Hemiscylliidae Chiloscyllium Current status Valid
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1 FAMILY Hemiscylliidae Gill, bamboosharks, epaulette sharks [=Hemiscylliinae, Chiloscylliinae] Notes: Hemiscylliinae Gill, 1862p:407, 408 [ref. 1783] (subfamily) Hemiscyllium [Gill 1893b:130 [ref ] selected Hemiscylliinae over Chiloscylliinae] Chiloscylliinae Gill, 1862p:407, 408 [ref. 1783] (subfamily) Chiloscyllium GENUS Chiloscyllium Müller & Henle, bamboosharks [=Chiloscyllium Müller [J.] & Henle [F. G. J.], 1837:112, Synchismus Gill [T. N.], 1862:407, 408] Notes: [ref. 3067]. Neut. Scyllium plagiosum Bennett, Type by subsequent monotypy (or designated by Gill 1861:407 [ref. 4910]). Appeared without species as above and in Müller & Henle 1837:395 [ref ], then apparently next in Müller & Henle in Smith 1837:85. Also appeared in Müller & Henle 1838:17 [ref. 3069]. Spelled Chyloscyllium and Cheiloscyllium in early literature. Valid as Chiloscyllium Müller & Henle, (Dingerkus & Defino 1983:7 [ref. 5386], Compagno 1984:189 [ref. 6474], Nakaya & Shirai in Masuda et al. 1984:8 [ref. 6441], Bass 1986:64 [ref. 5635], Kharin 1987 [ref. 6227], Cappetta 1987:72 [ref. 6348], Paxton et al. 1989:89 [ref ], Compagno 1999:476 [ref ], Compagno 2001:165 [ref ], Goto 2001:82 [ref ], Compagno et al. 2005:20 [ref ], Hoese et al. 2006:123 [ref ], Kimura 2009:7 [ref ], Moore et al. 2012:5 [ref ], Weigmann 2012:3 [ref ], Allen et al. 2013:124 [ref ], Kottelat [ref ]). Current status: Valid as Chiloscyllium Müller & Henle, Hemiscylliidae. (Synchismus) [ref. 1783]. Masc. Squalus tuberculatus Bloch & Schneider, Type by original designation (also monotypic). Also in Gill 1862:413 [ref. 4910]. Synonym of Chiloscyllium Müller & Henle, (Dingerkus & DeFino 1983:7 [ref. 5386], Compagno 1984:189 [ref. 6474], Kharin 1987:364 [ref. 6227], Cappetta 1987:72 [ref. 6348], Compagno 2001:166 [ref ], Goto 2001:82 [ref ], Kottelat [ref ]). Current status: Synonym of Chiloscyllium Müller & Henle, Hemiscylliidae.
2 Species Chiloscyllium arabicum Gubanov, in Gubanov & Shleib, Arabian carpetshark, confusing bambooshark, Arabian bambooshark [=Chiloscyllium arabicum Gubanov [E. P.], in Gubanov & Shleib, 1980:14, Figs. 6-7, Chiloscyllium confusum Dingerkus [G.] & DeFino [T. C.], 1983:9, Figs. 2, 4-7, 51, 57, 61-64] Notes: [Sharks of the Arabian Gulf; ref ] Persian Gulf, Current status: Valid as Chiloscyllium arabicum Gubanov, Hemiscylliidae. Distribution: Northwestern Indian Ocean. (confusum) [Bulletin of the American Museum of Natural History v. 176 (art. 1); ref. 5386] Offshore of Calicut (Kozhikode), Kerala State, India. Current status: Synonym of Chiloscyllium arabicum Gubanov, Hemiscylliidae. Distribution: India. Species Chiloscyllium burmensis Dingerkus & Defino, Burmese bambooshark [=Chiloscyllium burmensis Dingerkus [G.] & DeFino [T. C.], 1983:9, Figs. 3, 7, 59-60] Notes: [Bulletin of the American Museum of Natural History v. 176 (art. 1); ref. 5386] Off Rangoon, 15 04'N, 95 51'E, Myanmar, Anton Bruun, cruise 1, station 41A, depth meters. Current status: Valid as Chiloscyllium burmensis Dingerkus & DeFino, Hemiscylliidae. Distribution: Myanmar. Species Chiloscyllium caeruleopunctatum Pellegrin, blue spotted bamboo shark [=Chiloscyllium caeruleopunctatum Pellegrin [J.], 1914:230]
