Insecta: Diptera, Empididae

Save this PDF as:
 WORD  PNG  TXT  JPG

Size: px
Start display at page:

Download "Insecta: Diptera, Empididae"

Transcription

1 Insecta: Freshwater Diptera, Invertebrates Empididae of the Malaysian Region 805 Insecta: Diptera, Empididae Patrick Grootaert Royal Belgian Institute of Natural Sciences, Vautierstraat 29, B-1000 Brussels, Belgium ( INTRODUCTION The Empididae sensu stricto or true dance flies are small (2 mm) to large flies (15 mm), dark to yellow and generally with a sclerotized proboscis. They are very diverse in habitus (see Fig. 2) and have as a common characteristic a small indentation in the eye border at the level of the insertion of the antenna. They form a large family with an enormous radiation in the Palaearctic region. They are far less common in the tropics and will thrive only in mountainous areas. In the Malaysian region we expect the genera Hilara, Empis (Coptophlebia), Rhamphomyia which do not have true aquatic larvae and the Hemerodromiinae Hemerodromia, Chelifera and Clinocera which do have aquatic larvae. It is doubtful that other Palaearctic genera such as Chelipoda, and Wiedemannia are present here. GENERAL BIOLOGY The genus Hilara (Fig. 2A), an old dance fly genus that radiated in temperate regions, has an epigamic behaviour (to attract mates) associated with water bodies, but the larvae live mainly in the soil. The male hunts for small prey floating on the surface of pools, ponds, brooks or streams (Fig. 1). It picks up the prey and often wraps it in a silk cocoon, which is produced by silk glands in the enlarged fore basitarsi. Eventually the male carrying the cocoon joins a female swarm. Females tend to inspect the male with its present and when the female dives under the male to grab the prey, the male grabs the female s thorax and mating starts. Generally, the couple leaves the swarm for some vegetation. This behaviour is specific and has a number of variants. Light intensity of the area (Grootaert 1994) determined by the coverage of the canopy, the presence of markers in the landscape, the type of water surface (running or still, size) all determine where the male hunts and where female swarms are formed. Females sometimes swarm above water, but also terrestrial swarms are formed some distance from the place where the males hunt. Generally, the females fly in a horizontal plane in a rapid to-and-fro movement. However, triangular swarms are known as well, where the females dance up in a slow motion and at the top of the triangle, dive down vertically, and fly in a horizontal direction back to the point of departure and dance

2 806 Freshwater Invertebrates of the Malaysian Region Figure 1. Swarming behaviour of Hilara dance flies occurring in the gallery forest above the small streams. The male catches a prey floating on the water surface and enrobes it with silk. Eventually the male joins a swarm of females. The female inspects the offer, dives down below the male and grabs the cocoon while the male attempts mating. Figure 2. A Hilara isaanensis male (d: discal cell; arrow points to the upper branch of the fork); B Hemerodromia sp. from Northeast Thailand; C Empis (Coptophlebia) sp. from Thailand. (Arrow points to the upper branch of the fork)

3 Insecta: Diptera, Empididae 807 upwards again. The length of the swarm is limited by light zones (light-shade transition) and by location markers such as branches or vegetation. The genus Hemerodromia is associated with fast flowing streams and is predacious (Fig. 2B). The adults are small (3 5 mm), yellow or black coloured, and they are mainly active on boulders covered with moss or in the splash zone. They are present throughout the year and quite common. Their habitat is the small streams shaded by the gallery forest. This genus too is very sensitive to changing light conditions. Removing the evergreen gallery forest around the streams will destroy the populations of Hemerodromia. Empis (Coptophlebia) and Rhaphomyia are nectar feeders and can be caught when sweeping the vegetation. They are not associated with water. The larval stages of Empididae are usually found in soil, decaying wood, leaf litter and mosses, but some are aquatic. All are predacious. The larvae of Hemerodromia and Clinocera develop in the hyporheic zone of fast flowing streams at more than 10 cm under the water surface. They wander around in coarse substrate with a low content of silt, sometimes under loose rocks, in search of chironomid and simuliid lavae to prey on (Vaillant 1951, 1967; Vaillant and Gagneur 1998). The Hemerodromiinae take oxygen directly from the water while most other higher flies have to take oxygen out of the air with their spiracles. REGIONAL TAXA To our knowledge, there are no studies of aquatic empidids of Malaysia and knowledge of the Southeast Asian fauna on the whole is very poor (Smith 1975). Five species of Hilara have been described recently from Thailand (Grootaert and Verapong 2001) and eight Hemerodromiinae are known from Xishuangbanna (Yunnan, South China) (Grootaert et al. 2000), but it is doubtful that these species occur here. Six Hemerodromiinae are known also from Java: one Clinocera (Kowarzia), two Chelipoda and three Hemerodromia species. An intense search during three days in Endau-Rompin Park revealed the presence of only two Hemerodromia species (Grootaert in litt.). No larvae have been described yet from Southeast Asia and many adults are yet to be described as well. Since many species are indicators of environmental conditions and exhibit a zonation in the streams, research on their ecology is recommended in terms of site quality assessment studies. KEYS TO GENERA Identification of adults is done under a stereoscopic microscope. The genera can be recognized on the basis of the wing venation and the shape of the antennae. To identify to species level, the male secondary sexual characters are very characteristic as well as the structure of the male genitalia. Therefore preparation (dissection,

