Report of the Lake Erie Yellow Perch Task Group

Report of the Lake Erie Yellow Perch Task Group March 2 Members: Megan Belore Andy Cook, (Co-chairman) Don Einhouse, (Co-chairman) Travis Hartman Kevin Kayle Roger Kenyon Carey Knight Brian Locke Phil Ryan Bob Sutherland Mike Thomas Elizabeth Wright Ontario Ministry of Natural Resources Ontario Ministry of Natural Resources New York Department of Environmental Conservation Ohio Department of Natural Resources Ohio Department of Natural Resources Pennsylvania Fish and Boat Commission Ohio Department of Natural Resources Ontario Ministry of Natural Resources Ontario Ministry of Natural Resources Ontario Ministry of Natural Resources Michigan Department of Natural Resources Ontario Ministry of Natural Resources Presented to: Standing Technical Committee Lake Erie Committee Great Lakes Fishery Commission

Table of Contents Introduction... 2 Charge 1: 2 Fisheries Review and Population Dynamics... 3 Age Composition and Growth... ADMB Catch-Age Analysis and Population Estimates... Recruitment Estimator for Incoming Age 2 Yellow Perch... 6 2 Population Size Projections... 7 Charge 2: Harvest Strategy and RAH... 8 Stock Recruitment Simulation... 8 Stock Recruitment Simulation Sensitivity Analyses... 9 Harvest Strategies and RAH Determination...1 Charge 3: Yellow Perch Genetics........11 Charge : Eastern Basin (MU ) Sub-stock Delineation and Boundaries...11 Suggested New Charges for 2-26...12 Acknowledgments...12 Literature Cited...13 Note: The data and management summaries contained in this report are provisional. Every effort has been made to insure their correctness. Contact individual agencies for complete state and provincial data. Data reported in pounds for years prior to 1996 have been converted from metric tonnes. Please contact the Yellow Perch Task Group or individual agencies before using or citing data published herein.

Introduction From April 2 through March 2, the Yellow Perch Task Group (YPTG) addressed the following charges: 1) Maintain centralized time series of data sets required for population models including: a) fishery harvest, effort, age composition and biological parameters b) survey indices of adult abundance, size at age, and biological parameters c) recruitment indices and biological parameters of juvenile yellow perch 2) Support a sustainable harvest policy by: a) examining exploitation strategies b) recommending an allowable harvest (RAH) for 2 in each management unit c) supporting decision/risk analysis strategies for yellow perch management 3) Contribute to lake-wide genetic research on Lake Erie yellow perch stocks. ) Examine the issues of Eastern Basin (MU) sub-populations and explore whether there is support for re-defining boundaries within MU to manage as separate stocks. During the Coordinated Percid Strategy (CPMS) (21-23) the yellow perch task group had an independent review conducted by Myers and Bence (21). AD Model Builder (ADMB) software was adopted for catch-age analysis and was considered a significant improvement over the former CAGEAN approach. This new programming tool offered greater flexibility, allowing integration of survey data in the model which formerly relied exclusively on fishery data. In addition, commercial gill net selectivity became size dependent instead of simply age dependent. The task group explored a number of exploitation models including the Beverton- Holt yield per recruit, spawning stock biomass (Fx% SSB), Thompson-Bell F.1, spawner biomass per recruit and simulation based approaches. The task group, with endorsement from the LEC, concluded that a simulation based approach provided the most meaningful reference points on which to base harvest strategies. Drawbacks to this approach relate to assumptions and uncertainties common to most, if not all, fisheries models. Sensitivity analyses conducted in 2-2 quantified the implications of some simulation assumptions and are discussed in this report. New charges recommended for 2-26 are expected to address catch-age analysis uncertainties. A formal decision analysis for Lake Erie yellow perch is presently under consideration. Development of a Yellow Perch Management Plan (YPMP) is scheduled to take place this year. More details regarding charges including the most recent status of Lake Erie yellow perch stocks are described herein. 2

Charge 1: 2 Fisheries Review and Population Dynamics The lake-wide total allowable catch (TAC) in 2 was 11.27 million pounds. This allocation represented an 11% increase from a TAC of 9.96 million pounds in 23. For yellow perch assessment and allocation, Lake Erie is partitioned into four Management Units (Units, or MUs; Figure 1.1). The 2 allocation by management unit was 3.89,.37, 2.9 and.26 million pounds for Units 1 to, respectively. The lake-wide harvest of yellow perch in 2 was 9.739 million pounds, the highest observed since 199 (9.6 million lbs). The 2 harvest was % higher than reported in 23. Harvest by management unit was 2.9,.3, 2. and.2 million pounds for units 1 to respectively (Table 1.1). Harvest was near or below TAC in all management units, except in Management Unit 1, where the 2 harvest was about one million pounds below the TAC. The distribution of harvest among jurisdictions in 2 was similar to 23 lake-wide, but differed more within management units (Table 1.1, Figure 1.2). Harvest, fishing effort, and catch rates are summarized for the time period 199-2 by management unit, year, agency, and gear type in Tables 1.2 to 1.. Trends over a longer time series (197-2) are depicted graphically for harvest (Figure 1.2), fishing effort (Figure 1.), and catch rates (Figure 1.8) by management unit and gear type. The spatial distributions in 2 of harvest (all gears), and effort by gear are presented in Figures 1.3 and 1. to 1.7 respectively. Lake-wide, yield in 2 was almost identical to 23 for Ohio, but increased in Ontario (.2%), Michigan (12%), Pennsylvania (3%), and New York (232%). Compared to 23, harvest totals in 2 increased by 8% in MU 1, 2% in MU 2, 1% in MU 3 and 2% in MU. Ontario s 2 harvest increased in MU 1 (%) and MU (17%), but declined in MU 2 (3%), and MU 3 (13%). Michigan s 2 harvest (Unit 1) increased 12% from 23. In Ohio waters, harvest decreased in Unit 1 (22%), but increased in Units 2 (6%) and Unit 3 (37%). Pennsylvania s harvest increased in Unit 3 (38%) and Unit (16%). New York s 2 harvest (MU ) more than tripled from the previous year. Harvest from commercial trap nets increased in Units 1- (1%, 3%, 3%, and 273% respectively). Trap net effort (lifts) for 2 increased in Unit 1 (97%), Unit 2 (18%) but decreased in Unit 3 (2%) and Unit (2%). Within management units, Ontario s yellow perch harvest from large mesh gill nets (3 inch or greater) in 2 ranged from less than 1% to 8% of the gill net harvest. Harvest, effort and catch per unit effort from a) standard yellow perch effort (<3 inch stretched mesh) and b) 3