3 Notes: [Bulletin de la Société Zoologique de France v. 39; ref ] Fort Dauphin, Madagascar. Current status: Valid as Chiloscyllium caeruleopunctatus Pellegrin, Hemiscylliidae. Distribution: Madgascar. Species Chiloscyllium griseum Müller & Henle, grey bambooshark [=Chiloscyllium griseum Müller [J.] & Henle [F. G. J.], 1838:19, [Pl. 4], Chiloscyllium obscurum Gray [J. E.], 1851:35, Squalis (Scyliorhinus) unicolor Blainville [H. de], 1816:121] Notes: [Systematische Beschreibung der Plagiostomen; ref. 3069] Puducherry, India; Japan. Current status: Valid as Chiloscyllium griseum Müller & Henle, Hemiscylliidae. Distribution: Indo-West Pacific: Arabian Sea to East Indies region, north to southern Japan. (obscurum) [List of the specimens of fish in the collection of the British Museum.; ref. 4939] Current status: Synonym of Chiloscyllium griseum Müller & Henle, Hemiscylliidae. (unicolor) [Bulletin de la Société Philomathique de Paris v. 8; ref. 306] Current status: Synonym of Chiloscyllium griseum Müller & Henle, Hemiscylliidae. Species Chiloscyllium hasseltii Bleeker, Hasselt's bambooshark, Indonesian bambooshark [=Chiloscyllium hasseltii Bleeker [P.], 1852:19, Chiloscyllium dolganovi Kharin [V. E.], 1987:366, Fig. 5] Notes: [Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen. v. 24 (art. 12); ref. 333] Molucca Islands, Indonesia. Current status: Valid as Chiloscyllium hasseltii Bleeker, Hemiscylliidae. Distribution: Indo-West Pacific: East Indian Region.
4 (dolganovi) [Voprosy Ikhtiologii v. 27 (no. 3); ref. 6227] Gulf of Thailand, Vietnam, 9 12'01"N, '06"E, depth 12 meters. Current status: Synonym of Chiloscyllium hasseltii Bleeker, Hemiscylliidae. Species Chiloscyllium indicum (Gmelin, 1789) - ridgeback catshark, ridgebacked bambooshark, slender bambooshark, stone shark, time teller shark, frog shark [=Squalus indicus Gmelin [J. F.], 1789:1503, Squalus caudatus Gronow [L. T.], in Gray, 1854:8, Hemiscyllium colax Fowler [H. W.], 1941:89, Squalus (Scyliorhinus) variegatus Blainville [H. de], 1816:121, Squalus colax Meuschen [F. C.], 1781, Squalus (Scyliorhinus) dentatus Blainville [H. de], 1816:121, Squalus denticulatus Shaw [G.], 1804:351, Squalus gronovianus Shaw [G.], 1804:353, Chiloscyllium phymatodes Bleeker [P.], 1852:21, Squalus tuberculatus Bloch [M. E.] & Schneider [J. G.], 1801:137] Notes: [Caroli a Linné... Systema Naturae per regna tria naturae v. 1 (pt 3); ref ] Indian Ocean. Current status: Valid as Chiloscyllium indicum (Gmelin, 1789). Hemiscylliidae. Distribution: Indo-West Pacific: India and Sri Lanka to western Indonesia, north to China. Habitat: brackish, marine. (caudatus) [Catalogue of fish collected and described by Laurence Theodore Gronow; ref. 1911] Indian seas. (colax) [Bulletin of the United States National Museum No. 100, v. 13; ref. 6536] China Seas and