4 808 Freshwater Invertebrates of the Malaysian Region macerating in potassium hydroxide, mounting on slides) of the genitalia is necessary. The females are also characteristic but few authors have described them. So few keys allow the identification of females. The following key is limited to the empidid species associated with water bodies or marshes, except otherwise mentioned. Key to genera occurring along streams and ponds in Southeast Asia: adults 1. Fore legs raptorial: femora thickened with ventral spines (Fig. 2B) Fore legs not raptorial: femora of the usual shape Wing with discal cell (cf. Fig. 2A)... Chelifera - Wing without discal cell (Fig. 2B); running on rocks and boulders... Hemerodromia 3. Fourth vein not forked (nectar feeder not really associated with water)... Rhamphomyia - Wing with fourth vein forked (Fig. 2A,C) No proboscis, but thick lips (labellae) like in dolichopodids... Clinocera - Proboscis with a long pointed labrum (Fig. 2A,C) Proboscis long and slender (much longer than height of head, Fig. 2C); angle of fork on fourth vein acute, upper branch straight (Fig. 2C); 6 th vein not reaching the wing border (Fig. 2C); nectar feeder not associated with water... Empis - Proboscis short and stout (generally shorter than head is high; Fig 2A): angle of fork wider with upper branch elongated S-shaped (Fig. 2A); swarming above water bodies... Hilara Key to genera occurring along streams and ponds in Southeast Asia: larvae The larvae are very small, 3 6 mm when fully grown and have a whitish to yellowish colour. Hemerodromia larvae are dark pigmented. 1. Larvae amphipneustic with distinct posterior spiracles, without ventral pseudopods; terminal abdominal segment posteriorly rounded with a short tooth-like extension (Fig 3A arrow) Hilara, Empis, Rhamphomyia - Larvae apneustic (without respiratory spiracles), with conspicuous ventral abdominal pseudopods (Fig. 3B,C) Terminal abdominal segment posteriorly at most with small tubercles, each with a few long setae (otherwise the larva looks like the one of Hemerodromia)... Chelifera - Terminal abdominal segment with prominent posterior processes Terminal abdominal segment with a pair of large ventral pseudopods (Fig. 3C). Pupa (Fig. 3E) with long, thin lateral processes... Hemerodromia - Terminal abdominal segment with both dorsal and apical processes and a shorter ventral pseudopod (Fig. 3B)... Clinocera

5 Insecta: Diptera, Empididae 809 Figure 3. A Hilara larva (arrow indicates position of posterior spiracles and tooth-like projection); B Clinocera larva; C. Hemerodromia larva (p: pseudopods); D Clinocera head; E Hemerodromia pupa. Scale 1 mm. (Source: after Niesiolowski) REFERENCES Grootaert P. (1994) Biodiversity in insects Speciation and behaviour in Diptera. Proceedings Congress on Biodiversity. In Hoffman M. and Van der Veken P. (Eds) Proceedings of the Symposium on Biodiversity: Study, Exploration, Conservation, Ghent, 18 November Pp Grootaert P., Ding Yang and Saigusa T. (2000) Empididae (Diptera: Empidoidea) from Xishuangbanna, Yunnan (I): Hemerodromiinae. Bulletin van het Koninklijk Belgisch Instituut voor Natuurwetenschappen. Serie Entomologie 66: Grootaert P. and Kiatsoonthorn V. (2001) First records of the dance fly genus Hilara in Thailand with the description of five new species. Natural History Bulletin of the Siam Society 49: Smith K.V.G. (1969) The Empididae of southern Africa. Annals of the Natal Museum 19: Smith K.V.G. (1975) Superfamily Empidoidea Family Empididae (Empidae, Hybotidae). In Delfinado M.D. and Hardy D.E. (Eds) A Catalog of the Diptera of the Oriental Region. Volume II. Suborder Brachycera through division Aschiza, suborder Cyclorrhapha. University Press of Hawaii, Honolulu. Pp Vaillant F. (1951) Un empidide destructeur de simulies. Bulletin de le Société Zoologique de France 76: Vaillant F. (1967) La répartition des Wiedemannia dans les cours d eau et leur utilisation comme indicateurs de zones écologiques. Annales de Limnologie 3: Vaillant F. and Gagneur J. (1998) The Diptera Empididae Hemerodromiinae from western Algeria and the Middle Atlas of Morocco. Annales de la Société Entomologique (N.S.) 34:

Slide 1. Slide 1. Next. 5:30:08 AM

Slide 1. Slide 1. Next.  5:30:08 AM Slide 1 Slide 1 http://www3.utep.edu/leb/mosquito/larvslide1.htm10/27/2004 5:30:08 AM Slide 1 Slide 2 Recognition that the specimens are mosquito larvae is a prerequisite to identification of the genera.