larger mesh sizes, are distinguished in Tables 1.2 to 1.. Targeted gill net effort increased in MU 1 (62%) but decreased in MU 2 (7%), MU 3 (19%) and MU (%) compared to 23. Gill net effort remained generally low in 2 compared to the 199 s and earlier decades (Figure 1.). In 2, sport harvest decreased in MU 1 (28%) but increased by 12%, 37% and 16% in Units 2, 3 and, respectively. Angling effort decreased in MU 1 (2%) but increased in MU 2 (%), MU 3 (38%) and MU (17%). Due to the larger size (older age) composition of the sport harvest, angling catch rates expressed as kg harvested /angler hour (Figure 1.8) increased, in contrast to the number of fish harvested /angler hour which decreased in MU 1 (Table 1.2). Both gill net (12%) and trap net (1%) fisheries experienced lower catch rates in 2 compared to 23 in Unit 1. In MU 2, catch rates in 2 were similar to 23 for all gears with trap net values down 13%, sport success up by 12% and gill net harvest rates virtually unchanged (Table 1.3). In MU 3, sport catch rates contrasted between Ohio and Pennsylvania anglers, with increased fishing success in Ohio (39%), countered by a reduction of 26% in Pennsylvania waters (Table 1.). Commercial success rates improved in MU 3 for both gill net (8%) and trap net (91%) fisheries. With the exception of the Pennsylvania s sport fishing success down marginally in 2, catch rates improved dramatically in MU ; in both New York (sport by % and trap net by 8 fold) and Ontario s waters (gill net fishery by 23%). Ontario uses an ice allowance policy, first implemented in 22, by which 3.3% was subtracted from commercial landed weight. This step was taken so that ice was not debited from fishers quotas. Ontario s landed weights in the YPTG report have not been adjusted to account for ice content. Ontario s reported yellow perch harvest is represented exclusively by the commercial gill net fishery, described above. Reported sport harvests for Michigan, Ohio, Pennsylvania and New York are based on creel survey estimates, however, the sport harvest of yellow perch from Ontario waters is not routinely assessed. Additional fishery documentation is available in annual agency reports. Age Composition and Growth The yellow perch harvest in 2 consisted mostly of the 21 (age 3) and 1999 (age ) year classes, with older fish (i.e.: 1998 year class and earlier) more common in the catches of trapnet and sport fisheries farther east (MU 3 and MU ) (Table 1.6). There was limited, variable contribution from the 2 year class (age ) across management units. The harvest

of age 2 recruits (22 year class) was negligible in all Units. Ontario gill net harvest age composition is from targeted yellow perch harvest only. Differences between the age composition of the harvest between areas and gear types reflect different growth rates, factors affecting age interpretation, gear selectivity, and levels of abundance affected by recruitment and survival. Yellow perch growth trends differ among life stages and between basins (Figure 1.9). An abundance of yellow perch growth data exists among Lake Erie agencies. For simplicity, Figure 1.9 is comprised of young-of-the-year data from summer and fall interagency trawls, while age 1 and older data are from Ontario Partnership gill net surveys (MUs 1 and ) and Ohio fall trawls (MUs 2 and 3). Size at age time series results describe stable or improving length at age for ages - in management units 2, 3 and. Growth in management unit 1 appears to be stable or decreasing among ages. In the west basin, young-of-the-year yellow perch have been smaller than average for the last three consecutive years: that included two weak and one strong year classes. A general decline in size of YOY was evident since 199 in the west basin, with recent sizes comparable to the late 198s (not shown). Factors such as temperature, lake productivity, and invasive species, along with the proliferation of yellow and white perch may have contributed to the reduced size of young-of-the-year yellow perch in the west basin. The task group continues to update yellow perch growth data in: (1) weight-at-age values recorded annually in the harvest and (2) length and weight-at-age values taken from interagency trawl and gill net surveys. These values are applied in the calculation of population biomass and the forecasting of harvest in the approaching year. ADMB Catch-Age Analysis 2 Population size for each management unit was estimated by catch-at-age analysis using AD Model Builder, with the Commercial Selectivity Index (CSI) version, updated with 2 data. The approach was unchanged from last several years methodology and has been described in a previous Yellow Perch Task Group Report (22). Estimates of population size, biomass and parameters such as survival and exploitation rates are presented for 199-2 in Table 1.7 and for 197-2 in Figures 1.1 1.13. Mean weight-at-age from biological surveys was applied to abundance estimates to generate population biomass estimates (Table 1.8 and Figure 1.11). Population estimates are critical to monitoring the status of stocks and determining allowable harvest. Abundance estimates should be interpreted with several caveats. Inclusion of abundance estimates from 197 to 2 implies that the time series are continuous. Lack of

data continuity weakens the validity of this assumption. Survey data are represented in the latter part of the time series (generally 1989 to present), while methods of fishery data collection have also varied. Model parameters, constrained to constants, such as natural mortality, catchability and selectivity blocks, lessen our ability to directly compare abundance levels over three decades. In addition, commercial gill net selectivity was estimated independently in the latter part of the time series using gill net selectivity curves derived from index gillnet data by the method of Helser (1998), involving back calculation of length-at-age and weightings based on the monthly distribution of harvest-at-age. With catch-age analysis, the most recent year s data estimates inherently have wide error bounds. This is to be expected for cohorts that remain at-large in the population. Population estimates are derived by minimizing an objective function weighted by data sources including fishery effort, catch and survey catch rates. The weightings (or lambdas) of effort data are calculated by the ratio of variance of observed log-catch to log-effort (Quinn and Deriso, 1999). Weightings of fishery catch and survey catch rates are solved iteratively until convergence occurs; until lambdas remain relatively constant. While lambdas within similar parameter groups (i.e.: effort, catch and surveys) are solved and weighted unequally, the groups themselves are given equal weight. This can be problematic, as in the case of MU 1 for which fishery catch lambdas failed to converge. In order to address this lambda calculation process fully, a new charge has been recommended for 2-26 to review and examine methods of deriving lambdas. Data weightings are presented in Appendix Table 1. Plots of fishery and survey data residuals from catch-age analysis are presented in the Appendix Figures 1. Recruitment Estimator for Incoming Age 2 Yellow Perch Age 2 recruitment in 2 was predicted by linear regression of juvenile yellow perch trawl indices against catch-age analysis estimates of two-year-old abundance. Age 2 recruitment in 2 was calculated using the mean of values predicted from the indices listed in Appendix Table 2. Data from trawl index series for the time period examined are presented in Appendix Table 3 (geometric means) and Appendix Table (arithmetic means), while a key that summarizes abbreviations used for the trawl series is presented as a legend in the Appendix. The estimates of age 2 recruitment for 2 (the 23 year class) was strong in all management units (Table 1.7, Appendix Table 2). The 23 year class should contribute to fisheries in 2 to varying degrees among MUs, based on selectivity of the age 2 s in each MU. 6

They will contribute even more so in 26, as they become larger and more vulnerable to all harvest methods. 2 Population Size Projection Stock size estimates for 2 (ages 3 and older) were projected from catch-age analysis estimates of 2 population size and age-specific survival rates in 2 (Table 1.8). Projected age 2 recruitment from the 23 year class (method described above) was added to the 2 population estimate for older fish in each unit, producing the total standing stock in 2 (Table 1.8). Standard errors and ranges for estimates are provided for each age in 2, and following estimated survival (from ADMB), for 2. Descriptions of min, mean, and max population estimates refer to the estimates minus or plus one age-specific standard error. Stock size estimates projected for 2 were among the highest of the time series due to the 23 year class (Table 1.7 and Figure 1.1). Overall, projected 2 yellow perch abundance (2+) is 1%, 117%, 6% and 1% greater than 2 in management units 1 to, respectively. Estimates of abundance for age 3 and older yellow perch in 2, however, were not as favorable by comparison. Abundance of perch ages 3 and older in 2 was projected to be reduced by % or more than estimated for 2. As a function of population estimates and mean weight-at-age, biomass estimates in 2 were among the highest in the time series (Figure 1.11). Total biomass estimates for 2 increased from 2, except in MU where biomass was comparable between 2 and 2. Yellow perch biomass estimates for 2 (ages 2 and older) increased %, 9% and 6% in MU s 1 to 3, respectively, but decreased slightly by 7% in MU. Biomass of ages 3 and older yellow perch decreased significantly- by 3%, %, 32% and 22% in Units 1 to, respectively. Estimated survival of yellow perch ages 2 and older in 23 were 8%, 38%, 6% and 62% in MU 1, 2, 3 and, respectively (Figure 1.12). In 2, estimated survival was 8%, 38%, % and 63% in Units 1 through. As expected, survival rates were higher for fish ages 2 and older, than ages 3 and older, since new recruits are less vulnerable to fishing mortality. Albeit with fluctuations, estimated survival has improved gradually in all management units since early to mid 199s. Estimated exploitation rates in 23 were 23%, 37%, 1% and 6% in Management Units 1, respectively, for ages 3 and older. Exploitation rates for 2 were estimated at 23%, 37%, 1% and % for yellow perch ages 3 and older (Figure 1.13). Exploitation rates of 7

