SUBCLASS ELASMOBRANCHII

Transcription

1 INFRAPHYLUM Gnathostomata Gegenbaur, jawed vertebrates CLASS Chondrichthyes Huxley, cartilaginous fishes SUBCLASS ELASMOBRANCHII Bonaparte, 1838 INFRACLASS Euselachii Hay, 1902 SUBDIVISION Selachii (no authority discovered) SUPERORDER Galeomorphii Compagno, 1973 ORDER Heterodontiformes Berg, bullhead sharks FAMILY Heterodontidae Gray, bullhead and horn sharks [=Cestraciontini, Heterodontina, Centraciontidae] Notes: Name in prevailing recent practice, Article 35.5 Cestraciontini Bonaparte, 1835:[4] [ref ] (subfamily) Cestracion Bosc [genus inferred from the stem, Article ; senior objective synonym of Heterodontina Gray, 1851; family name sometimes seen as Cestraciondontidae] Heterodontina Gray, 1851:65 [ref. 4939] (tribe) Heterodontus [junior objective synonym of Cestraciontini Bonaparte, 1835, but Heterodontidae in prevailing recent practice; Heterodontidae used as valid by: Jordan 1923 [ref. 2421], Schultz with Stern 1948 [ref ], Kamohara 1967, Lindberg 1971 [ref ], Nelson 1976 [ref ], Shiino 1976, Lagler, Bardach, Miller & May Passino 1977, Nelson 1984 [ref ], Smith & Heemstra 1986 [ref. 5715], Paxton et al [ref ], Robins et al. 1991a [ref ], Allen & Robertson 1994 [ref ], Last & Stevens 1994 [ref ], Nelson 1994 [ref ], Eschmeyer 1998 [ref ], Compagno 1999 [ref ], Chu & Meng 2001,

2 Compagno 2001 [ref ], Nelson et al [ref ], Baldwin 2005 [ref ], Hoese et al. 2006, Nelson 2006 [ref ], Last & Stevens 2009; family name sometimes seen as Heterondontidae] Centraciontidae Garman, 1913:13, 180 [ref. 1545] (family) Centracion GENUS Heterodontus Blainville, bullhead sharks, horn sharks [=Heterodontus (subgenus of Squalus) Blainville [H. de], 1816:121, Centracion Gray [J. E.], 1831:5, Cestracion Bosc [L. A. G.] (ex Cuvier), 1816:598, Gyropleurodus Gill [T. N.], 1862:331, Molochophrys Whitley [G. P.], 1931:310, Tropidodus Gill [T. N.], 1863:489, 490, Wuia (subgenus of Heterodontus) Fowler [H. W.], 1934:233] Notes: [ref. 306]. Masc. Squalus philippi Bloch & Schneider, Type by monotypy. Misspelled Heterodontes by Gill 1862:403 [ref. 1783]. Valid as Heterodontus Blainville, (Taylor & Castro-Aguirre 1972 [ref. 7158], Compagno 1984:155 [ref. 6474], Nakaya & Shirai in Masuda et al. 1984:3 [ref. 6441], Bass 1986:48 [ref. 5635], Cappetta 1987:70 [ref. 6348], Paxton et al. 1989:95 [ref ], Gomon et al. 1994:90 [ref ], Castro-Aguirre et al. 1999:41 [ref ], Compagno 1999:475 [ref ], Compagno 2001:34 [ref ], Compagno et al. 2005:15 [ref ], Hoese & Gates 2006:60 [ref ], White 2008:32 [ref ], Castro 2011:174 [ref ]). Current status: Valid as Heterodontus Blainville, Heterodontidae. (Centracion) [ref. 1880]. Neut. Centracion zebra Gray, Type by monotypy. Perhaps intended for Cestracion but unclear from text. Synonym of Heterodontus Blainville, 1816 (Compagno 1984:155 [ref. 6474]). Current status: Synonym of Heterodontus Blainville, Heterodontidae. (Cestracion) [ref. 5126]. Squalus philippi Bloch & Schneider, Type by subsequent monotypy. Also latinized as Cestracion by Oken (ex Cuvier) 1817:Cestracio by Cloquet 1817:20 [ref. 852], and Cestralion by Müller & Henle 1838:85 [ref. 3068]. Cestracion Gray, 1831:5 [ref. 1880] is not an original genus description [contra Compagno 2001:34 [ref ]].