More information

TOMINEMOURA, A NEW STONEFLY GENUS FROM SABAH, EAST MALAYSIA (PLECOPTERA: NEMOURIDAE)

TOMINEMOURA, A NEW STONEFLY GENUS FROM SABAH, EAST MALAYSIA (PLECOPTERA: NEMOURIDAE) TOMINEMOURA, A NEW STONEFLY GENUS FROM SABAH, EAST MALAYSIA (PLECOPTERA: NEMOURIDAE) Ignac Sivec 1 and Bill P. Stark 2 1 Slovenian Museum of Natural History, Prešernova 20, P.O. Box 290, SLO-1001 Ljubljana,

More information

Instructions: CLASSIFY ANIMALS AS BLOOD FEEDERS OR NON-BLOODFEEDERS 1. Take out the ADULT CARDS, and place them on the correct boxes 2.

Instructions: CLASSIFY ANIMALS AS BLOOD FEEDERS OR NON-BLOODFEEDERS 1. Take out the ADULT CARDS, and place them on the correct boxes 2. Instructions: CLASSIFY ANIMALS AS BLOOD FEEDERS OR NON-BLOODFEEDERS 1. Take out the ADULT CARDS, and place them on the correct boxes 2. March the LARVA CARDS with the correct adult cards 3. March the HABITAT

More information

Contribution to the morphology of the third-instar larvae of Laccophilus poecilus KLUG (Coleoptera: Dytiscidae)

Contribution to the morphology of the third-instar larvae of Laccophilus poecilus KLUG (Coleoptera: Dytiscidae) Genus Vol. 15(1): 31-36 Wroc³aw, 30 III 2004 Contribution to the morphology of the third-instar larvae of Laccophilus poecilus KLUG (Coleoptera: Dytiscidae) EUGENIUSZ BIESIADKA and IWONA KA KAŹMIERSKA

More information

Identifying Some Pest and Beneficial Insects on Your Sticky Cards

Identifying Some Pest and Beneficial Insects on Your Sticky Cards Identifying Some Pest and Beneficial Insects on Your Sticky Cards Leanne Pundt University of Connecticut Department of Extension An Equal Opportunity Employer and Program Provider Scouting Methods Sticky

More information

Taxonomy. An Introduction to the Taxonomy and Ecology of EPT Families

Taxonomy. An Introduction to the Taxonomy and Ecology of EPT Families An Introduction to the Taxonomy and Ecology of EPT Families Prepared by Dave Penrose North Carolina State University Biological and Agricultural Engineering Department Soil & Water Environmental Technology

More information

Stactobia gozmanyi sp. n.

Stactobia gozmanyi sp. n. Acta Zoologica Academiae Scientiarum Hungaricae 53 (Suppl. 1), pp. 225 229, 2007 A NEW SPECIES OF THE GENUS STACTOBIA MCLACHLAN FROM ETHIOPIA (TRICHOPTERA: HYDROPTILIDAE) MEY, W. Museum für Naturkunde,

More information

Identifying Some Pest and Beneficial Insects on Your Sticky Cards Leanne Pundt UConn Extension

Identifying Some Pest and Beneficial Insects on Your Sticky Cards Leanne Pundt UConn Extension Identifying Some Pest and Beneficial Insects on Your Sticky Cards Leanne Pundt UConn Extension www.ipm.uconn.edu An Equal Opportunity Employer and Program Provider Scouting Use yellow sticky cards to trap

More information

Noldo kaprusii n. sp., a new species from Crimea (Protura: Nipponentomidae)

Noldo kaprusii n. sp., a new species from Crimea (Protura: Nipponentomidae) Genus Vol. 17(2): 151-15 Wroc³aw, 30 VI 2006 Noldo kaprusii n. sp., a new species from Crimea (Protura: Nipponentomidae) JULIA SHRUBOVYCH 1 & ANDRZEJ SZEPTYCKI 2 1 State Museum of Natural History, Ukrainian

More information

Checklist of the Collembola: Key to common surface dwelling species of Collembola from the UK

Checklist of the Collembola: Key to common surface dwelling species of Collembola from the UK http://www.collembola.org/key/fkuk.pdf - Last updated on 2009.03.14 by Frans Janssens Checklist of the Collembola: Key to common surface dwelling species of Collembola from the UK This key is still under

More information

Live Foods from the Wild Part IV Types of Wild Foods

Live Foods from the Wild Part IV Types of Wild Foods Live Foods from the Wild Part IV Types of Wild Foods A presentation for The Angelfish Society March 20, 2010 by Tamar Stephens 1 About this presentation This is Part IV in a series of presentations on

