PURSE-SEINE AND BOAT-SEINE {JHANGU VALA) FISHERY FOR THE OIL SARDINE OFF COCHIN Central Marine Fisheries Research Institute, Cochin-18

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PURSE-SEINE AND BOAT-SEINE {JHANGU VALA) FISHERY FOR THE OIL SARDINE OFF COCHIN-1968-1971 V. BALAN' Central Marine Fisheries Research Institute, Cochin-18 ABSTRACT The data on the catches,fishingeflfort, length and sex composition relating to the purse seinefisheryfor the oil sardine, Sardinella longiceps V^l. off Cochin, during the three seasons from 1968-69 to -71 are presented. A comparison of catch rates of the sardine landed by this gear with those of the boat seine thmgu vala during the same period has been made. A comparative study of the length range, the modal and the mean length distributions of the oil sardine catches obtained in the two gear revealed a fairly good similarity. Application of Chi-square test on the sex composition data of the sardine shoalsfishedby purse seine during 1969-71 indicated that there was no significant dominance of either sex- INTRODUCTION The exploitation of the oil sardine fishery around Cochin has hitherto depended solely on the use of traditional gear such as thaugu vala and chala vala Recently, the purse seine was introduced in this area by the Indo-Norwegian Project for the oil sardine and mackerel fisheries. The traditional gear are usually operated from close inshore to about 16 m depth. Generally, the purse seiners also operate within the same depth range; but, occasionally the operations extend up to 20 m depth. This paper embodies the results of a comparative study of the operation of the purse seine and thangu vala for the exploitation of the oil sardine along with studies on such biological aspects as the size, sex composition, etc. MATERIAL AND METHODS Regular oil sardine samples, of at least 50 fish each, were collected from the purse seine catches on every fishing day at Cochin from 1969. length, mean length, body weight, stages of maturation and sex composition were noted. Catch details such as depth of fishing, duration of operation of the gear and number of hauls were collected from the log of the fishing vessels. 'Present address: Central Marine Fisheries Research Institute, Substation. Kozhikode-S.

PURSE SEINE FISHERY FCM S. THE FISHERY LONGICEPS The purse seine employed aboard the 36 foot vessels (M2/M3) measures 320 m in length and 20-28'm in width. It is made of nylon knotless webbing and is provided with a thick twined bunt at one end. The head rope and foot rope made of terylene measured 320 m and 330 m respectively in length and the purse line was about 500 m in length. Details of the mode of fishing and dimensions of this net are given by Menon (). The design and operation of thangu vala are described by Kuriyan, George and Menon (1962). 71 TABLE 1. Oil sardine catches of Purse seine (Jndo-Norwegian Project) from 1968 to 1971, off Cochin. Months Oil sardine (kg.) Catch per haul (kg.) Catch per hour (kg.) September Oct. Nov. Dec. January Feb. 1968 f>, >» 1969» 864.0 26737.0 10099.0 16186.0 22179.0 2303.0 432.0 1273.2 561.1 899.2 1056.1 1919 216.00 656.12 294.17 553.36 603.51 112.34 78368.0 Monthly average 851.8 Monthly average 473.29 October November December January February March April 1969 11 >» fl t»»»» 25978.0 31618.0 54.0 25743.0 30174.6 24202J) 20965.0 3790.0 1731.9 1129.2 543.9 1430.2 2.3 968.1 1397.7 758.0 731.77 715.14 362.60 725.15 815.51 562.83 742.12,, 1 505.33 182524.0 1267.5 659.53 October November December January February March April *f 1* 1971 >l l> J» fi 15788.0 24953.0 15236.0 5500.0 14554,0 21966.0 12091.0 7332.0 1754.? 1134.2 896.2, 785.7 519.8 523.0 287.9 1466.4 853.40 :, 594.11 534-59 323,52 312.98 327.85 177.is 733.20 117420.0 682,7 394.36 At the deepest region in the middle of the seine, it is 28 m wide-