3 Objective synonym of Heterodontus Blainville, (Compagno 1984:155 [ref. 6474], Cappetta 1987:70 [ref. 6348], Compagno 2001:34 [ref ]). Current status: Synonym of Heterodontus Blainville, Heterodontidae. (Gyropleurodus) [ref. 1668]. Masc. Cestracion francisci Girard, Type by monotypy. Synonym of Heterodontus Blainville, (Compagno 1984:155 [ref. 6474], Compagno 2001:34 [ref ]). Current status: Synonym of Heterodontus Blainville, Heterodontidae. (Molochophrys) [ref. 4672]. Fem. Cestracion galeatus Günther, Type by original designation (also monotypic). Synonym of Heterodontus Blainville, (Compagno 1984:155 [ref. 6474], Compagno 2001:34 [ref ]). Current status: Synonym of Heterodontus Blainville, Heterodontidae. (Tropidodus) [ref. 1671]. Masc. Cestracion pantherinus Valenciennes, Type by original designation (also monotypic). Type designated in footnote on p Misspelled Tropidopus by Beebe & Tee-Van 1941:118 [ref ]. Synonym of Heterodontus Blainville, 1816 (Compagno 1984:155 [ref. 6474], Compagno 2001:34 [ref ]). Current status: Synonym of Heterodontus Blainville, Heterodontidae. (Wuia) [ref. 1416]. Fem. Centracion zebra Gray, Type by original designation (also monotypic). Objective synonym of Centracion Gray, Synonym of Heterodontus Blainville, (Compagno 1984:155 [ref. 6474], Compagno 2001:34 [ref ]). Current status: Synonym of Heterodontus Blainville, Heterodontidae. Species Heterodontus francisci (Girard, 1854) - California horn shark [=Cestracion francisci Girard [C. F.], 1855:196, Heterodontus californicus Herald [E. S.], 1961:49] Notes: [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7 (no. 6); ref ] Monterey Bay, California, U.S.A. Current status: Valid as Heterodontus francisci (Girard, 1855). Heterodontidae. Distribution: Eastern Pacific.

4 (californicus) [Living fishes of the World; ref ] Current status: Synonym of Heterodontus francisci (Girard, 1855). Heterodontidae. Species Heterodontus galeatus (Günther, 1870) - crested bullhead shark, crested hornshark [=Cestracion galeatus Günther [A.], 1870:416] Notes: [Catalogue of the fishes in the British Museum v. 8; ref. 1995] New South Wales, Australia. Current status: Valid as Heterodontus galeatus (Günther, 1870). Heterodontidae. Distribution: Australia. Species Heterodontus japonicus Maclay & Macleay, Japanese bullhead shark [=Heterodontus japonicus Miklouho-Maclay [N. de] & Macleay [W.], 1884:428, Pl. 20] Notes: [Proceedings of the Linnean Society of New South Wales v. 8 (pt 4); ref ] Tokyo, Japan. Current status: Valid as Heterodontus japonicus Miklouho-Maclay & Macleay, Heterodontidae. Distribution: Western North Pacific. Species Heterodontus mexicanus Taylor & Castro-Aquirre, Mexican hornshark [=Heterodontus mexicanus Taylor [L. R., Jr.] & Castro-Aguirre [J. L.], 1972:125, Figs. 1-2, 5-6] Notes: [Anales de la Escuela Nacional de Ciencias Biológicas México v. 19; ref. 7158] Cerro Colorado, Sonora, Mexico, Gulf of California, depth 20 meters. Current status: Valid as Heterodontus mexicanus Taylor & Castro-Aguirre, Heterodontidae. Distribution: Eastern Pacific: Gulf of California and Mexico to Peru.