yellow perch ages 2 and older are lower since new recruits are less vulnerable to fishing. Charge 2: Harvest Strategy and RAH Harvest Strategy Methodology In 2, a suite of exploitation strategies was presented to the Lake Erie Committee (LEC) to support TAC determination. This year, the LEC directed the YPTG to present a single yield strategy, the F.1 spawner-recruit (S/R) for management units 1, 2 and 3. The F.1 rates calculated in 2 remain unchanged this year for MUs 1-3 (Tables 2.1.1-2.1.3, 2.2.1). F.1 values were derived based on the ratio of average yield to average recruitment plotted against fishing rates. F.1 was determined from the fishing rate at which the slope was 1% of the initial slope of the curve. This approach does not rely on the assumption of knife-edge recruitment, and it incorporates a gamma stock-recruitment relationship. Parameters include mean weightat-age from harvest (recent two-year mean), age specific selectivities (recent two-year mean) from catch-age analysis weighted by sharing formula, and survey maturity data. The simulation assumes that the targeted fishing rates will be realized for all gear types. Simulation methodology and risk assessment is described below. Stock-Recruitment Simulation This simulation approach documented in 2 remains essentially the same this year, although a number of sensitivity analyses were conducted. Spawner-recruit (S/R) relationships were described by gamma functions (Reish et al. 198 in Quinn et al. 1999) with the recognition that environmental factors exert major influence on recruitment. The YPTG created population simulations based on gamma stock recruitment functions, influenced by environmental factors. Environment Factors (EF) were derived from residuals of the S/R relationship as: EF = (observed recruitment)/(predicted recruitment) Two years of recent abundance estimates were used to initiate simulations. Recruitment for each year was estimated from the S/R function, and then multiplied by an EF selected randomly from the observed distribution of residuals (EFs). This process extended over 2 years and 1 replicates under a broad range of fishing mortality rates ( to 2) to produce measures of risk. Other model parameters included were consistent with ADMB catch-at-age 8

analysis. This process, applied to populations in each management unit, allowed the YPTG to quantify risk associated with various fishing rates, while giving consideration to stockrecruitment patterns and environmental influences experienced by yellow perch during recent decades in Lake Erie. Biological reference points including spawner biomass (as a fraction of an unfished population), survival rates, and the probability of attaining low levels of abundance comparable to 1993-9 were included as outputs. A further refinement in 2 included averaging the results of simulations over 1 multiple runs. This is described in detail under Sensitivity Analyses below. Results are presented for each management unit in Tables 2.1.1-2.1.3. Stock-Recruitment Simulation Sensitivity Analyses In order to address concerns from the LEC regarding the sensitivity of risk indicators to F.1 S/R model assumptions, the YPTG conducted sensitivity analyses relating to 1) model configuration, 2) time frames used in model (that relate to recruitment potential), 3) stock recruitment model applied to the data, and ) single vs. multiple simulations to describe average outcomes at different fishing rates. Simulation results using data input from two different catch at age models were compared. The first inputs used catch-at-age analyses results that solved commercial gill net selectivity, and the second inputs used results from the current catch at age model in which selectivity was calculated independently of catch-age analysis. Using MU 1 as an example, the F.1 rate was higher with selectivity calculated independently, but would have produced a lower TAC in 2 (21%) due to lower population estimates. Time frames compared included the entire series (197-23) vs. one that began when target phosphorus loading targets were achieve circa 1982, with the belief that conditions were more favorable for recruitment prior to this period. The F.1 rate derived from the shorter (stock-recruit) time series (1982-23) was lower compared to that derived using the full time series (197-23) in MU1, but was higher in MU2. Risk indicators were more apparent at lower fishing rates in the 1982-23 model. Gamma, Ricker, and Beverton-Holt stock recruitment relationships were used to calculate F.1 rates with time series of various lengths and risk indicators. The S/R model that triggered risk indicators at the lowest fishing rates was Beverton-Holt, followed by Ricker and Gamma models which alternated between MUs. Derivation of F.1 rates were insensitive to truncating the time series for Ricker or Gamma S/R models (from 1982-23 down to 1982-1998). 9

The simulation is driven in part by the environmental factors (EFs) used in the model. The arrangement of these EFs can affect the risk levels associated with different fishing rates. Therefore, it was suggested that an average of results from multiple simulations with different selections of EF's be used to describe the risk associated with each fishing rate. Although F.1 rates that were derived in 2 are presented (Tables 2.1.1 2.1.3, 2.2.1), the simulations were run again multiple times (1) and averaged to better describe risk associated with various fishing rates. In 26, following the lambda review and development of the Yellow Perch Management Plan, application of the results of the sensitivity analyses will be given further consideration. Harvest Strategies and RAH Determination A harvest strategy is a plan that should be robust to the unpredictable and/or uncontrolled biological fluctuations that are expected from the stock. A harvest strategy involves biological, economic, social and political decisions (Hilborn and Walters, 1992). The task group described biological risk associated with various fishing intensities. The YPTG calculated target fishing rates (F.1 S/R) believed to be sustainable based on a simulation approach that is subject to assumptions which overlap with those of catch-age analysis. These may be essential elements of a harvest strategy, but do not by themselves constitute a complete one until economic, social and political considerations have been satisfied. The LEC, supported by the YPTG, deemed that the F.1 harvest projections for 2, presented in Table 2.2.1, satisfied these elements in the interim, until outstanding uncertainties have been addressed and incorporated into a Yellow Perch Management Plan (YPMP). Since 2, the Management Unit harvest strategy has been pursued separately from other Units, and established more directly by the LEC as a rehabilitation strategy. Based on the improved status of yellow perch reported in Management Unit, the LEC proposes a % increase in allowable harvest for MU in 2. This proposal should not impose excessive biological risk to yellow perch or other species, and represents a compromise between diverse economic and social interests. If the proposed TAC were adopted in 2, the outcomes would be monitored by assessment programs and reported by the YPTG. Further considerations for MU are discussed in Charge. Also of note, 2 marks the end of a transition from historic pattern to lake area allocation sharing by jurisdiction which began in 1993. In 2, lake area-based allocation shares by management unit and jurisdiction are: 1