More information

SALAMANDERS, MUDPUPPY AND NEWT OF WISCONSIN

SALAMANDERS, MUDPUPPY AND NEWT OF WISCONSIN SALAMANDERS, MUDPUPPY AND NEWT OF WISCONSIN Information from Wisconsin DNR Spotted Salamander (Ambystoma maculatum) Size: 4 to 7 in. Description: The dorsal (back) side of the body is generally black or

More information

CORE CURRICULUM CONTENT STANDARDS. Language Arts 1(12), 2(2,5) Science 3(1-3), 4(1,2), 5(1,4,6), 6(1,8) Arts 3(1,2), 5(1)

CORE CURRICULUM CONTENT STANDARDS. Language Arts 1(12), 2(2,5) Science 3(1-3), 4(1,2), 5(1,4,6), 6(1,8) Arts 3(1,2), 5(1) Lesson 11 Water Creatures How do insects survive under water? How are insects indicators of the health of a stream? Farmington River Watershed Education Curriculum: Elementary GOAL OBJECTIVES MATERIALS

More information

Key to the Species of Anamixis

Key to the Species of Anamixis Key to the Species of Anamixis (Except A. stebbingi*) 1. Article 2 of gnathopod 2 with 3 or more long setae on anterior margin (1)...2 1. Article 2 of gnathopod 2 lacking long setae on anterior margin...3

More information

RHOPALOPSOLE ALOBATA (PLECOPTERA: LEUCTRIDAE), A NEW STONEFLY SPECIES FROM VIETNAM

RHOPALOPSOLE ALOBATA (PLECOPTERA: LEUCTRIDAE), A NEW STONEFLY SPECIES FROM VIETNAM RHOPALOPSOLE ALOBATA (PLECOPTERA: LEUCTRIDAE), A NEW STONEFLY SPECIES FROM VIETNAM Audrey B. Harrison 1 & Bill P. Stark 2 1 Department of Biology, Box 4045, Mississippi College, Clinton, Mississippi, 39058,

More information

males, but they are less susceptible to infectious diseases and

males, but they are less susceptible to infectious diseases and General information- In nature, this specie's males have shorter fins that those we usually see in shops,these are called by the Thai "Plakad" (meaning- "Fighter"), and are used in Thailand for show-fights

More information

BENSON PARK POND FISH SPECIES

BENSON PARK POND FISH SPECIES BENSON PARK POND FISH SPECIES Bluegill (Lepomis macrochirus) From the Greek, lepomis means scaled gill cover and macrochirus means large hand, in reference to its body shape and size. Average adult size

More information

Four new species of the spider genus Pinelema (Araneae, Telemidae) from caves in South China

Four new species of the spider genus Pinelema (Araneae, Telemidae) from caves in South China ISSN 2336-9744 (online) ISSN 2337-0173 (print) The journal is available on line at www.biotaxa.org/em Research Article https://zoobank.org/urn:lsid:zoobank.org:pub:a978b551-dcd5-46a0-8cfb-2e1927a24c8d

More information

Blood Sucking Insects

Blood Sucking Insects An Introduction ti to Study of Blood Sucking Insects Part I Dipterans Comprehensive Workshop kh on Medical Entomology 23 February 2012 Prepared by MB Ghavami Associated Professor of Medical Entomology

More information

THEJ FIRST ZOEA OF PORCELLANA. By W K. BROOKS and E. B. WILSON. With Plate* VI and VII.

THEJ FIRST ZOEA OF PORCELLANA. By W K. BROOKS and E. B. WILSON. With Plate* VI and VII. 7 "83?* {Jn-fJatoj 11 "T THEJ FIRST ZOEA OF PORCELLANA. By W K BROOKS and E. B. WILSON. With Plate* VI and VII. 7z O THE FIRST ZOEA OF PORCELLANA. By W K. BROOKS and E. B. WILSON. With Plates VI and VII.

More information

FAO SPECIES IDENTIFICATION SHEETS SYNODONTIDAE. Lizardfishes

FAO SPECIES IDENTIFICATION SHEETS SYNODONTIDAE. Lizardfishes click for previous page SYNOD 1474 FAO SPECIES IDENTIFICATION SHEETS FISHING AREAS 57,71 (E Ind. Ocean) (W Cent. Pacific) SYNODONTIDAE Lizardfishes Body elongate, usually cylindrical and with adipose fin.

More information

[USP5655] [USP5650] [USP5653] [USP5649] [USP5654] [USP5648] 121 [USP5652] grazers 33. predator grazers 124 [USP5647]

[USP5655] [USP5650] [USP5653] [USP5649] [USP5654] [USP5648] 121 [USP5652] grazers 33. predator grazers 124 [USP5647] Appendix Author(s): Source: A Rapid Biodiversity Assessment of the Nakauvadra Range, Ra Province, Fiji:-. 0. Published By: Conservation International URL: http://www.bioone.org/doi/full/0.896/0.07.00 BioOne

More information

Received 12 December 2014; received in revised form 29 December 2014; accepted 30 December 2014

Received 12 December 2014; received in revised form 29 December 2014; accepted 30 December 2014 Tropical Biomedicine 32(3): 504 510 (2015) Description of male, pupa and larva of Simulium (Asiosimulium) wanchaii (Diptera: Simuliidae) from Thailand, with keys to identify four species of the subgenus

More information

Dragonhu Drago nt nh er unter Dragonflies of N. Va. Kevin Munroe, 2012 Dragonflies of N. Va. - text and photos by K. T.

