: From moderate sized, subcylindrical; integument dark reddish brown; pubescence sparse, yellowish. Head coarsely, irregularly punctate on vertex, with a shallow depression between eyes, pubescen ce sparse, short, depressed; antennae almost as long as elytra, segments three to seven spined at apices, spine of thrid segment rather short, about 1/4 as long as fourth segment, basal segments sparsely clothed with short, appressed pubescence, becoming denser and shorter toward apex, long, erect hairs numerous on basal segments, restricted to apices at segments eight to ten, third segment longer than fourth, fifth longer than fourth but shorter than thrid, eleventh longer than tenth, appendiculate. Pronotum about as long as broad, sides sinuately rounded; apex narrowly impressed, base broadly impressed; disk with an irregular, longitudinal glabrous, impunctate area at middle; punctures around middle moderately coarse, irregular, subconfluent, coarse and confluent at si des; pubescen ce consisting of long erect setae arising out of punctures; prosternum subglabrous, transversely rugose before coxae, transversely plicate near apex, pubescence sparse, erect, prosternal process moderately expanded at apex, coxal cavities open behind; mesosternum micropunctate, dense! y clothed with very fine,

30 FoLIA ENTOMOL. MEx. 69 ( 1986) Fig. 3. Megapsyrassa linsleyi Chemsak & Giesbert, male.

CHEMSAK-GIESBERT. CERAMBYCIDAE FROM jalisco 31 appressed, grayish pubescen ce; metasternum sparsely pune tate medial! y, si des densely clothed with pale silky pubescense. Scutellum densely, very finely pubescen t. Elytra almost 3 1/2 times as lond as broad; each elytron vague! y bicostate on disk; punctures at base moderately coarse, separated, becoming denser behind and finer and sparser near apex; pubescen ce very short, depressed, with longer, suberect hairs sparsely interspersed; apices obliquely truncate, strongly bispinose, outer spines large. Legs modera te; femora coarsely, confluently punctate, moderately clothed with yellowish, subdepressed pubescence. Abdomen sparsely punctate and pubescent; last sternite trunca te to shallowly emarginate at apex. Length, 24 27 mm. Female: Form similar. Antennae shorter than elytra. Abdomen with last sternite rounded at apex. Length, 27-29 mm. Type material. Holtype male and 43 paratypes (28 males, 15 females from Estación Biología de Charnela, 20-27 July, 1984, at lights (.J.A. Chemsak, J. T. Doyen), 8-16 July, 1985, at lights (J. A. Chemsak, H. Katsura, A. & M. Michelbacher), 10-20 July, 1985 (E. Giesbert), 16 June and 4 July, 1983, (S. H. Bullock). An additional male paratype from 6 km SE Charnela, 23 July, 1978 (J. O. Matson). Remarks. The smaller spine of the third antennal segment, shorter ercct setae and presence of small depressed hairs on the elytra will separate this species from M. xestioides (Bates). M. linsleyi differs from M. auricomis Chemsak & Linsley and M. puncticollis Chemsak & Linsley by the much sparser pubescence, shorter antennal spines and open front coxal cavities. Etymology. In recognition of his many contributions to the study of Cerambycidae we dedícate this species to E. G. Linsley. Megapsyrassa atkinsoni, new species Male: Form modera te sized, subcylindrical; integument oragne-testaceus, nitid, with antennae except scapte, tibiase, and tarsi orange-brown to piceous; pubescen ce sparse, golden. Head sparsely punctate on vertex with an indistinct depression between eyes, pubescence very sparse, inconspicuous; antennae about as long as body, segment 3 to 7 spined at apices, spine of third segment long, 1/ to 2/3 as long as fourth segment, spine of fourth segment somewhat shorter, basal segments obtusely carinate, thinly clothed with fine, short, appres-