Allocation by Management Unit and Jurisdiction, 2: MU 1 : MI 8.1% OH 9.6% ONT 2.3% MU 2: OH 7.% ONT 2.% MU 3: OH 31.9% PA 11.9% ONT 6.1% MU : NY 27.6% PA 17.2% ONT.2% With the advent of geographic information software (GIS) technology, the Standing Technical Committee (STC) is redefining yellow perch management unit delineations using modern approaches. Implications to sharing formulas based on lake surface area have yet to be addressed. Charge 3: Yellow Perch Genetics During 2 the YPTG supported genetic stock discrimination research by collecting yellow perch tissue samples for Dr. Carol Stepien at the University of Toledo. In recent years this support has become an annual endeavor by the YPTG with expectation that this research will expand our understanding of yellow perch genetic stock structure. Ongoing tissue collections from spawning concentrations should assemble a database that represents a stock library for Lake Erie yellow perch. The YPTG thanks Dr. Carol Stepien and her associates for their continued efforts. Charge : Eastern Basin (MU ) Sub-stock Delineation and Boundaries Yellow perch in eastern Lake Erie have been treated as a single stock for assessment and allocation purposes since the 198s. However, MU is notable among Lake Erie s yellow perch management units as the area where yellow perch fisheries are more often spatially isolated within the basin, and yellow perch habitat remains more clearly partitioned by lake bathymetry. Also, there has been evidence of differing recruitment patterns within various parts of the basin. Finally, the Myers and Bence (21) independent review of YPTG stock assessment efforts identified MU as a special case where stock definition seemed evident within the basin. Currently, eastern basin yellow perch stock assessment is being examined as part of a thorough technical review being pursued by the Ontario Ministry of Natural Resources, Eastern Lake Erie Technical Report Draft, December 2. At present, this draft document supports the YPTG s ongoing practice of treating the east basin yellow perch resource as one 11

unit, i.e. "MU", for stock assessment purposes. Nevertheless, there remains enough evidence for sub-stocks within MU that yellow perch assessments in this area should pursue approaches capable of detecting, describing and managing discrete stocks. Suggested New Charges for 2-26 1) Lambda review- In 2-26 the YPTG & STC will initiate a review of methods that can be used to generate data set weighting lambdas for catch-age analysis. The objective of the review is to identify, describe and apply the most scientifically defensible method of generating lambdas that influence population estimation. 2) Yellow Perch Management Plan In 2-26, the LEC, the YPTG, and the STC will formulate, with stakeholder input, a yellow perch management plan that documents historic methods and outlines an appropriate exploitation strategy with measurable performance indicators. Acknowledgments The task group wishes to thank the following people for providing support to the task group during the past year: Tim Bader (Ohio Department of Natural Resources, Division of Wildlife), Mike Bur (US Geological Survey- Biological Resources Division), Bruce Morrison (Ontario Ministry of Natural Resources), Dr. Carol Stepien (University of Toldeo), Jeff Tyson (Ohio Department of Natural Resources, Division of Wildlife), and Larry Witzel (Ontario Ministry of Natural Resources) The YPTG report could not be completed without the contributions of all Lake Erie staff from the Michigan Department of Natural Resources, Ohio Division of Wildlife, Pennsylvania Fish and Boat Commission, New York Department of Environmental Conservation, US Geological Survey- Biological Resources Division, and the Ontario Ministry of Natural Resources. In addition, the YPTG expresses thanks to the Great Lakes Fishery Commission for their continued support. 12

Literature Cited Helser, T.E., J.P. Geaghan and R.E. Condrey. 1998. Estimating gill net selectivity using nonlinear response surface regression. Can. J. Fish. Aquat. Sci. : 1328-1337. Hilborn, R. and C.J. Walters. 1992. Quantitative Fisheries Stock Assessment : Choice, Dynamics and Uncertainty. Routledge, Chapman and Hall, Inc. NY. MacGregor, R.B. and L.D. Witzel. 1987. A twelve year study of the fish community in the Nanticoke Region of Long Point Bay, Lake Erie: 1971-1983 Summary Report. Lake Erie Fisheries Management Unit. Report 1987-3. 61 pp. Myers, R.A. and J.R. Bence. 21. The 21 assessment of perch in Lake Erie; a review. Presented to the Lake Erie Committee, Great Lakes Fishery Commission. Ontario Ministry of Natural Resources. 2. Eastern Lake Erie Technical Report Draft. Lake Erie Management Unit. Quinn, T.J. and R.B. Deriso. 1999. Quantitative Fish Dynamics. Oxford University Press. NY. Reish, R.L., R.B. Deriso, D. Ruppert, and R.J. Carroll. 198. An investigation into the population dynamics of Atlantic Menhaden (Brevoortia tyrannus). Can. J. Fish. Aquat. Sci. 2: 17-17. Yellow Perch Task Group (YPTG). 22. Report of the Yellow Perch Task Group to the Standing Technical Committee, Lake Erie Committee of the Great Lakes Fishery Commission. Yellow Perch Task Group (YPTG). 2. Report of the Yellow Perch Task Group to the Standing Technical Committee, Lake Erie Committee of the Great Lakes Fishery Commission. 13

Table 1.1. Lake Erie yellow perch harvest in pounds by management unit (Unit) and agency, 199-2. Ontario* Ohio Michigan Pennsylvania New York Total Catch % Catch % Catch % Catch % Catch % Catch Unit 1 199 71,1 9 3,38 36 66,1 -- -- -- -- 1,21, 199 2,79 38 78,98 7 77,17 6 -- -- -- -- 1,386,9 1996 7,167 36 1,12,716 7 13,81 7 -- -- -- -- 1,96,693 1997 1,91,8 8 1,71,2 7 111,819 -- -- -- -- 2,27,688 1998 1,17,33 2 968,82 3 132,1 6 -- -- -- -- 2,271,26 1999 1,8,1 1 98,8 11,9 -- -- -- -- 2,8,197 2 98,323 7 1,38,6 67,1 3 -- -- -- -- 2,8,983 21 813,66 91,61 1 7,91 -- -- -- -- 1,799,617 22 1,,1 1,316,3 17,6 -- -- -- -- 2,917,723 23 1,179,667 1,6,38 3 8,878 3 -- -- -- -- 2,67,93 2 1,698,761 9 1,9,669 38 9,732 3 -- -- -- -- 2,88,162 Unit 2 199 1,3,9 1,,17 -- -- -- -- -- -- 2,36,12 199 1,73,83 7 8,82 3 -- -- -- -- -- -- 1,878,66 1996 1,29,998 61 823,2 39 -- -- -- -- -- -- 2,11,23 1997 1,826,18 63 1,79,882 37 -- -- -- -- -- -- 2,96,62 1998 1,797,8 7 627,9 26 -- -- -- -- -- -- 2,2,2 1999 1,72,829 62 97,123 38 -- -- -- -- -- -- 2,6,92 2 1,8,12 6 1,169,23 -- -- -- -- -- -- 2,63,39 21 1,79,27 1 1,77,69 9 -- -- -- -- -- -- 3,1,3 22 2,19,621 2 1,986,73 8 -- -- -- -- -- --,177,31 23 2,17,639 2,113,28 -- -- -- -- -- --,22,92 2 2,1,73 8 2,26,26 2 -- -- -- -- -- --,297,737 Unit 3 199 379,26 8 39,1 -- --,12 7 -- -- 793,8 199 6,2 8 83,79 1 -- -- 3,87 -- -- 79,91 1996 12,293 72 186,69 26 -- -- 9,1 1 -- -- 78,29 1997 829,33 77 219,66 2 -- -- 23,36 2 -- -- 1,72,377 1998 811,93 73 27,993 2 -- -- 28,27 3 -- -- 1,11,23 1999 66,73 6 32,63 3 -- -- 8,92 1 -- -- 1,27,263 2 771,66 62 3,2 36 -- -- 32,613 3 -- -- 1,27,9 21 999, 6 6,811 3 -- -- 91,211 6 -- -- 1,,72 22 1,192,691 6 6,1 32 -- -- 1,821 7 -- -- 1,973,616 23 1,667,133 72 81,8 21 -- -- 177,16 8 -- -- 2,326,27 2 1,3,19 62 69,7 28 -- -- 2,63 1 -- -- 2,36,929 Unit 199 2,92 8 -- -- -- -- -- -- 1,21 16 63,13 199 33,7 8 -- -- -- -- -- -- 8,12 2 1,87 1996 3,9 82 -- -- -- -- 2,2 6,72 12 37,172 1997 36,171 87 -- -- -- -- 3,9 7 2,387 6 1,67 1998 8,7 93 -- -- -- -- 38 1 3,17 6 2,17 1999 9,82 92 -- -- -- -- 2,216 3 3,23 6,292 2 3,686 73 -- -- -- -- 1,9 22 2,8 9,9 21 3,893 6 -- -- -- -- 8,337 1 1,319 26 9,9 22 87,1 -- -- -- -- 6,93 29 26,93 17 161,37 23 8,772 6 -- -- -- -- 39,821 28 16,11 12 11,1 2 98,733 9 -- -- -- -- 6,3 23,862 27 199,939 Lakewide 199 2,3,1 1,838,97 2 66,1 1,12 1 1,21 <1,13,99 Totals 199 2,96,9 1,673,9 3 77,17 2 3,87 1 8,12 <1 3,886,67 1996 2,37,93 3 2,13,836 13,81 3 11,26 <1,72 <1,82,317 1997 3,783,8 6 2,37,71 38 111,819 2 26,9 <1 2,387 <1 6,29,73 1998 3,828,31 6 1,871,779 32 132,1 2 29,6 <1 3,17 <1,86,21 1999 3,36,7 9 2,23,36 39 11,9 2 11,11 <1 3,23 <1,697,7 2 3,271,78 2,61,13 67,1 1 3,63 1 2,8 <1 6,3,9 21 3,62,68 2 3,127,21 7,91 1 99,8 1 1,319 <1 6,9,982 22,92,98 3 3,93,387 3 17,6 2 187,72 2 26,93 <1 9,23,37 23,39,211,1,228 3 8,878 <1 217,337 2 16,11 <1 9,39,16 2,32,386 3,996,38 1 9,732 1 29,7 3,862 <1 9,738,767 * processor weight 1