5 Species Heterodontus omanensis Baldwin, Oman bullhead shark [=Heterodontus omanensis Baldwin [Z. H.], 2005:262, Figs. 1-2] Notes: [Copeia 2005 (no. 2); ref ] Northeast of Ra's al Madrakah, Gulf of Masira, central Oman, Arabian Sea, 19 46'N, 58 19'E, depth 72 meters. Current status: Valid as Heterodontus omanensis Baldwin, Heterodontidae. Distribution: Northwestern Indian Ocean: Oman. Species Heterodontus portusjacksoni (Meyer, 1793) - Port Jackson shark [=Squalus portusjacksoni Meyer [F. A. A.], 1793:71, Heterodontus bonaespei Ogilby [J. D.], 1908:26, Cestracion heterodontus Sherrard [J. E.], 1896:42, 88, Squalus jacksoni Suckow [G. A.], 1799:102, Squalus philippi Bloch [M. E.] & Schneider [J. G.], 1801:134, Squalus philippinus Shaw [G.], 1804:341, Squalus superciliosus Forster [J. R.], 1794:131] Notes: [Systematisch-summarische Uebersicht der neuesten zoologischen Entdeckungen in Neuholland und Afrika.; ref ] Port Jackson, New South Wales, Australia. Current status: Valid as Heterodontus portusjacksoni (Meyer, 1793). Heterodontidae. Distribution: Australia and New Zealand. (bonaespei) [Proceedings of the Royal Society of Queensland v. 21; ref. 3286] Table Bay, South Africa [locality questioned in literature] (heterodontus) [Illustrated official handbook to the aquarium; ref ] Hobson's Bay, Victoria,

6 Australia. (jacksoni) [Naturgeschichte der Thiere v. 4, pt. 1; ref ] Port Jackson, New South Wales, Australia. (philippi) [M. E. Blochii, Systema Ichthyologiae; ref. 471] Botany Bay, New South Wales, Australia. (philippinus) [General zoology or systematic natural history; ref. 4015] Botany Bay, New South Wales, Australia. (superciliosus) [Die Neuesten Reisen nach der Botany Bay; ref ] Botany Bay, New South Wales, Australia. Species Heterodontus quoyi (Freminville, 1840) - Galapagos bullhead shark, Peruvian hornshark [=Cestracion quoyi Fréminville [C. P. P.], 1840:3, Pl. 3, Cestracion pantherinus Valenciennes [A.], 1846:no p., Pl. 10 (fig. 2), Gyropleurodus peruanus Evermann [B. W.] & Radcliffe [L.], 1917:2, Pl. 1 (fig. 1)] Notes: [Magazin de Zoologie, d'anatomie Comparée et de Paléontologie (Sér. 2). v. 2, Pt 5; ref ] Galápagos Islands.

7 Current status: Valid as Heterodontus quoyi (Fréminville, 1840). Heterodontidae. Distribution: Eastern Pacific. (pantherinus) [Voyage autour du monde sur la frégate "la Vénus"; ref. 6165] Galápagos Islands. Current status: Synonym of Heterodontus quoyi (Fréminville, 1840). Heterodontidae. (peruanus) [Bulletin of the United States National Museum No. 95; ref. 1284] Lobos de Tierra Island, Peru. Current status: Synonym of Heterodontus quoyi (Fréminville, 1840). Heterodontidae. Species Heterodontus ramalheira (Smith, 1949) - whitespotted bullhead shark [=Gyropleurodus ramalheira Smith [J. L. B.], 1949:367, Fig. 1] Notes: [Annals and Magazine of Natural History (Series 12) v. 2 (no. 17) (art. 30); ref. 4079] Off Inhambane, Mozambique, western Indian Ocean. Current status: Valid as Heterodontus ramalheira (Smith, 1949). Heterodontidae. Distribution: Western Indian Ocean. Species Heterodontus zebra (Gray, 1831) - striped catshark, zebra bullhead shark, barred bullhead shark, barred shark, little shark, striped bullhead shark, zebra hornshark, zebra Port Jackson shark [=Centracion zebra Gray [J. E.], 1831:5, Cestracion amboinensis Regan [C. T.], 1906:436, Cestracion philippi var. japonica Duméril [A. H. A.], 1865:426] Notes: [Zoological Miscellany 1831 (art. 6); ref. 1880] Swatow, China.

8 Current status: Valid as Heterodontus zebra (Gray, 1831). Heterodontidae. Distribution: Indo-West Pacific. (amboinensis) [Annals and Magazine of Natural History (Series 7) v. 18 (no. 108); ref ] Ambon Island, Molucca Islands, Indonesia. Current status: Synonym of Heterodontus zebra (Gray, 1831). Heterodontidae. (japonica) [Histoire naturelle des poissons ou ichthyologie générale v. 1; ref. 1150] Current status: Synonym of Heterodontus zebra (Gray, 1831). Heterodontidae. Habitat: marine