5 other areas. (variegatus) [Bulletin de la Société Philomathique de Paris v. 8; ref. 306] Current status: Synonym of Chiloscyllium colax Fowler, Hemiscylliidae. (colax) [Index to Gronovius, Zoophylacium Gronovianum; ref ] (dentatus) [Bulletin de la Société Philomathique de Paris v. 8; ref. 306] (denticulatus) [General zoology or systematic natural history; ref. 4015] No locality. (gronovianus) [General zoology or systematic natural history; ref. 4015] Indian Seas. (phymatodes) [Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen. v. 24 (art. 12) ref. 333] Semarang, Java, Indonesia. (tuberculatus) [M. E. Blochii, Systema Ichthyologiae; ref. 471] No locality. Species Chiloscyllium plagiosum (Bennett, 1830) - whitespotted bambooshark, bluespotted bambooshark [=Scyllium plagiosum Anonymous [Bennett [E. T.]], 1830:694, Chiloscyllium plagiosum var. interruptum Bleeker [P.], 1852:18,
6 Chiloscyllium margaritiferum Bleeker [P.], 1863:243, Scyllium ornatum Gray [J. E.], 1830:no page number, Pl. 98 (fig. 2)] Notes: [Memoir of the life and public services of Sir Thomas Stamford Raffles; ref. 259] Java Sea, 5 58'S, 'E, Sumatra, Indonesia. Current status: Valid as Chiloscyllium plagiosum (Anonymous [Bennett], 1830). Hemiscylliidae. Distribution: Indo-West Pacific: Madagascar and India to East Indian Region and ranging north to Japan. (interruptum) [Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen. v. 24 (art. 12); ref. 333] Jakarta and Semarang, Java, Indonesia. Current status: Synonym of Chiloscyllium plagiosum (Anonymous [Bennett], 1830). Hemiscylliidae. (margaritiferum) [Nederlandsch Tijdschrift voor de Dierkunde v. 1; ref ] Obi Island, Molucca Islands, Indonesia. Current status: Synonym of Chiloscyllium plagiosum (Anonymous [Bennett], 1830). Hemiscylliidae. (ornatum) [Illustrations of Indian zoology; ref. 1878] China seas. Current status: Synonym of Chiloscyllium plagiosum (Anonymous [Bennett], 1830). Hemiscylliidae. Species Chiloscyllium punctatum Müller & Henle, brownbanded bambooshark, brownspotted catshark, grey carpetshark, spotted catshark [=Chiloscyllium punctatum Müller [J.] & Henle [F. G. J.] (ex Kuhl & van Hasselt), 1838:18, [Pl. 3], Scylia russelii Billberg [G. J.], 1833:51,
7 Squalus (Scyliorhinus) russellianus Blainville [H. de], 1816:121] Notes: [Systematische Beschreibung der Plagiostomen; ref. 3069] Off Jakarta, Java, Indonesia. Current status: Valid as Chiloscyllium punctatum Müller & Henle, Hemiscylliidae. Distribution: Indo-West Pacific: India east to East Indian region, and to Japan and Australia. (russelii) [Linnéska Samfundets Handlingar v. 1 (for 1832); ref ] Vizagapatnam, India. Current status: Synonym of Chiloscyllium punctatum Müller & Henle, Hemiscylliidae. (russellianus) [Bulletin de la Société Philomathique de Paris v. 8; ref. 306] Current status: Synonym of Chiloscyllium punctatum Müller & Henle, Hemiscylliidae. GENUS Hemiscyllium Müller & Henle, epaulette sharks [=Hemiscyllium Müller [J.] & Henle [F. G. J.], 1838:34] Notes: [ref. 3066]. Neut. Squalus ocellatus Bonnaterre, Type by monotypy. Predates Hemiscyllium Müller & Henle, in Smith, (13 Feb. 1838), Müller & Henle apparently published 2 Jan. 1838; see Dingerkus & DeFino 1983:35 [ref. 5386]. Also in Müller & Henle 1838:83 [ref. 3068]. Valid as Hemiscyllium Müller & Henle, (Dingerkus & DeFino 1983:35 [ref. 5386], Compagno 1984:195 [ref. 6474], Cappetta 1987:73 [ref. 6348], Paxton et al. 1989:89 [ref ], Compagno 1999:476 [ref ], Compagno 2001:177 [ref ], Goto 2001:81 [ref ], Hoese et al. 2006:123 [ref ], Allen & Erdmann 2008:94 [ref ], Allen & Dudgeon 2010:20 [ref ], Allen et al. 2013:123 [ref ]). Current status: Valid as Hemiscyllium Müller & Henle, Hemiscylliidae. Species Hemiscyllium freycineti (Quoy & Gaimard, 1824) - Indonesian speckled carpetshark, Freycinet's epaulette shark [=Scyllium freycineti Quoy [J. R. C.] & Gaimard [J. P.], 1824:192, Scyllium malaisianum Lesson [R. P.], 1831:94, Poissons Pl. 6]