Table 1.2. Catch, effort and catch per unit effort summaries for Lake Erie yellow perch fisheries in Management Unit 1 (Western Basin) by agency and gear type, 199-2. Michigan Ohio Unit 1 Sport Trap Nets Sport Small Mesh Large Mesh 199 66,1 16,37 269,1 71,1 Catch 199 77,17 18, 676,93 2,79 (pounds) 1996 13,81 2,313 92,3 7,167 1997 111,819 211,876 89,19 1,91,8 1998 132,1 18,12 78,7 1,17,33 1999 11,9 2,939 77,69 1,8,1 2 67,1 2,1 798,19 98,323 21 7,91 179,23 736,7 711,7 11,321 22 17,6 337,829 978,72 1,39,637 9,68 23 8,879 2,6 1,1,929 1,11,38 28,39 2 9,732 289,136 81,33 1,637,88 61,273 199 3 7 122 322 Catch 199 3 9 37 238 (Metric) 1996 61 91 2 319 (tonnes) 1997 1 96 39 9 1998 6 8 36 31 1999 6 91 321 7 2 3 19 362 21 32 81 33 323 6 22 67 13 617 3 23 38 11 2 22 13 2 3 131 36 73 28 199 22,7,937 69,99 11,73 Effort 199 123,616,13 98,977 11,136 (a) 1996 193,733,869 772,78 8,61 1997 192,6,8 83,93 13,7 1998 183,882,6 863,336 19,9 1999 18,71,18 91,3 12,86 2 122,7,26 96,628 6,71 21 97,761 1,18 686,937 2,167 2,12 22 19,73 2,71 9,289,6 739 23 121,638 2,213 1,182,69 3,72 39 2 26,92,31 833,69 6,2 91 199 1.1 12.6 2.2 27. -- Catch Rates 199 2.8 9.6.3 21. -- (b) 1996 3.3 18.7.9 37. -- 1997 2.8 17.2 3.7 36.1 -- 1998 3.2 1.3 3.8 27.8 -- 1999 2.1 17.6 3.3 37. -- 2 2.2 27.1 3. 66. -- 21 2.9 3. 3. 19.1 21. 22 2. 6. 3. 13.7 8.2 23 2. 1.3 3. 1.1 32. 2 1.6 3.1 3. 122.7 3.8 (a) sport effort in angler-hours; gill net effort in km; trap net effort in lifts (b) catch rates for sport in fish/hr, gill net in kg/km, trap net in kg/lift Ontario Gill Nets 1

Table 1.3. Catch, effort and catch per unit effort summaries for Lake Erie yellow perch fisheries in Management Unit 2 (western Central Basin) by agency and gear type, 199-2. Unit 2 Ohio Ontario Gill Nets Trap Nets Sport Small Mesh Large Mesh 199 3,29 7,88 1,3,9 Catch 199 27,98 6,8 1,73,83 (pounds) 1996 323,33,91 1,29,998 1997 98,9 8,937 1,826,18 1998 3,661 323,283 1,797,8 1999 389,973 8,1 1,72,829 2 6,9 6,22 1,8,12 21 9,88 81,891 1,93,7 2,71 22 1,99,971 886,79 1,892,7 298,1 23 1,2,2 88,8 2,19,617 88,22 2 1,287,77 98,17 1,893,871 17,62 199 138 336 9 Catch 199 117 28 87 (Metric) 1996 17 227 8 (tonnes) 1997 226 263 828 1998 138 17 81 1999 177 26 713 2 26 27 673 21 1 382 723 91 22 99 2 88 13 23 69 389 916 2 8 3 89 71 199 7,139 38,977 23,1 Effort 199 6,67 388,238 18,337 (a) 1996,83 316,736 1,72 1997 8,721 7,36 2,97 1998 7,93 22,176 23,823 1999 7,2 63,819 13,179 2,272 61,712 6,266 21,77 81,118 3,,97 22 7,67 68,799,786 3,29 23 1,21 632,813,311 1, 2 12,23 69,,929 2,787 199 19.3 3.3 2.2 -- Catch Rates 199 18.1 3. 26.6 -- (b) 1996 2.1.2.1 -- 1997 2.9 2.8 33.2 -- 1998 17. 2.6 3.2 -- 1999 23.6 3..1 -- 2 8.6 2.9 17. -- 21 86. 3.2 29.9 18.3 22 6. 3.1 179.3 2.1 23.7 3.3 172. 2.7 2 8.6 3.7 17.3 2.6 (a) sport effort in angler-hours; gill net effort in km; trap net effort in lifts (b) catch rates for sport in fish/hr, gill net in kg/km, trap net in kg/lift 16