Dragonhu Drago nt nh er unter Dragonflies of N. Va. Kevin Munroe, 2012 Dragonflies of N. Va. - text and photos by K. T. Dragonhunter Dragonflies of N. Va. Kevin Munroe, 2012 Dragonflies of N. Va. - text and photos by K. T. Munroe, 2010 Waiting for dinner to fly by One of Northern Virginia s most impressive insects. Flight

More information

Four Undescribed Species of Waterstriders (Hemip.-Gerridae)

Four Undescribed Species of Waterstriders (Hemip.-Gerridae) The Ohio State University Knowledge Bank kb.osu.edu Ohio Journal of Science (Ohio Academy of Science) Ohio Journal of Science: Volume 25, Issue 1 (January, 1925) 1925-01 Four Undescribed Species of Waterstriders

More information

CHAPTER 4. EPHEMEROPTERA (Mayflies)

CHAPTER 4. EPHEMEROPTERA (Mayflies) Guide to Aquatic Invertebrate Families of Mongolia 2009 CHAPTER 4 (Mayflies) Draft June 17, 2009 Chapter 4 45 4 ORDER Mayflies Mayfly larvae are found in a variety of locations including lakes, wetlands,

More information

The immature stages of two Caladomyia Säwedall, 1981 species, from São Paulo State, Brazil (Chironomidae, Chironominae, Tanytarsini)

The immature stages of two Caladomyia Säwedall, 1981 species, from São Paulo State, Brazil (Chironomidae, Chironominae, Tanytarsini) Revista Brasileira de Entomologia 7(): 97-602 The immature stages of two Caladomyia Säwedall 97.XII.200 The immature stages of two Caladomyia Säwedall, 98 species, from São Paulo State, Brazil (Chironomidae,

More information

THE RAFFLES BULLETIN OF ZOOLOGY 2009

THE RAFFLES BULLETIN OF ZOOLOGY 2009 THE RAFFLES BULLETIN OF ZOOLOGY 2009 57(2): 255 277 Date of Publication: 31 Aug.2009 National University of Singapore DIVERSITY OF CHELIPODA MACQUART, 1823 (DIPTERA: EMPIDIDAE: HEMERODROMIINAE) IN NORTHERN

More information

CHAPTER 2 DIVERSITY OF HONEYBEES IN TAMIL NADU, INDIA

CHAPTER 2 DIVERSITY OF HONEYBEES IN TAMIL NADU, INDIA CHAPTER 2 DIVERSITY OF HONEYBEES IN TAMIL NADU, INDIA 2.1 INTRODUCTION Tamil Nadu is the extreme southern state of India, bounded by Andhra Pradesh on the North, Karnataka and Kerala on the west, Indian

More information

Chapter 30 Nonvertebrate Chordates, Fishes, and Amphibians Name

Chapter 30 Nonvertebrate Chordates, Fishes, and Amphibians Name Chapter 30 Nonvertebrate Chordates, Fishes, and Amphibians Name Lab Dissecting a Perch Background Information Fish are the largest group of vertebrates found in fresh and salt water. In fact, over 25,000

More information

REICHENBACHI A STAATLICHES MUSEUM FUR TIERKUNDE IN DRESDE N

REICHENBACHI A STAATLICHES MUSEUM FUR TIERKUNDE IN DRESDE N REICHENBACHI A STAATLICHES MUSEUM FUR TIERKUNDE IN DRESDE N Bd. 13 Ausgegeben : 17. Oktober 197 0 Nr. 14 A new Phalangid from Cuba, Trinimontius darlingtoni gen. n., sp. n. (Opilionidea, Cosmetidae ) with

More information

Unit 18.2: Mollusks and Annelids

Unit 18.2: Mollusks and Annelids Unit 18.2: Mollusks and Annelids Lesson Objectives Describe invertebrates in the phylum Mollusca. Summarize the characteristics of annelids. Vocabulary Annelida deposit feeder gills heart mantle Mollusca

More information

Chapter XII Order Diptera

Chapter XII Order Diptera Chapter XII Order Diptera (Two-winged or true flies ) (Flies, Mosquitoes, Midges) (Williams & Feltmate, 1992) Superphylum Arthropoda (jointed-legged metazoan animals [Gr, arthron = joint; pous = foot])

More information

Fly Fishing for Trout in Southeast Minnesota Streams

Fly Fishing for Trout in Southeast Minnesota Streams Fly Fishing for Trout in Southeast Minnesota Streams Fly Fishing for the First Time? Equipment Basics Mid-weight fly rod, 5 or 6 weight, with fly line to match Reel Leader Extra tippet Flies Glasses or