72 V. BALAN Traditionally, the sardine fishery by thangu vala is from August to April. Either due to the poor availability of the oil sardine shoals in the usual fishing areas or the inclement weather affecting the fishing operations, due to the catches are usually poor between and September. The Indo-Norwegian Project purse seines were generally operated from October to, off Cochin. The details of the purse seine catch, catch per haul and catch per hour are furnished in Table 1; those pertaining to thangu vala for the same period are furnished in Table 2. For comparison of effort, the catch per haul of the purse seine and catch per unit of thangu vala have been used. The sardine catches In the 1968-69 season, the purse seine fishery began in September and ended in February (Table 1). The monthly catch per haul varied front 1919. to 1273.2 kg, the highest (127i3.2 kg) being in October. The monthly catcli per hour was on an average about 500 kg. ' Mfhe 1969-70 season, the Ihsery began in^ October with good landings. The highest annual catch per effort was obtained dunng this season; the catches remained good throughout the season excepting in is/[$^ The monthly catch per haul ranged from 543.9 to 2.3 kg, the highest )b!eing in February; monthly catch per hour also was the highest during thai Baonth. In -71 season, the fishery was moderate. The heaviest catch j^vas made in November, the highest catch per haul (1754.2 kg) and the highest hatch per hour (853.4 kg.) being in October. The catch and catc}i per unit of oil sardine in thangu vala os Cochin were very poor between ana August during the three seasons studied (Table 2). TABLE 2. Monthly catch and effort (kg) of oil sardine o/thangu vala from 1968-69 to -71, off Cochin. Months 1968-69 season No. of Catch per catch units unit June July Aug. Sept. Oct. Nov. Dec. Jan. Feb. Mar. Apr. 22,468 138,355 2,429,373 343,497 190,274 191,300 235,119 144,829 37,078 85,277 45,333 1591,332 679 1516 1412 668 719 652 675 605 540 206 325 314 8,311 330 91.3 1.7 ^ 232.6 477.7 291.8 2834 388.6 268.2 180.0 262.4 144.4 191.5 1969-70 season No. of Catch per catch units unit 51,705 140.342 13,437 141,678 333,228 432,421 30,903 63,4 2 217,399 171,203 4,250 8,550 1608,528 954 1020 957 646 1053 1052 419 380 714 733 128 192 8,248 54.2 137.6 14.0 219.3 316.5 411.0 73.8 166.9 304 5 233.6 33.2 44-5 195.0-71 season No. of Catch per catch units unit - 7,340 74,154 239,963 276,395 225,340 657,570 642.660 990,379 389,096 181,734 101,215 139,789 3925,635 744 1,188 1,364 1,099 1,070 1,210 1,528 1,348 1.250 1,200 1,271 1,038 14,310 9.9 62.4 175.9 251.5 210.6 543.4 520.6 734.7 311.3 151.4 79.6 134.7 274.3

PURSE SEINE FISHERY FOR 5. LONGICEPS 73 During 1968-69, the catch per unit ranged from 1.7 to 477.7 kg. In 1969-70, catch per unit was fairly good from September to March excepting in December and January. In -71, it was remarkably good from September to February made in and yielded the highest figure for catch per unit (734.7 kg), in January. The catch per haul of purse seine and catch per unit of thangu vala were fairly good from October to January in 1968-69. Though the catches per haul of purse seine were remarkably good throughout 1969-70, they were below average in thangu vala during that season excepting from October-November and February. In -71, though the catch per haul figures in purse seine were average throughout, the corresponding figures in thangu vala were just above average for about five months only with a sharp decline from March to. I Ml ISO 8 I ;^ 49 19 loll 15 26 22 ^,.,.n?c23 18 j 12931 26?5 nil.. ' > 2 5 4 6 19 12 123 3192022 lu JAN 1971 FEB 1971 MA ft ^71 APR 97f MAY I97t MODAL LENGTH MEAN LENGTH 28 24 25 20 : zs NOV FEB MAR APR 1969 20 17 23 NOV 27 Ml M 11.. ' I.. 10 le 17 OEC 21221 31 FiO' 1. Length distributions of oil sardine samples from daily total catches of purse seine (dates marked against length ranges) from November 1969 to 1971, off Coctiin-