Table 1.. Catch, effort and catch per unit effort summaries for Lake Erie yellow perch fisheries in Management Unit 3 (eastern Central Basin) by agency and gear type, 199-2. Ohio Ontario Gill Nets Pennsylvania Trap Nets Sport Small Mesh Large Mesh Gill Nets Trap Nets Sport 199 11,12 218,29 379,26,12 Catch 199 63,9 19,8 6,2 3,87 (pounds) 1996 13,1 83,281 12,293,292 3,79 1997,776 16,888 829,33 7,398 1,962 1998 9,82 18,911 811,93,291 23,236 1999 16,28 26,377 66,73 2,9 6,2 2 16,1 286,7 771,66,93 26,683 21,72 6,339 98,622,828 2,62 96,96 22 6,1 1,9,89 97,797 2,9 138,812 23 81,9 1,67,7 2,86, 172,67 2 69,7 1,3,31 1,1 7,73 236,31 199 6 99 172 2 Catch 199 29 9 211 1 (Metric) 1996 7 38 232 2. 1.7 (tonnes) 1997 2 7 376 3. 7.2 1998 1 8 368 2. 11 1999 8 112 32 1.3 2.7 2 71 13 3 2.7 12 21 2. 29 3 23 1.2 22 29 97.9 63 23 218 77 9.1 2.3 78 2 299 6.6 3. 17 199 7,376 31,881 12,72 1,978 1991,16,67 12,27 2,18 1992 3,361 8, 1, 1,321 1993 2,61 96,619 1,17 62 199 3,3 173,76 8,169 1,2 Effort 199 3,28 2,23 6,83 1,6 (a) 1996 2,73 69,887 6,18 18 12,8 1997 2, 126,3 9,23 1 3,377 1998 2,12 111,2 1,89 3 3,612 1999 2,388 176,63,338 23 28,8 2 1,6 21,82 2,32 231 8,61 21 32 27,217 2,1 1,7 17 9,21 22 16,3 2,9 1, 9 123,287 23 26,89,617 316 87 138,72 2 368,37 3,7 268 7 17,96 199 21. 2.3 21.1 -- 17.3 -- -- Catch Rates 199 8.9 1.3 3.8 -- 9.6 -- -- (b) 1996 17.2 2.8 37. -- -- 13..8 1997 1.1 3.1 39.9 -- -- 7.6.9 1998 16.3 3.6 3. -- -- 7.9 1. 1999 2.2 3. 69.6 -- --. 1.3 2 3.3 3. 19. -- -- 11.6 1.9 21 63. 2.9 17. 22. -- 6.7 2.6 22 -- 2.7 199.6 1.7 -- 9.6 3.6 23 -- 3.1 161.8 28.8 -- 26.3.3 2 --.3 17.6 17.1 --.2 3.9 (a) sport effort in angler-hours; gill net effort in km; trap net effort in lifts (b) catch rates for sport in fish/hr, gill net in kg/km, trap net in kg/lift Unit 3 17

Table 1.. Catch, effort and catch per unit effort summaries for Lake Erie yellow perch fisheries in Management Unit (Eastern Basin) by agency and gear type, 199-2. Unit New York Ontario Gill Nets Pennsylvania Trap Nets Sport Small Mesh Large Mesh Trap Nets Sport 199,1,8 2,92 Catch 199 3,122,89 33,7 (pounds) 1996 2,822 1,6 3,9 2,2 1997 1,21 1,16 36,171 3,9 1998 1,3 1,83 8,7 38 1999 69 2, 9,82 2,216 2 62 1,833 3,686 1,9 21 27 1,292 3,28 1,68 8,337 22 1,91 2,92 8,93 1,66 29 6,87 23 1,8 1,6 8,68 12 39,822 2 3,97,9 98,716 17 9,1 199 2. 2.6 2. Catch 199 1. 2.2 1. (Metric) 1996 1.3.7 13.8 1. (tonnes) 1997.6. 16. 1. 1998.6.8 22..2 1999.3 1.2 27.1 1. 2.3.8 16.2. 21.1 6.9 1..7 3.8 22.9 11.3 39..7.1 21.3 23. 7. 38..6 18.1 2 1.8 23.1.8.1 1. 199 1,8 1,62 Effort 199 32 12,11 1,37 (a) 1996 33 6,3 1,63 7,292 1997 292 8,9 1,73 13,77 1998 178 7,73 1,81 3,78 1999 118,1 872 13,623 2 2,66 31 21,16 21 39 22,9 128 28 12,1 22 89,27 22 28 9 61,73 23 91 33,162 373 21 32,2 2 73,6 3 3.2 62,639 199 3.6. 1.6 Catch Rates 199 2.7.8 1.9 (b) 1996 2.. 13..6 1997 1.9. 1.3 1. 1998 3..7 2.3.3 1999 2.7.8 31.1. 2 6..2 1. 1.7 21.3 1.8 121. 26. 1. 22 9.9 1.3 17. 2. 1. 2. 23.2.9 12.9 2.9 1.9 2.3 1. 126.1 2. 1.7 (a) sport effort in angler-hours; gill net effort in km; trap net effort in lifts (b) catch rates for sport in fish/hr, gill net in kg/km, trap net in kg/lift 18

Table 1.6. Lake Erie 2 yellow perch harvest in numbers of fish by gear, age and management unit (Unit). Unit 1 Unit 2 Unit 3 Unit Lakewide Gear Age Number % Number % Number % Number % Number % 1..... 2,98 1. 193,366 2.9 1,7. 2,826 1.2 266,8 1.6 3 2,91,76 36.6,29,8 63. 1,,7. 11,8 2. 7,89,83 8.7 Gill Nets 1,367,16 23.9 1,8,873 1.9 318,167 9.2 3,697 1.9 2,729,8 16.8 1,26,73 26.7 893,798 13.2 1,116,92 32.2 9,73 2.8 3,97, 22.2 6+ 68,223 11.9 383,9.7 61,69 17.7,21 16.8 1,72,62 1.6 Total,721,772 6,77,62 3,71,169 2,196 16,28,99 1..... 2 19,768 2.... 19,768. 3 96,39 61. 2,397,698 6.1 6.7 9 1. 2,99,88 6.9 Trap Nets 119,81 12.2 29,2 6.8,6 71.7 97 16.7 16,23 7.9 17,382 16.1 93,866 21.9 893 11.3 1, 17.9 1,9,186 2.8 6+ 83,833 8.6 6,62 1.2 3,869 9.1 2,92. 71,278 1.1 Total 977,3,27,2 7,887,89,26,979 1 7,69.3 1,879.1.. 9,69.1 2 76,29 2.6 19,711.8,61.2 1,29 1. 11,276 1.3 3 2,3,38 68.2 1,86,3 7.2 81,9 38. 1,31 18.1,319,8.8 Sport 161,7. 13,8.9 12,37 7.2 17,13 2.2 8,87 6.3 397,231 13. 2,21 16.3 38,27 16.9 1,22 17.7 1,19,8 1. 6+ 291,987 9.9 13,73 19.8 786,79 37.2 3,83 2.3 1,627,82 21. Total 2,938,2 2,99,6 2,11,361 8,61 7,737,771 1 7,69.1 1,879... 9,69. 2 11,92 1.6 213,77 1.6 18,766.3,121 1.2 387,889 1.3 3,691,2 8.7 8,179,82 9.9 2,22,61 39.7 118,16 3.7 1,29,7 2.1 All Gear 1,68,27 17.1 1,1,977 1.6 76,36 8. 3,82 16.3 3,63,66 12. 2,81,33 21.6 2,2,68 16. 1,76,19 26. 7,6 22.9,887,777 2.2 6+ 1,6,3 11. 1,8,16 11.3 1,,97 2.1 79,18 23.9,88,823 1. Total 9,629,29 13,69,266,97,393 33,686 29,21,32 19