More information

FAO SPECIES IDENTIFICATION SHEETS. FAMILY: SIGANIDAE FISHING AREA 51 (W. Indian Ocean) Siganis rivulatus Forsskål, 1775

FAO SPECIES IDENTIFICATION SHEETS. FAMILY: SIGANIDAE FISHING AREA 51 (W. Indian Ocean) Siganis rivulatus Forsskål, 1775 click for previous page SIGAN Sigan 1 1983 FAO SPECIES IDENTIFICATION SHEETS FAMILY: SIGANIDAE FISHING AREA 51 (W. Indian Ocean) Siganis rivulatus Forsskål, 1775 OTHER SCIENTIFIC NAMES STILL IN USE : Teuthis

More information

Contents. A Field Guide to the Dragonflies and Damselflies of Sri Lanka 13

Contents. A Field Guide to the Dragonflies and Damselflies of Sri Lanka 13 Contents Foreword 08 Preface 09 Acknowledgements 11 1. A Brief Introduction to Odonata (Dragonflies and Damselflies) 15 Etymology 15 Basic Morphology 16 Zygoptera or Anisoptera 17 Differentiation between

More information

A NEW SPECIES OF NEVSKYELLA OSSIANNILSSON, 1954 (HOMOPTERA, APHIDIDAE)

A NEW SPECIES OF NEVSKYELLA OSSIANNILSSON, 1954 (HOMOPTERA, APHIDIDAE) A NEW SPECIES OF NEVSKYELLA OSSIANNILSSON, 1954 (HOMOPTERA, APHIDIDAE) by D. H I L L E RIS L A M B E R S Bladluisonderzoek T.N.O., Bennekom, Netherlands and R. V A N D E N B O S C H University of California,

More information

Question # The question The answer Bugs to use Fact or ID sheets Magnify? needed needed Difficulty

Question # The question The answer Bugs to use Fact or ID sheets Magnify? needed needed Difficulty A B Here is a mount of a common insect s life cycle. a. Identify this insect. b. Review the life cycle mount and determine if it is complete or incomplete metamorphosis. c. List the stages in order of

More information

www.montessorinature.com/printables How To Use Montessori Nomenclature 3 -Part Cards Montessori Three-Part Cards are designed for children to learn and process the information on the cards. The Montessori

More information

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi MARINE BIORESOURCES FORMS DATA ENTRY: Form- 1(general ) (please answer only relevant fields;add additional fields

More information

Checklist of the tiger beetle genus Calochroa Hope, 1838 from Laos and a description of a new species (Coleoptera: Cicindelidae)

Checklist of the tiger beetle genus Calochroa Hope, 1838 from Laos and a description of a new species (Coleoptera: Cicindelidae) Entomologica Basiliensia et Collectionis Frey 34 47 51 2013 ISSN 1661 8041 Checklist of the tiger beetle genus Calochroa Hope, 1838 from Laos and a description of a new species (Coleoptera: Cicindelidae)

More information

TAKE HOME BOOKLET KANANASKIS COUNTRY WETLANDS: PLUNGE IN! POND LIFE DISCOVERY PACK A LOOK AT POND LIFE

TAKE HOME BOOKLET KANANASKIS COUNTRY WETLANDS: PLUNGE IN! POND LIFE DISCOVERY PACK A LOOK AT POND LIFE KANANASKIS COUNTRY POND LIFE DISCOVERY PACK TAKE HOME BOOKLET WETLANDS: PLUNGE IN! A LOOK AT POND LIFE October 1999 Pub. No. I/781 ISBN 0-7785-0880-3 (printed edition) ISBN 0-7785-0881-1 (on-line edition)

More information

Life Cycles: Egg, Nymph, Adult!

Life Cycles: Egg, Nymph, Adult! Life Cycles: Egg, Nymph, Adult! National Mississippi River Museum & Aquarium Environmental Education Curriculum Target Grades: K 5 Key Words: Subject Area: Duration: Life Cycle, Metamorphosis, Native Species,

More information

Stoneflies Order Plecoptera True Alaskans?

Stoneflies Order Plecoptera True Alaskans? Stoneflies Order Plecoptera True Alaskans? Perhaps, while snowshoeing early in the spring in Alaska, you ve noticed an insect on the surface of the snow. On closer examination, you realize that the bug

More information

Salamanders of Idaho. Ambystomatidae Mole Salamanders Long-toed Salamander Barred Tiger Salamander. Dicamptodontidae Idaho Giant Salamander

Salamanders of Idaho. Ambystomatidae Mole Salamanders Long-toed Salamander Barred Tiger Salamander. Dicamptodontidae Idaho Giant Salamander Salamanders of Idaho Ambystomatidae Mole Salamanders Long-toed Salamander Barred Tiger Salamander Dicamptodontidae Idaho Giant Salamander Plethodontidae Lungless Salamanders Coeur d Alene Salamander Salamandridae

