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Larry Kuehn Genetic Prediction Preliminary estimates of breed differences from recent sampling in the Germplasm Evaluation Project Larry Kuehn, Warren Snelling, Mark Thallman Research Geneticists Uses USMARC Germplasm Evaluation () data to adjust for breed differences The USDA is an equal opportunity employer. SIRE BREEDS USED IN THE GERMPLASM EVALUATION PROGRAM AT THE USMARC Cycle II 73-74 Jersey S. Devon Limousin Simmental Charolais Red Poll Braunvieh Gelbvieh Maine Anj. Chianina Cycle III Cycle IV 75-76 86-9 Cycle VI 97-98 F1 Crosses ( or dams) a Brahman Sahiwal Pinzgauer Tarentaise 3-way crosses Brahman Brangus Devon Santa Gertrudis Holstein Cycle V 92-94 Longhorn Salers Galloway Nellore Shorthorn Piedmontese Charolais Gelbvieh Pinzgauer Tuli Boran Belg. Blue Brahman Piedmontese Across-breed EPD adjustment factors have been computed for BIF since 1993 Account for differences in EPD base in the genetic evaluations of each breed involved USDA, ARS, U.S. Meat Animal Research Center Cycle I 7-72 Background - ABEPD Wagyu Norweg. Red Sw. Red&Wh. Friesian a Sire breeds mated to and females, and Composite MARC III (1/4,, Red Poll and Pinzgauer) cows in Cycles V, VI, VII and VIII. Cycle VII Cycle VIII 99-1-2 Red Limousin Charolais Simmental Gelbvieh Beefmaster Brangus Bonsmara Romosinuano Across EPD adjustment factors Calculated by scaling the current breed difference (Bi) to EPD base Current breed difference is 211 this year Base difference calculated by subtracting the 211 average EPD from the 211 breed average EPD For i: MARC III Cows Factori = Bi - B - (EPDi - EPD) Adjusting sample for current EPD Across-breed EPD program Estimate breed differences (Bi) from using animal model Adjust records for bull EPD Bi = USMARC i / b + (EPDi,YY EPDi,USMARC ) EPDi,YY is the breed average EPD (current) EPDi,USMARC weighted average USMARC sire EPD b is a scaling factor to convert USMARC (USMARCi) solution to an industry scale 1

14 12 1 8 6 4 2-2 -4 1972 1974 Gene$c Trends for Birth Weight, lb 1976 1978 198 1982 1984 1986 1988 Red Simmental Gelbvieh Adapted from Spring 213 Genetic Trends from Associations 2 22 24 26 28 21 7 6 5 4 3 2 1-1 -2 1972 Gene$c Trends for Weaning Weight, lb 1974 1976 1978 198 1982 1984 1986 1988 Red Simmental Gelbvieh Adapted from Spring 213 Genetic Trends from Associations 2 22 24 26 28 21 1 75 5 25-25 -5 1972 1974 Gene$c Trends for Yearling Weight, lb 1976 1978 198 1982 1984 1986 1988 Red Simmental Gelbvieh Adapted from Spring 213 Genetic Trends from Associations 2 22 24 26 28 21 35 3 25 2 15 1 5-5 -1-15 Gene$c Trends for Maternal Milk, lb 2 Red Simmental Gelbvieh Adapted from Spring 213 Genetic Trends from Associations 22 24 26 28 21 differences Some inquiry into ABEPD system Since we re using the whole database, are we biasing current breed difference with old bulls? We rely on the genetic trend to estimate current breed differences from older bulls Use weighted average of EPDs from bulls we ve used to adjust USMARC data 2