Table 1.7. Yellow perch stock size (millions of fish) in each Lake Erie management unit. The years 199 to 2 are estimated by ADMB catch-age analysis. The 2 population estimates use age 2 values derived from regressions of ADMB age 2 abundance against YOY and yearling trawl indices. Age 199 199 1996 1997 1998 1999 2 21 22 23 2 2 Unit 1 2 8.87 21.6 2.33 2.392 39.89 1.663 33.32 3.86 8.16 2.71 6.3 77.997 3 1.831. 13.39 1.668 12.8 2.73 6.8 21.132 22.72.2 33.639.216 1.919.82 2.71.8 7.8 6.8 13.61 3.787 12.622 12.131 2.998 17.2.31.13.232.669 1.8 2.97 2.882 6.63 2.62.289.73 1.3 6+.33.86.178.16.173.8 1.8 1.731.38 2.39 3.31 3.32 2 and Older 12.98 28.73.622 1.68 61.738.826 7.9 68.366 9.872 77.7 2.226 1.33 3 and Older.8 6.868 16.19 21.289 21.879 3.163 2.63 33.28 1.766 2.1.723 26.36 Unit 2 2 12.38 12.922 27.37 17.86 8.866 1.21. 39.279 8.96 66.138 3.97 9.7 3 3.78 7.28 7.22 13.98 9.19 3.728 8.71 29.893 22.91.1 37.898 2.2 2.879.933 2.139 2.218 3.23 3.292 1.729.6 1.9 1.33 2.16 1.98.81.66.2.9..793 1.63 7.7 2.166 6.7 3.7.768 6+.8.129.176.91.69.76.32.889.23 2.2 3.88 2.1 2 and Older 18.92 21.677 37.11 33.78 71.921 9.1 76.3 81.933 3.18 9.7 1.13 11.666 3 and Older 6.18 8.7 9.7 1.916 13. 3.89 26.2 2.6.62 2.332 7.16 2.191 Unit 3 2.818 6.21 11.98 8.66 32.933 1.36 37.77 21.8.19 23.6 1.271 31.772 3 1.11 3.37 3.92 7.6.37 21.2 6.671 23.762 13.672 2.81 1.19.81.967.73 1.969 2.292 3.97 3.177 13.29.19 1.8 8.329 1.676 8.62.19.336.381.986 1.86 2.137 1.963 7.97 2.27 8.87.61.877 6+.21.238.296.33.6.813 1.77 2.183 6.17.17 7.1 6.3 2 and Older 8.816 11.122 18.7 19.836 3.9 37.679 6.91 9.63 1.7 8.373 3.21 8.38 3 and Older 2.998.71 6.98 11.176 1.971 27.333 23.833 38.73 37.16 2.73 28.939 16.613 Unit 2.121.998.632.271 3.31 1.12 1.31 2.7 1.8 7.683.77 3.7 3.167.77.66.1.178 2.219.7 6.68 1.387 1.61.13.31.28.76..37.231.116 1.388.9.7.913.69 3.2..9.37.21.17.1.71.892.328 2.89.7.22 6+.81.37.21.26.22.11.1.1.679.611 2.16 1.2 2 and Older.2 1.198 1.39 1.18 3.919 3.73 12.39 1.281 8.36 13.128 8.977 9.18 3 and Older.321.2.78.836.6 2.93 2.36 8.212 6.82. 8..33 2

Table 1.8. Projection of the 2 Lake Erie yellow perch population. Stock size estimates are derived from ADMB and age 2 estimates for 2 are derived from regressions of ADMB age 2 abundance against YOY and yearling trawl indices. Standard errors are produced from the ADMB catch-age analysis report. 2 Parameters Rate Functions 2 Parameters Stock Biomass Survival Mean Stock Size (numbers) Mortality Rates Rate Stock Size (numbers) Weight in millions kg millions lbs. Age Mean Std. Err. Min. Max. (F) (Z) (A) (u) (S) Age Mean Min. Max. Pop. (kg) 2 2 2 Unit 1 2 6.3 3.83 2.66 1.36.33.33.32.27.68 2 77.997.128 1.866.66.1.18 11.31 3 33.639 1.3 19.1 8.173.22.62.79.18.21 3.216 1.72 6.78.9 3.128.379.837 2.998 1.119 1.879.117.3.83.66.29.3 17.2 9.9 2.89.11.318 1.997..73 2.186 3.8 7.92.3.93.61.37.38 1.3.81 1.78.18.96.2.3 6+ 3.31 1.372 1.979.723.76 1.16.669.27.331 6+ 3.32 2.23.616.222.72.737 1.62 Total 2.226 23. 29.172 7.28.28.68.9.26. Total 1.33 69.61 139.6.81.86 8.67 18.669 (3+).723 19.211 26.12 6.93.32.72.16.231.8 (3+) 26.36 1.13 38.199.126.76 3.319 7.319 Unit 2 2 3.97 2.21 1.76 6.18.7.7.37.6.62 2 9.7 6.86 11.13.86.293 7.87 17.21 3 37.898 1.276 22.622 3.17.28.928.6.3.39 3 2.2 1.67 3.818.133.96.32.717 2.16.863 1.3 3.279.76 1.16.682..318 1.98 8.92 21.19.218.7 3.261 7.189 3.7 1.3 2.36..831 1.231.78.78.292.768.9 1.2.29 1.33.223.92 6+ 3.88 1.123 1.96.211.812 1.212.72.71.298 6+ 2.1 1.27 2.726.31.97.628 1.38 Total 1.13 2.83 3.21 71.816.27.927.6.33.396 Total 11.666 77.623 13.78.111 7.7 12.23 26.996 (3+) 7.16 18.62 28. 6.78.76.976.623.368.377 (3+) 2.191 11.777 28.6.22 7.12.37 9.783 Unit 3 2 1.271.73.36 2.6...38.3.62 2 31.772 22.133 1.11.76.93 2.2.36 3 1.19 6.8 8.629 21.89.161.61.29.123.71 3.81.3 1.287.13 1.889.11.23 1.676.63 1.1 2.311.27.67.76.182.2 8.62.923 12.318.21.32 1.73 3.81.61 1.686 2.928 6.3.267.667.87.19.13.877. 1.21.28 1.283.26.2 6+ 7.1 2.98.93 1.139.21.61.79.18.21 6+ 6.3.8 8.2.339 2.71 2.133.73 Total 3.21 18.76 2.3.198.98..19. Total 8.38 32.2 6.7.137 6.28 6.6 1.69 (3+) 28.939 17..338.2.6..1.6 (3+) 16.613 9.892 23.33.2 6.188.219 9.33 Unit 2.77.376.11.83.1.1.3.12.66 2 3.7 2..11.83.6.296.6 3.13 3.29 1.836 8.2...366..63 3.31.67.63.18.78..19.69.7.283 1.97.91.91.388.72.612 3.2 1.16.3.22.11.717 1.8.7.329.26.9.16.6.32.126.68.22.173.671.263.1.111.2 6+ 2.16 1.193.913 3.299.1..23.11.77 6+ 1.2.666 2.17.33.691.8 1.12 Total 8.977 3.378 1.76.8.8.38.67.616 Total 9.18.111 1.16.18 1.813 1.682 3.79 (3+) 8. 3.277 13.723.88.88.386.7.61 (3+).33 2.71 8.996.2 1.766 1.38 3. 21