More information

RIVER CONONISH INVERTEBRATE SURVEY Dr Kjersti Birkeland

RIVER CONONISH INVERTEBRATE SURVEY Dr Kjersti Birkeland RIVER CONONISH INVERTEBRATE SURVEY 26 Dr Kjersti Birkeland Background The Cononish, the upper reaches of the River Dochart, should be very important for spring salmon but surveys by the Tay District Salmon

More information

THE GENUS PHOTIS (CRUSTACEA: AMPHIPODA) FROM THE TEXAS COAST WITH THE DESCRIPTION OF A NEW SPECIES, PHOTIS MELANICUS

THE GENUS PHOTIS (CRUSTACEA: AMPHIPODA) FROM THE TEXAS COAST WITH THE DESCRIPTION OF A NEW SPECIES, PHOTIS MELANICUS THE GENUS PHOTIS (CRUSTACEA: AMPHIPODA) FROM THE TEXAS COAST WITH THE DESCRIPTION OF A NEW SPECIES, PHOTIS MELANICUS L. D. McKinney Texas A&M University at Galveston, Moody College of Marine Technology,

More information

Biology & Ecology of large branchiopods

Biology & Ecology of large branchiopods Biology & Ecology of large branchiopods Carol W. Witham, VernalPools.Org Talk Overview Classification of large branchiopods Anatomy of a fairy shrimp Biology of a fairy shrimp Ecology of large branchiopods

More information

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi For office use: MARINE BIORESOURCES FORMS DATA ENTRY: Form- 1(general ) Ref. No.: (please answer only relevant

More information

IPM Fun with Insects, Weeds and the Environment. Lesson #2 Insect IPM. The New York State Integrated Pest Management Program

IPM Fun with Insects, Weeds and the Environment. Lesson #2 Insect IPM. The New York State Integrated Pest Management Program IPM Fun with Insects, Weeds and the Environment Lesson #2 Insect IPM The New York State Integrated Pest Management Program Learn how you can use the 6 steps of integrated pest management to help protect

More information

Mollusks Soft-bodied Invertebrates

Mollusks Soft-bodied Invertebrates Mollusks Soft-bodied Invertebrates Phylum Mollusca Very diverse - more species of molluscs than any other group in the ocean. Phylum includes: Bivalves (2 shells); ex. Clam Gastropods (1 shell, coiled);

More information

Stream Insects of the Pacific Northwest

Stream Insects of the Pacific Northwest Stream Insects of the Pacific Northwest Patrick Edwards 2008 Acknowledgments - Jeff Adams, Rick Hafele, Dr. Ian Waite and Dr. Robert W. Wisseman for sharing their knowledge of aquatic insects. Dr. Trygve

More information

Biology. Slide 1 of 43. End Show. Copyright Pearson Prentice Hall

Biology. Slide 1 of 43. End Show. Copyright Pearson Prentice Hall Biology 1 of 43 2 of 43 What Is a Mollusk? What are the defining features of mollusks? 3 of 43 What Is a Mollusk? What Is a Mollusk? Mollusks are soft-bodied animals that usually have an internal or external

More information

Life in freshwater stream - Mayfly Nymphs

Life in freshwater stream - Mayfly Nymphs Life in freshwater stream - Mayfly Nymphs Group members F.5 Chu Tsz Fung F.5 Wong Wing Kwan F.4 Ho Sum Yat F.4 Lee Kwun Chak F.4 Lau Wai Sum Report on mayfly nymphs, Page 1 Content page 1 Abstract 1.1

More information

Wave, Tide or Ride, Follow the Guide

Wave, Tide or Ride, Follow the Guide Wave, Tide or Ride, Follow the Guide KEY TERMS Catch pool: A landing pool located at the bottom of a water slide. Current: The continuous movement of water. Dispatcher: The lifeguard at the top of a water

More information

Dead Perch Parts. ACADEMIC STANDARDS: 4 th Grade B. Know that living things are made up of parts that have specific functions.

Dead Perch Parts. ACADEMIC STANDARDS: 4 th Grade B. Know that living things are made up of parts that have specific functions. Dead Perch Parts Fish Anatomy Adapted from: An original Creek Connections activity created from the Fish Anatomy model. Grade Level: Intermediate or advanced Duration: 30 minutes Setting: classroom Summary:

More information

NOTES AND NEWS A NORTHERN INDIAN OCEAN RECORD OF A BOPYRID, PARAPENAEON JAPONICA (THIELEMANN, 1910) PARASITIC ON SPECIES OF PENAEID PRAWNS (PENAEUS)

NOTES AND NEWS A NORTHERN INDIAN OCEAN RECORD OF A BOPYRID, PARAPENAEON JAPONICA (THIELEMANN, 1910) PARASITIC ON SPECIES OF PENAEID PRAWNS (PENAEUS) Pakistan Journal of Marine Sciences, Vol.3(2), 171-175, 1994. NOTES AND NEWS A NORTHERN INDIAN OCEAN RECORD OF A BOPYRID, PARAPENAEON JAPONICA (THIELEMANN, 1910) PARASITIC ON SPECIES OF PENAEID PRAWNS