8 Notes: [Voyage autour du monde; ref. 3574] Waigeo [Waigiou]; Irian Jaya, Indonesia. Current status: Valid as Hemiscyllium freycineti (Quoy & Gaimard, 1824). Hemiscylliidae. Distribution: Raja Ampat Islands, West Papua, Indonesia. (malaisianum) [Voyage autour du monde Zool. v. 2 (pt 1); ref. 2776] Offack Bay, Waigeo [Waigiou], Irian Jaya, Indonesia. Current status: Synonym of Hemiscyllium freycineti (Quoy & Gaimard, 1824). Hemiscylliidae. Habitat: marine Species Hemiscyllium galei Allen & Erdmann, walking shark [=Hemiscyllium galei Allen [G. R.] & Erdmann [M. V.], 2008:96, Figs. 2B, 3-6] Notes: [Journal of Ichthyology and Aquatic Biology v. 13 (nos 3-4); ref ] Reef near Rumberpon Village, 'S, 'E, western Cenderwasih Bay, Papua Batat Province, Indonesia. Current status: Valid as Hemiscyllium galei Allen & Erdmann, Hemiscylliidae. Distribution: Cenderwasih Bay, West Papua, Indonesia. Species Hemiscyllium hallstromi Whitley, Papuan epaulette shark [=Hemiscyllium hallstromi Whitley [G. P.], 1967:178] Notes: [Australian Zoologist v. 14 (no. 2); ref. 8001] New Guinea. Current status: Valid as Hemiscyllium hallstromi Whitley, Hemiscylliidae. Distribution: Papua New Guinea. Species Hemiscyllium halmahera Allen et al., Halmahera epaulette shark [=Hemiscyllium halmahera Allen [G. R.], Erdmann [M. V.] & Dudgeon [C. L.], 2013:126, Figs. 2-7] Notes: [aqua, International Journal of Ichthyology v. 19 (no. 3); ref ] Northwestern Ternate,
9 N, E, Halmahera, Indonesia, depth 10 meters. Current status: Valid as Hemiscyllium halmahera Allen, Erdmann & Dudgeon, Hemiscylliidae. Distribution: Halmahera, Ternate. Species Hemiscyllium henryi Allen & Erdmann, Triton Bay epaulette shark, Henry's epaulette shark [=Hemiscyllium henryi Allen [G. R.] & Erdmann [M. V.], 2008:102, Figs. 2B, 6-9] Notes: [aqua, Journal of Ichthyology and Aquatic Biology v. 13 (nos 3-4); ref ] Northwestern portion of Selat Iris, 'S, "E, Papua Barat Province, Indonesia. Current status: Valid as Hemiscyllium henryi Allen & Erdmann, Hemiscylliidae. Distribution: West Papua, Indonesia. Species Hemiscyllium michaeli Allen & Dudgeon, Michael's epaulette shark, leopard epaulette shark, Milne Bay carpetshark [=Hemiscyllium michaeli Allen [G. R.] & Dudgeon [C. L.], 2010:22, Figs. 2-4] Notes: [aqua, International Journal of Ichthyology v. 16 (no. 1); ref ] Coral reef in Harvey Bay, 'S, 'E, east of Oro Bay, Papua New Ginea. Current status: Valid as Hemiscyllium michaeli Allen & Dudgeon, Hemiscylliidae. Distribution: Eastern Papua New Guinea and Trobriand Islands. Species Hemiscyllium ocellatum (Bonnaterre, 1788) - blind shark, epaulette shark [=Squalus ocellatus Bonnaterre [J. P.] (ex Broussonet), 1788:8,