32 FOLlA ENTOMOL. MEX. 69 ( 1986) sed pubescence, which is denser on remaining segments, long erect hairs numerous on inside and at apices of basal segments, sparser on distal segments, third segment subequal to fourth, shorter than fifth, eleventh longer than tenth, feebly appendiculate. Pronotum longer than broad, sides subparallel, slightly tumid laterally behind narrow apical impression, base broadly impressed; disk shining, somewhat flattened, very sparsely, irregular! y punctate, glabrous except for sparse, long, erect setae arising from punctures; prosternum micropubescent and coarsely punctate befare coxae, transversely plicate near apex, coxal cavities closed behind; mesosternum finely micropubescent; metasternum shining, nearly impunctate, sides clothed with very fine, pale, silky appressed pubescence. Scutellum clothed with very fine pubescence. Elytra about 3 times as long as broad; surface shining, sparsely unevenly punctate, with coarser punctures forming vague, widely spaced longitudinal lines; pubescence consisting of sparse, long, suberect hairs arising from punctures; a pices emarginate with hind angles denta te. Legs modera te; femora sparsely punctate with moderately sparse, short, subdepressed, and long erect hairs mixed; tibiae strongly carinate. Abdomen shining, sparsely punctate and pubescent; terminal sternite truncate to shallowly emarginate at apex. Length 13.5-19.5 mm. Female: Form similar to male. Antennae shorter than elytra. Prosternum with punctation befare coxae much reduced. Abdomen with terminal sternite roundad at apex. Length 19 mm. Type material. H olotype mal e and 2 male paratypes from Estación Biología de Charnela, 10-20 July, 1985, at lights (E. Giesbert). One maje paratype from San Jerónimo, Chiapas, México, 21 August, 1971. Six additional specimens, 5 males, 1 female from "La Pacífica", 3 km NW Cañas, Guanacaste Prov., Costa Rica, 9-12 June, 1978, 2-7 ]une, 1980, and 8 June (E. Giesbert). Remarks. The smaller size, orange-testaceous integument, and dentate elytral apices will separate this species from others in the genus. The Costa Rican population exhibits somewhat less melanism of the appendages (redbrown rather than piceous), slightly longer antennal spines, and a more thickly pubescent scutellum. Etymology. We take pleasure in dedicating this species to Dr. Thomas H. Atkinson.

CHEMS.AK-GIESBERT. CERAMBYCIDAE FROM JALISCO 33 Nemognathomimus michelbacheri, new species (Fig. 4) Male: Forro small; integument black, pronotum orange, elytra occasionally orange or partly orange. Head with front short, about as broad as long; frons short, subparallel; median line short, shallow; vertex broad, fin el y punctate; neck abruptly constricted behind eyes; pubescence modera te, golden, subappressed; eyes large, notchcd, widely separated above; antennae inserted on sides of front at top of inner margin of lower lobe, extending to about third abdominal segment, outer segments thickened, short, basal segments shining, clothed with dark, subdepressed setae, segments from seventh densely clothed with short, dark, depressed hairs, third segment equal to scape, fourth shorter than third, outer segments slightly longer than broad. Pronotum broader than long, hind angles acute, extending over humeri; front margin very narrow, very narrowly constricted; basal margin broadly lobed at middle, shallowly impressed at sides of middle; disk convex, shallowly inflated behind apex, densely, moderately coarsely punctate except for vague median line behind middle; pubescence golden, depressed, moderately dense; prosternum subglabrous; meso- and metasternum finely, densely punctate, densely clothed with depressed silvery pubescence. Elytra slightly more than two times longer than broad, tapering, sides shallowly impressed near middle; punctures larger than those on pronotum. dense, separated; pubescence dense, dar k, subdepressd; apices narrowly rounded, slightly dehiscent. Legs slender; posterior tarsi longer than tibiae, firts segment longer than remaining segments together, third segment slender, cleft to middle. Abdomen finely, densely punctate at sides, densely clothed with fine silvery pubescence; last sternite emarginate at apex. Lengt, 6-9 mm. Female. Forro similar, slightly more robust. Integument except antennae, apices of tibiae and tarsi usually orange. Antennae extending to first abdominal segment. Abdomen sparsely puncta te and pubescent; last sternite truncate at apex. Length, 7-8 mm. Type material. H olotype mal e, allotype and 17 paratypes ( 11 males, 6 females) from Estación Biología de Charnela, 8-16 July, 1985 (J. Chemsak, H. Katsura, aa & M. Michelbacher); 10-20 July, 1985 (E. Giesbert). Remarks. The color variation of this species is similar to that of N. pallidulus (Linsley). The female are usually orange with black antennae and partially black tarsi and tibiae. There is an occasional partial infuscation of the elytra. The males are usually black with orange pronota. In sorne cases the elytra are orange or partially orange. This species resembles adults of the meloid, Ne-