Table 2.1.1. Management Unit 1 yellow perch biological references from simulations and projected population size in 26 for a range of fishing rates. Biological reference points include mean spawner biomass as a fraction of an unfished population, mean survival of age 2+ and 3+ fish, and the probability of attaining low population levels observed in 1993- for ages 2+ (12.9 million) and 3+ (.1 million). The harvest strategy applied in the "Harvest 2" column, is based on the S/R F.1 stock recruitment simulation from the 2 YPTG report (using ADMB abundance estimates from 197-23). Refer to Table 2.2.1 for summary of F.1 fishing rates and 2 recommended harvest by management unit. Simulation Projections at Different Fishing Rates % Spawner Biomass (of Unfished) Survival 2+ Survival 3+ Prob %. 1993 2+ Prob. % 199 3+ F Harvest (lbs x 1 6 ) 2 Population 2+ (millions) 26 Population 3+ (millions) 26 Harvest Strategy Reference 1 67% 67%.. 71.6 69.9 86 6% 63%.1.6 7. 68. 7 62% 9%.2 1.177 68. 66.8 7 61% 8%.2 1.2 67.8 66.1 66 6% 6%.1.3 1.72 67.1 6. 62 9% %.3.3 1.98 66. 6.7 9 8% 3%.. 2.23 6.7 6. 6 7% 2%.. 2.82 6. 63. 6% %.. 2.72 6. 62.8 1 6% 9%.6. 2.99 63.8 62.1 9 % 8%.7.6 3.188 63.2 61. 7 % 7% 1..2.6 3.12 62.6 6.9 3% 6% 1..3.7 3.631 62. 6.3 3% % 1.6.3.72 3.716 61.8 6.1 F.1 SR 3 3% % 1.7.3.7 3.8 61. 9.8 2 2% 3% 2...8.2 6.9 9.2 2% 2% 2...8.2 6.3 8.7 39 1% 1% 2.8.6.9.3 9.8 8.1 37 1% 1% 3..9.9.66 9.3 7.6 36 % % 3.2 1.1 1..83 8.8 7.1 32 8% 36% 6. 2.7 1.2.9 6.8.2 29 7% 33% 9.8.1 1. 6.22.1 3. 26 % 31% 13. 9.3 1.6 6.82 3. 1.8 2 % 28% 16.7 1.1 1.8 7.3 1.9.3 22 2% 26% 2. 19.3 2. 7.86. 8.9 Parameters in Computations 2 Stock Size (numbers x 1 6 ) 26 Recruitment Age s(age) Weight (kg) Age Mean Min. Max. Millions Age 2s 2.8.1 2 77.997.128 1.866 1.6 3.388.12 3.216 1.72 6.78.72.12 17.2 9.9 2.89.76.172 1.3.81 1.78 6.81.21 6+ 3.32 2.23.616 (2+) 1.33 69.61 139.6 (3+) 26.36 1.13 38.199 22

Table 2.1.2. Management Unit 2 yellow perch biological references from simulations and projected population size in 26 for a range of fishing rates. Biological reference points include mean spawner biomass as a fraction of an unfished population, mean survival of age 2+ and 3+ fish, and the probability of attaining low population levels observed in 1993- for ages 2+ (17.8 million) and 3+ (7. million). The harvest strategy applied in the "Harvest 2" column, is based on the S/R F.1 stock recruitment simulation from the 2 YPTG report (using ADMB abundance estimates from 197-23). Refer to Table 2.2.1 for summary of F.1 fishing rates and 2 recommended harvest by management unit. Simulation Projections at Different Fishing Rates % Spawner Biomass (of Unfished) Survival 2+ Survival 3+ Prob %. 1993 2+ Prob. % 199 3+ F Harvest (lbs x 1 6 ) 2 Population 2+ (millions) 26 Population 3+ (millions) 26 Harvest Strategy Reference 1 67% 67%.. 76. 7.2 9 6% 63%.1.1.761 7. 72.3 82 61% 9%..2 1.83 72.7 7. 78 6% 7%.8.2 1.83 71.8 69.6 7 8% %.9.1.3 2.169 71. 68.8 72 7% % 1.7.1.3 2. 7.1 67.9 69 6% 2% 2.1.3. 2.823 69.3 67.1 67 % 1% 2.7.. 3.138 68. 66. 6 % 9% 3.6.9. 3. 67.8 6.6 62 3% 8%.2 1.2. 3.7 67. 6.8 6 2% 6%.6 2.1.6.39 66.3 6.1 8 2% % 7.2 3.2.6.32 6.6 63. 8 1% % 7. 3..661.387 6. 63.2 F.1 SR 6 1% % 8.1.2.7.6 6.9 62.7 % 3% 8.7.9.7.879 6.2 62. 3 9% 2% 1. 7..8.17 63. 61.3 2 9% 1% 12.3 9.2.8.9 62.8 6.7 8% % 13.9 11..9.66 62.2 6. 9 7% 39% 1.6 13.2.9.916 61.6 9. 8 7% 38% 16.1 1.9 1. 6.161 6.9 8.8 3 % 3% 2.3 23.2 1.2 7.9 8.6 6. 39 2% 31% 28.1 3.3 1. 7.96 6..2 36 % 28% 3.2 7. 1.6 8.7. 2.2 32 38% 26% 1.6 9.6 1.8 9.97 2..3 3 36% 23% 7.2 69.8 2. 1.192.7 8. Parameters in Computations 2 Stock Size (numbers x 1 6 ) 26 Recruitment Age s(age) Weight (kg) Age Mean Min. Max. Millions Age 2s 2.11.12 2 9.7 6.86 11.13 2.192 3.86.17 3 2.2 1.67 3.818.83.161 1.98 8.92 21.19.86.183.768.9 1.2 6.819.227 6+ 2.1 1.27 2.726 (2+) 11.666 77.623 13.78 (3+) 2.191 11.777 28.6 23

Table 2.1.3. Management Unit 3 yellow perch biological references from simulations and projected population size in 26 for a range of fishing rates. Biological reference points include mean spawner biomass as a fraction of an unfished population, mean survival of age 2+ and 3+ fish, and the probability of attaining low population levels observed in 1993- for ages 2+ (6.9 million) and 3+ (2.9 million). The harvest strategy applied in the "Harvest 2" column, is based on the S/R F.1 stock recruitment simulation from the 2 YPTG report (using ADMB abundance estimates from 197-23). Refer to Table 2.2.1 for summary of F.1 fishing rates and 2 recommended harvest by management unit. Simulation Projections at Different Fishing Rates % Spawner Biomass (of Unfished) Survival 2+ Survival 3+ Prob %. 1993 2+ Prob. % 199 3+ F Harvest (lbs x 1 6 ) 2 Population 2+ (millions) 26 Population 3+ (millions) 26 Harvest Strategy Reference 1 67% 67%.. 3.6 32. 89 6% 63%.1.68 33. 31.3 79 61% 9%.2 1.12 32. 3.3 7 9% 7%.2 1.37 32. 29.8 72 8% %.3 1.63 31. 29.3 69 7% 3%.1.3 1.83 31. 28.8 66 % 2%.2. 2.7 3. 28. 63 % %.7. 2.3 3.1 27.9 6 3% 8%.9.1. 2.18 29.7 27. 8 2% 7% 1.7.1. 2.73 29.2 27.1 6 1% 6% 2.7.6.6 2.936 28.8 26.7 % % 3.1.9.6 3.13 28. 26.3 2 9% 3% 3. 1.8.7 3.329 28.1 2.9 2 9% 3% 3.7 1.8.73 3.3 28. 2.9 F.1 SR 9% 2%.9 2.7.7 3.17 27.7 2. 8 8% 1% 6.7 3.3.8 3.7 27.3 2.1 7 7% % 7.7.7.8 3.878 27. 2.8 6% 38% 9. 7..9. 26.6 2. 6% 37% 11.8 1.1.9.218 26.3 2.1 3 % 36% 1.2 13.1 1..381 26. 23.8 38 2% 33% 21.7 27. 1.2.99 2.7 22.6 3 % 29% 31.3.8 1..36 23.6 21. 31 38% 27% 3.8 63. 1.6 6.27 22.6 2. 28 36% 2% 2.1 78. 1.8 6.72 21.7 19. 26 3% 22% 61.3 88. 2. 6.87 3. 2.9 Parameters in Computations 2 Stock Size (numbers x 1 6 ) 26 Recruitment Age s(age) Weight (kg) Age Mean Min. Max. Millions Age 2s 2.13.12 2 31.772 22.133 1.11 2.177 3.32.17 3.81.3 1.287.79.179 8.62.923 12.318.823.222.877. 1.21 6.789.28 6+ 6.3.8 8.2 (2+) 8.38 32.2 6.7 (3+) 16.613 9.892 23.33 2