10 Squalus oculatus Banks [J.] & Solander [D. C.], in Gray, 1827:436 Notes: [Tableau encyclopédique et méthodique des trois règnes de la nature... Ichthyologie; ref. 4940] Cooktown, Queensland, Australia, Coral Sea, southwestern Pacific. Current status: Valid as Hemiscyllium ocellatum (Bonnaterre, 1788). Hemiscylliidae. Distribution: Northern Australia. (oculatus) [List of fish of Australia; ref ] Cooktown, Queensland, Australia. Current status: Synonym of Hemiscyllium ocellatum (Bonnaterre, 1788). Hemiscylliidae. Species Hemiscyllium strahani Whitley, hooded carpetshark [=Hemiscyllium strahani Whitley [G. P.], 1967:176, Fig. 1] Notes: [Australian Zoologist v. 14 (no. 2); ref. 8001] Vicinity of Port Moresby, Papua New Guinea. Current status: Valid as Hemiscyllium strahani Whitley, Hemiscylliidae. Distribution: Papua New Guinea. Species Hemiscyllium trispeculare Richardson, speckled carpetshark, speckled catshark, speckled epaulette shark [=Hemiscyllium trispeculare Richardson [J.], 1843:5, Pl. 1 (fig. 2)] Notes: [Icones piscium, or plates of rare fishes; ref. 6537] North West Australia. Current status: Valid as Hemiscyllium trispeculare Richardson, Hemiscylliidae. Distribution: Northern Australia. Habitat: marine ****************************** Unavailable names Chiloscyllium ornatum var. astigma Bleeker [P.], 1863:136 [Nederlandsch Tijdschrift voor de Dierkunde v. 1; ref. 396]
11 Amoy, China. Holotype (unique): whereabouts unknown. Not treated by Dingerkus & DeFino 1983 [ref. 5386]. Hemiscylliidae. Squalis (Scyliorhinus) lambarda Blainville [H. De], 1816:121 [Bulletin de la Société Philomathique de Paris v. 8; ref. 306] Not available, name only (see Dingerkus & DeFino 1983:56 [ref. 5386]). Questionably in the synonymy of Chiloscyllium colax [not available] -- (Kharin 1987:367 [ref. 6227]). Current status: Synonym of Chiloscyllium. Hemiscylliidae. Chiloscyllium ornatum var. melanostigma Bleeker [P.], 1863:136 [Nederlandsch Tijdschrift voor de Dierkunde v. 1; ref. 396] Singapore. Holotype (unique): whereabouts unknown. Not treated by Dingerkus & DeFino 1983 [ref. 5386]. Hemiscylliidae. Chiloscyllium indicum var. obscura Günther [A.], 1870:413 [Catalogue of the fishes in the British Museum v. 8; ref. 1995] Apparently not an original description; intended for Scyllium obscurum of authors. See Dingerkus & DeFino 1983:17 [ref. 5386]. Hemiscylliidae. Squalus ocellatus Gmelin [J. F.], 1789:1494 [Caroli a Linné... Systema Naturae per regna tria naturae v. 1 (pt 3); ref ] Objectively invalid; preoccupied by Squalus ocellatus Bonnaterre, Hemiscylliidae. Squalus ocellatus Walbaum [J. J.] 1792:516 [Petri Artedi sueci genera piscium Part 3; ref. 4572] No locality stated. Objectively invalid; preoccupied by Squalus ocellatus Bonnaterre, 1788 and Gmelin, Hemiscylliidae. Habitat: marine. Squalus oculatus Banks [J.], in Müller & Henle, 1838:16 [Systematische Beschreibung der Plagiostomen; ref. 3069] Not available; manuscript name mentioned under Hemiscyllium ocellatum.
Hemiscylliidae Chiloscyllium Current status Valid Chiloscyllium arabicum Current status Valid Distribution
FAMILY Hemiscylliidae Gill, 1862 - bamboosharks, epaulette sharks [=Hemiscylliinae, Chiloscylliinae] Notes: Hemiscylliinae Gill, 1862p:407, 408 [ref. 1783] (subfamily) Hemiscyllium [Gill 1893b:130 [ref.
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