34 FoLIA ENTOMOL. MEx. 69 ( 1986) Fig. 4. Nemognat h omzmus. michelbacheri Chemsak & Giesbert, maje.

CHEMSAK-GIESBERT. CERAMRYCIDAE FROM jalisco 35 mognatha. N. michelbacheri differs from N. pallidulus by the shorter, broader body size, densely punctate pronotum, shorter, subdepressed pubescence, and shining rather than dull elytra. Adults of N. michelbacheri are found on flowers of Croton on the grounds of the Estación. Etymology. We dedícate this species to A. E. Michelbacher for his friendship, support, and inspirational efforts in the field. Oncidercs albomarginata charnela, new subspecies Form and s1ze of O. albomarginata albomarginata Thomson. Pubescence grayish, head and pronotum brownish variegated lf elytra basally orange-brown, disk densely grayish pubescent with numerous irregular orange patches interspersed, narrow, angulate, brownish, transverse fasciae present behind middle, lateral white bands very densely pubescent, humeri dark brownish pubescent beneath, underside densely grayish pubescent medially, sides brownish variega ted, legs rather uniformly brownish pubescent with flecks of grayish interspersed. Elytra with small, glabrous asperites sparsely interspersed on anterior portions of low gibbosities and on sides behind humeri, punctures behind small, interspersed over grayish portions, larger, glabrous puntures outlining white lateral vittae sparse. Scutellum densely brownish pubescent. Length. Type material. Holotype male from Estación Biología de Charnela, Jalisco. 12 October, 1983 (S. H. Bullock). Paratypes, all from the same locality as follow: 3 miles, 1 female, 14 November, 1982 (Bullock); 1 male, 27 Novem ber, 1983 (Bullock); 1 male, 18 November, 1983 (Bullock); 1 male, 2 December, 1983 (Bullock); 1 male, 5 December, 1983 (Bullock); 1 female, 12 October, 1984 (Bullock). Additional paratypes incluye 3 males, 1 female reared from Spondias purpurea and Magnifera indica by Atkinson and Noguera. Those adults emerged in December and April..\dditional specimens assignable to this population include 1 male, Nayarit, November, 1977; 1 male, Candelaria Loxicha, Oaxaca, 12 September, 1973 (E. Welling); 1 female, Ocozocoautla, Chiapas, 26-28 September, 1971 (D. E. Breedlove) and 1 male, Selva del Ocotc, 37 km NW Ocozocoautla, 29 August, 1972 (Beedlove). Remarks. This subspecies may be readily separated from the typical form by the dense grayish pubescence of the elytra with the narrow transverse, brownish fasciae. The basal asperites are fewer and smaller in number and the

36 FoLIA ENTOMOL. MEx. 69 ( 1986) remammg punctures smaller and sparser. The lateral white fasciae are also outlined by fewer glabrous punctures. Etymology. The name is a noun in opposition pertaining to the locality. Acgomorphus chamelac, new species (Fig. 5) Male: Form modera te sized, slightly tapering; integument piceous, antennae pale annulate; pubescence dense, short, appressed, black, brownish and grayish. Head with front feebly convex, rectangular, sparsely, irregular punctate, browni>h pubescent; vertex convex, punctate along eyes, brownish pubescent; eyes small, rather finely faceted, lo bes connected by a!in e; genae subparallel, as long as lower eye lo bes; antennae about as long as body, thrid segment white biannulate, segments from fourth pale annulate at basal halves, basal segments lightly fringed beneath, tenth and eleventh segments lightly fringed, partially glabrous beneath. Pronotum broader than long, lateral tubercles large, broad, obtuse at apices; disk with an elongate median, globrous callus which narrowly extends to apical margin, each side with a large tumid callus; punctures sparse, located along margins and around calluses; pubescence dense appressed, mottled pale and dar k brownish with a gray spot at middle on basal margin; prosternum finely pale brownish pubescent, intercoxal process arcuate; meso- and metasternum finely, densely pubescent, mesosternal process declivous. Elytra slightly more than 1-1/2 times as long as broad; basal gibbosities very shallow, broad; basal punctures coarse, sparse, irregular, denser behind scutellum; pubescence dense, fine, appressed, mottled brownish and gray with small, black spots irregularly interspersed, two larger black spots behind middle and two near apex, usually with vague whitish fasciae before middle and at apical one-third, suberect setae very short; apices emarginate- truncate. Legs stout, femora strongly clava te; tibiae pale triannulate; tarsi pale pubescent, front pair fringed lateral! y. Abdomen finely densely pubescent, medial! y subglabrous; apex of last sternite emarginate. Length, 16-24 mm. Female: Form similar, sides subparallel. Antennae shorter than body apical segments not fringed nor glabrous beneath. Legs less robuts, front tarsi not expanded nor fringed. Abdomen with als sternite subtruncate at apex, with a narroy median line. Length, 16-21 mm. Typc material. Ji olotype m ale and 31 paratypes ( 14 males, 17 females) from Estación Biología c~c Chamela, Jalisco, México 8-16 July 1985 (Chemsak, H.

CHEMSAK-GJESBERT. CERAMBYCIDAE FROM JALISCO 37 Fig. 5. Aegomorphus chamelae Chemsak & e 1es 1 Jcrt, female.