74 V. BALAN LENGTH COMPOSITION OF THE OIL SARDINE CATCHES OF PURSE SEINE As the purse seine is able to catch almost the entire shoals encountered, it is reasonable to infer that the catches obtained in this net give a complete picture of the length composition of the individuals constituting the shoals. The modal and the mean length of the sardine samples (taken from daily total catches) examined on different fishing days show good similarity (except in March ). (Fig. 1). In November 1969, with the appearance of older age group in the catches, the mm modal length dominated. In February and April, with the increase of abundance of young age groups, the mode dropped to mm (the 170 mm mode was relatively less important in February). In March, a higher year class (170 mm mode) was dominant, the mm mode being less important. When the purse seining started again in November, on account of adundant recruitment of older age group consisting mostly of" spent" sardine, the 160 mm modal length (mode being the same in the succeeding month) became prominent. Mode progressed slightly, then remained static in January and February of 1971, owing to a large-scale influx of older year-class of" spent-recovering" fish. From March to, modal length slightly declined and stood static at mm. The mean TABLE 3. Monthly length distribution of oil-sardine in Purse seine and thangu vala catches from November 1969 to 1971 off Cochin Months Nov. Dec. Jan. Feb. Mar. Apr. June July Aug. Sept. Oct. Nov. Dec. Jan. Feb. Mar. Apr. Purse seine length Mean length range (itim) (mm) 1969 150-175 168.3 N. A. do 115-180 146.5 110-180 153.7 110 180 144 9 N. A. N. F. N.F. N.F. N.F. N. A. 115-190 160.7 115 195 162.1 1971 125 185 161.2 125-195 157.6 115 195 160.8 125-195 156 4 135-195 161.1 N..A.- Material not available. N. F.--No fishing. Modal length (ram) 170 133 163 160 thangu vala i length Mean length range (mm) (mm) 110-180 100-185 110 185 UP 180 115 180 120-170 115 170 110-175 -180 45 195 75 185 120 195 115 195 115 195 115-195 120-190 120-190 ; 120-190 125 195 159.6 129 8 135.3 139.0 146.8 136.0 1322 142.8 150.1 115.1 116.2.4 158.5 160.7 161.0 158,5 159.2.3 160.2 Modal length (mm> 120 120 145 145 1(0 140 150 160 150

PURSE SEINE FISHERY FOR S. LONGICEPS 75 length values of the sardine catches of purse s«4ne and thangu vala showed a fairly good similarity from November to 1971. The modal lengths of the oil sardine caught in the purse seine during the period of this study showed a fair degree of similarity with those of the fish landed by thangu vala excepting in March and D^ember and March 1971, the slight differences observed in certain months being possibly due to commingling of slightly different age-groups in thefishinggrounds (Table 3). SEX COMPOSITION A total of 3,754 oil sardine was"examined for studying the sex composition during the period of this investigation. There were small variations in the percentages between the two sexes in theiiidiyidii%l san^les. TJje average proportion p of females for the entire period of study tvas found to be 0.i5013, i. e. about half. To test if this average proportion was being, maintained during the different months, of the period, Chi-square test was made xising the formula J p q i M \,,^ where p is proportion of females to the'totftl^fpr all th6 months, qis (1-p), a is the individual number of females in each month%tid n, fs theto^al number of females examined during this investigation. The X' value'was found to be 12.66 which is non-significant, showing that there is, no significant difference in the sex ratios between months (Table 4). a ^ * 1 " DISCUSSION ' ' ; A comparison of the sardine cttches made by the purse seine and thangu vala in the inshore waters off Cochin indicated slight differences during^he three seasons covered. In addition to differences in the number of hauls made, in hours spent in searching for shoals, in the availability of shoals in the fishing grounds etc., in the case of purse seine, factors sucli as (1) total area of webbing of the net, (2) total area covered during each operation of the net, (3) horse power and size of the mechanized vessel used and (4) the efficient manoeuvrability in shooting and hauling up of the net are also to l)e taken into consideration while comparing it with the indigenous gear. In the case of thangu vala since several hauls were made and several units operated on different fishing days, and all the hauls made by each unit were pooled together in the respective units before the catches were landed on each fishing day. Heijce, a, comparison of catch per haul of purse seine with the catch per unit of tharig^ yah would obviously have its own limitations.! j ' Homell and Nayudu (1923);vi-fco*ded dom&ance of females up to the size at first maturity, the sexual segre atio4 gettihg reduced among the ripe fish, and suggested that there was gres ev. ii of-talifyj amo g females after spawning.