differences Can we address concern by looking only at recent sampling for Continuous evaluation of in 26 Shift from exotic germplasm to influential breeds Began resampling breeds 14 of 18 in 26-27 Added Brangus and Beefmaster in 28-29 Added South Devon and Tarentaise in 211 Target Population Structure AI Sires: AN, HH, SM, CH, AR, LM, GV, SH, BN, BM, MA, BR, CI, SG, SA, BV PB & F 1 Steers PB & F 1 Bulls Dams: AN, HH, SM, CH, AR, LM, GV, SH, BN, BM, MA, BR, CI, SG, SA, BV PB & F 1 Heifers Natural Service PB, F 1, & F 1 2 Steers & Heifers Current AI-sired Calves Produced 1/2 3/4 7/8 PB Total 131 7 16 25 422 Beefmaster 178 3 1 29 Brahman 189 43 232 Brangus 184 38 3 225 Braunvieh 185 61 7 253 Charolais 92 46 14 243 295 Chiangus 184 76 8 268 Gelbvieh 97 139 51 2 289 132 45 8 235 42 Limousin 95 14 48 2 285 Maine Anjou 223 55 3 281 Red 95 134 46 5 28 Salers 184 77 4 265 Santa Gertrudis 181 68 7 256 Shorthorn 2 86 7 293 Simmental 14 58 11 19 399 Total 249 1166 234 882 4772 Analysis of data only Compared to cumulative data Used same analysis as ABEPD Results are shown relative to of sire differences 1/2 of actual breed effect Shown as differences from These results are PRELIMINARY They will likely change soon Birth Weight breed of sire effect... 4.4 4.5 -.1 Red.2.8 -.7 Shorthorn 5.1 6.5-1.3 Beefmaster 3.5 4.8-1.4 Brahman 11.9 11.1.8 Brangus 3.4 3.5 -.1 Santa Gertrudis 4.7 5.4 -.6 Braunvieh 2.3 2.6 -.3 Charolais 7.1 7.5 -.4 Chiangus 3. 3.6 -.6 Gelbvieh 2.5 2.4.1 Limousin 2.6 3.6 -.9 Maine Anjou 2.2 4.5-2.3 Salers.2 1.8-1.6 Simmental 3.8 4.3 -.5 Weaning Wt breed of sire effect... -21.5-5.5-16. Red -11.5-15.4 3.9 Shorthorn -28.6-2.4-8.2 Beefmaster -8.3-1.3-7. Brahman 4.2 1.8-6.6 Brangus -11.6-8.7-2.9 Santa Gertrudis -9.4-6.5-2.9 Braunvieh -26.6-27.5.9 Charolais 1.9 15.5-13.6 Chiangus -26.2-3.7 4.5 Gelbvieh -14. -1.5-12.4 Limousin -9.2-2.3-6.9 Maine Anjou -22.1-22.9.8 Salers -17. -1.5-6.5 Simmental 4.1 9.2-5. 3