38 FoLIA ENTOMOL. MEx. 69 ( 1986) Katsura, A. & M. Michelbacher); 20-22 July, 1984 (Chemsak & J. Doyen), 23 June, 1984, 29 June, 1983, 18 May, 1983, 4 July, 1983 (S. Bullock), 22 June, 1985 (F. Noguera); 10-20 July, 1985 (E. Giesbert). Remarks. The type series is fairly uniform in coloration and pattern. U sually two whitish fasciae are evident on the elytra to the naked eye. These are rather vague and not well differentiated. The color pattern, robust size and shape and the broad tubercles of the pronotum will separate this species from i ts relatives. Etymology. The name is geographical indicating the occurrence of this species. Tetraopes ineditus, new species Tetraopes comes Chemsak (not Bates), 1963, Univ. California Pubs. Entorno!., 30:27, fig. 5. Male: Form stout, robust, sides subparallel; integument black, elytra partially reddish; pubescence dense, short, cinereous* erect hairs short, dense. Head moderately coarsely, irregular! y, sparsely punctate; pubescence dense, blucgray with brownish overtones in obligue light, short, erect hairs numerous; antennae black, slightly shorter than elytra, scape sparsely clothed above with short, appressed, blue-gray pubescence, segments from first densely clothed beneath by short, appressed, blue-gray pubescence, segments from third clothed above with very short, brownish pubescence, segments with long, erect ciliae beneath. Pronotum black, lacking black spots; umbone abruptly elevated, prominent, longer than broad, delimited posterior! y by a deep sulcus; si des broadly rounded, deeply impressed at base of umbone; punctures modera te sized, dense toward inner half of umbone, scattered at sides; pubescence dense, blue-gray, short with long, erect, black hairs numerous, forming a dense brush on umbone; prosternum dense! y blue-gray pubescent; mesoand metasternum densely clothed with appressed blue-gray pubescence. Scutellum black. Elytra about twice as long as broad; slightly more than basal onc-half reddish; humeri with black spots, disk two black spots before middle, each about egua] in diameter to lenght of eighth antennal segment; apex black, chevron beginning at about middle at suture and arching back toward margins, each side before margin with a small black spot at anterior margin of chevron; punctures moderately coarse, somewhat serially arranged, rows well separated, punctures becoming finer on black portions; pubescence dense, very short, appressed, grayish on red part and blue-gray on black,

CHEMSAK-Gn:SBERT. CERAMBYCIDAE FROM jalisco 39 erect black hairs numerous, longer at base; apices rounded. Legs short, dense! y blue-gray pubescent; tibiae with erect, black setae along outside edge. Abdomen densely blue-gray pubesccnt; last segment with numerous, long, erect setae, rounded at apex. Length, 7-5-12 mm. Female: Form more robust. Antennae extending to about third abdominal segment. Abdomen with last sternite broadly subtruncate at apex, linearly impressed at middle. Length, 11-13 mm. Type material. H olotype male and 11 paratypes (9 males, 2 females) from Estación Biología de Charnela, Jalisco, México 8-16 July, 1985 (J. Chemsak, H. Katsura, A. & M. Michelbacher); 10-20.July, 1985 (E. Giesbert). Additional paratypes incluye: 1 male, Suchiapa, Chiapas, Mexico, 18 July, 1957 (J. Chemsak); 1 female, Playa de Oro Rd., 19 miles NW Manzanillo, Colima, México, 25 August, 1970. Remarks. This species differs from T. paracomes Chemsak and T. comes Bates by the broader, suturally anteriorly directed black chevron of the alytra and prescence of two black spots at the anterior edge of the chevron. The elytral punctures are finer and subserially arranged in T. ineditus, T. cleroides Thomson differs by the small apical, dark chevron which is directed back from the suture. The discal black spots are smaller, the humeral spots reduced and the elytral punctures more irregular in T. cleroides. Adults of T. ineditus are associated with an asclepiad vine on the grounds of the station at Charnela. Etymology. The term is Latin for unpublished. AcKNOWLEDGEMENTs We are grateful to Luis Alfredo Pérez J., S. Bullock, and T. Atkinson for the opportuning to undertake this project. For field assistance and collecting efforts, thanks are due to Sonia G. Navarro Pérez, J. T. Doyen, H. Katsura Chemsak and A. & M. Michelbacher. LITERATURA CITED CHEMSAK, ]ohn A. 1963. Taxonomy and bionomics of the genus Tetraopes. Univ. California Pubs. Entorno!., 30:1-90, 17 figs., 9 pis. CHEMSAK, ]ohn, A. AND E. G. LINSLEY. 1963. Synopsis of the known Mexican species of Aneflus. Bu/l. Brooklyn Entorno!. S oc., 63:80-96, 3 pis.