0\ TABLE 4. Monthly sex ratio of oil-sardine in purse seine catches during I969-'71 off Cochin Femaies Maks Ratio of Females Nov 1969 38 38 76 OSOOO Feb 77 100 177 04350 Mar 9y 103 100 203 05074 Apr 99 82 82 164 05000 Nov 9> 189 206 395 04785 Dec 99 250 219 469 05330 Jan 1971 93 80 173 05376 Feb 244 243 487 05010 Mar 305 262 567 05379 Apr 373 390 763 04889 >> 128 152 280 04571 Examined 1882 1872 3754 0-5013 > t-l >

PURSE SEINE FISHERY FOR s. LONGICEPS 77 Chidambaram (1950) reported eiqual proportions of sexes upto the length of 200 mm, but beyond that size, the females were found to outnumber the males. Nair (1960) found no disparity in sex ratio. Antony Raja (1969, p. 54) stated, " although the individual samples showed dominance of either sex or their equal distribution, there were no seasonal differences in the ratio of males to females, nor dominance of either sex among juvenile stock. But the females were distinctly more in the overall population of recovering spawners." A statistical examination of the available sex composition data in the present study clearly established that there was no significant predominance of either sex in the oil sardine catches made by the purse seine, during the seasons 1969-71 off Cochin. ACKNOWLEDGEMENTS I am grateful to Dr. S. Z. Qasim, the Director, and Shri S. K. Banerji of C. M. F. R. Institute, Cochin, for suggesting improvements in the typescript. My thanks are also due to Shri S. K. Dharmaraja for his help in application of Chi-square test. I also gratefully acknowledge receipt of the catch data of purse seiners from the Indo-Norwegian Project, Cochin. REFERENCES ANTONY RAJA, B. T. 1969. The Indian oil sardine. Bull. 16, Cent. mar. Fish. Res. Inst., Mandapam Camp, November 1969. BALAN, V. 1971. The fecundity.and sex composition of Sardinella longiceps Val. along the Cochin Coast. Indian J. Fish, 12 A (2): 473-491 (October 1965). CHIDAMBARAM, K. 19S0. Studies on the length frequency of the sitdine, Sardinella longiceps Cuv. and Val.. and on certain factors influencing their appearance on the Calicut coast of Madras Presidency. Proc. Indian Acad. Sci., 31 (5): 252-286.'^ HORNEUU, J. AND M. R. NAYUDU. 1923. A contribution to the life history of the Indian sardine with notes on the plankton of the Malabar coast. Madras Fish. Bull. 17 (5): 129-197. KuRiYAN, G. K., V. C. GEORGE, AND T. R. MENON. 1962. Design and operation of the so-called " Thangu vala " a single boat seine. I. P. F. C- Occasional paper 63/9 ; 1-17. MENON, M. D.. PurKS seining oil sardines from medium vessels. Souvenir- Central Inst- Fish. Operatives. Pp. 2S-i\. (February ). NAIR, R. V I960. Synopsis on the biology and fishery of the Indian sardine- Proc- World Scl- Meeting on the Biology of Sardines and related species, 2- Species synopsis (11), Fisheries Biology synopsis 16:329-414, FAO-, Rome-