Yearling Wt breed of sire effect... -44.4-35.6-8.8 Red -28.7-32.3 3.6 Shorthorn -42.2-22.4-19.8 Beefmaster -44.1-42.5-1.7 Brahman -42.8-56. 13.2 Brangus -44.7-36.9-7.8 Santa Gertrudis -29.5-31.4 1.9 Braunvieh -59.6-63.5 3.9 Charolais -17.9 2.4-2.3 Chiangus -55.3-58.3 3. Gelbvieh -1.8-18.2 7.3 Limousin -38.1-37.7 -.5 Maine Anjou -55.6-44.5-11.1 Salers -49.6-25.9-23.7 Simmental 1.2-6.5 7.7 Milk breed of sire effect... -13.3-22.1 8.7 Red -7.8-7. -.9 Shorthorn 1.1 -.5 1.6 Beefmaster 5.5-11.1 16.6 Brahman 2.2 6.6-4.4 Brangus 1.8-6. 7.8 Santa Gertrudis -.4-9.9 9.4 Braunvieh 22. 11.1 1.9 Charolais -7.4-9.2 1.8 Chiangus -3.7-7.4 3.6 Gelbvieh 24.6 6.1 18.6 Limousin.4-9.6 1.1 Maine Anjou -21.3-1.1-11.2 Salers 3.7-1.3 5. Simmental 3.4 -.9 4.3 Marbling breed of sire effect... -.91 -.73 -.18 Red -.43 -.38 -.5 Shorthorn -.75 -.64 -.11 Santa Gertrudis -1.8-1.12.5 Braunvieh -.59 -.63.4 Charolais -.93 -.87 -.6 Chiangus -.66 -.72.5 Gelbvieh -.87 -.83 -.4 Limousin -1.17-1.19.2 Maine Anjou -1.23-1.9 -.13 Salers -.27 -.36.9 Simmental -.76 -.8.4 Ribeye area breed of sire effect... -.26 -.25 -.1 Red -.5 -.35.31 Shorthorn.23 -.14.37 Santa Gertrudis -.21 -.46.25 Braunvieh.89.51.38 Charolais 1.39.8.59 Chiangus.37.12.25 Gelbvieh.94.66.28 Limousin 1.45 1.21.24 Maine Anjou.59.68 -.8 Salers.6.4.2 Simmental.85.7.16 Fat thickness breed of sire effect... -.87 -.58 -.29 Red -.35 -.41.6 Shorthorn -.174 -.162 -.12 Santa Gertrudis -.112 -.124.12 Braunvieh -.192 -.179 -.14 Charolais -.211 -.23.19 Chiangus -.155 -.162.7 Gelbvieh -.16 -.189.29 Maine Anjou -.246 -.239 -.7 Salers -.199 -.217.18 Simmental -.163 -.29.45 Why differences? EPD on bulls not accounting for differences in sampling between old cycles of and new sampling TREND and connectedness Accuracy structure of new Cycle VII not included s samples of bulls better connected Error structure of breed differences (USMARC) and EPDs (accuracy) Need to look further! 4

BREED GROUP MEANS (DEVIATIONS FROM HA & AH) FOR MATURE WEIGHT (ADJUSTED TO CONDITION SCORE OF 5.5) OF F1 CROSS COWS IN CYCLES I AND II (BIRTH YEARS: 197-74) COMPARED TO CYCLE VII (BIRTH YEARS 1999-2), LB erences in Cow Weights 16 14 () (- 6) (- 11) (- 92) (- 24) (- 44) 1412 146 141 1388 1368 132 Are we changing efficiency through selection for growth? 12 1 8 6 () 121 (64) (75) (33) 185 196 154 (118) 1137 Cycle I & II Cycle VII 4 HA + AH Red Ang Simm Gelb Lim Char Cow weight differences same? Compare Cycle VII females to new sampling ~8 years between bull sampling Oldest cows only 5.5 in new sampling Mature weight limited Earlier weights are a proxy (highly correlated) Weights at ~55 and 92 d (1.5, 2.5 yr) Data not precise yet but give an indication Palpation weight (~1.5 yr) weight Cycle VII weight 95 923 913 956 Red 921 921 Charolais 952 969 Gelbvieh 96 97 Limousin 916 912 Simmental 949 979 Beefmaster 917 Brahman 95 Brangus 941 Braunvieh 87 Chiangus 886 Maine Anjou 895 Santa Gertrudis 948 Salers 88 Shorthorn 926 Palpation weight (~2.5 yr) weight Cycle VII weight 1192 1166 1171 1199 Red 1154 1141 Charolais 1225 1211 Gelbvieh 1183 112 Limousin 1144 1154 Simmental 1178 1213 Beefmaster 1126 Brahman 1187 Brangus 1134 Braunvieh 991 Chiangus 1124 Maine Anjou 1171 Santa Gertrudis 115 Salers 1126 Shorthorn 1111 Cow Weights Some breeds have moderated while others are larger than at Cycle VII Seems to be a real opportunity for breed complementarity These results are preliminary Would like a few more years of data. 5

Thoughts Questions Demonstrates need for continued research for breed differences USMARC not enough More to the country than central Nebraska Southern location most important Use of common sires between locations will strengthen overall power Exploring options to improve delivery of ABEPD 6