More information

TWO NEW SPECIES OF THE GENUS DYNAMENELLA FROM THE NORHTERN ARABIAN SEA (ISOPODA)

TWO NEW SPECIES OF THE GENUS DYNAMENELLA FROM THE NORHTERN ARABIAN SEA (ISOPODA) TWO NEW SPECIES OF THE GENUS DYNAMENELLA FROM THE NORHTERN ARABIAN SEA (ISOPODA) BY WAQUAR JAVED and ROSHAN AHMED Department of Zoology, University of Karachi, Karachi-32 RESUME Deux nouvelles espèces,

More information

West African Lungfish A living fossil s biological and behavioral adaptations

West African Lungfish A living fossil s biological and behavioral adaptations VideoMedia Spotlight West African Lungfish A living fossil s biological and behavioral adaptations For the complete video with media resources, visit: http://education.nationalgeographic.org/media/west-african-lungfish/

More information

Pass the Water Fleas, Please

Pass the Water Fleas, Please Tracy Armand Grolee Elementary Opelousas, La. Grade Level Middle School (6-8) Duration Four to six weeks Subject Area Inquiry Life Science Materials List Daphnia spp. Computer with internet access and

More information

Cylicocyclus spp. Walls of the Buccal Capsule:

Cylicocyclus spp. Walls of the Buccal Capsule: Figure 8a Cylicocyclus ashworthi Figure 9a Cylicocyclus auriculatus Figure 0a Cylicocyclus brevicapsulatus Figure a Cylicocyclus elongatus Figure 3a Cylicocyclus insigne Figure 4a Cylicocyclus radiatus

More information

An Annotated and Illustrated Key to Multistage Larvae of Ohio Salamanders

An Annotated and Illustrated Key to Multistage Larvae of Ohio Salamanders The Ohio State University Knowledge Bank kb.osu.edu Ohio Journal of Science (Ohio Academy of Science) Ohio Journal of Science: Volume 64, Issue 4 (July, 1964) 1964-07 An Annotated and Illustrated Key to

More information

The Animal Kingdom. Animal Kingdom. Characteristics of All Animals. Major Characteristics Used To Classify Animals

The Animal Kingdom. Animal Kingdom. Characteristics of All Animals. Major Characteristics Used To Classify Animals The Animal Kingdom Animal Kingdom Phylums: 1. Sponges 2. Cnidaria Jelly Fish, Hydra 3. Flatworms Flukes, Tapeworms 4. Roundworms- Hookworms 5. Segmented Worms- Earthworms 6. Rotifera- Rotifers 7. Mullusca

More information

Two new species of the genus C y a n o se sia G o r b u n o v &

Two new species of the genus C y a n o se sia G o r b u n o v & N ach r. en to m o l. V er. A p o llo, S u p p l. 1 7 : 4 5 7-4 6 4 (1 9 9 8 ) 457 Two new species of the genus C y a n o se sia G o r b u n o v & A rita, 1995 (Lepidoptera, Sesiidae) from the Oriental

More information

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi MARINE BIORESOURCES FORMS DATA ENTRY: Form- 1(general ) Ref. No.: (please answer only relevant fields;add additional

More information

Tips for Identifying Common Fish Species in the Bush River

Tips for Identifying Common Fish Species in the Bush River Tips for Identifying Common Fish Species in the Bush River Juvenile Largemouth bass Spot Threadfin shad Juvenile White perch The Herrings (Family Clupeidae) Alewife American shad Blueback herring Atlantic

More information

A new Neotropical species of Tanytarsus van der Wulp, 1874 (Diptera: Chironomidae), with an unusual anal process

A new Neotropical species of Tanytarsus van der Wulp, 1874 (Diptera: Chironomidae), with an unusual anal process Zootaxa 1654: 61 67 (2007) www.mapress.com/zootaxa/ Copyright 2007 Magnolia Press ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) A new Neotropical species of Tanytarsus van der

More information

ZOOPLANKTON. Zooplankton: 2. Crustaceans Copepods. Diverse -- protozoans and others

ZOOPLANKTON. Zooplankton: 2. Crustaceans Copepods. Diverse -- protozoans and others ZOOPLANKTON Diverse -- protozoans and others Nutrition modes: Herbivores (graze on phytoplankton) Carnivores (predators) Radiolaria Foraminifera Zooplankton: 1. Protozoans: Foraminifera and Radiolarians

More information

Lesson 2-4: Nechako White Sturgeon Habitat and Humans

Lesson 2-4: Nechako White Sturgeon Habitat and Humans 2-27 Lesson 2-4: Nechako White Sturgeon Habitat and Humans Time of Lesson: 1.5 hours Instruction Objectives: Students understand the basic habitat needs of Nechako white sturgeon. Students can identify

More information

Title. Author(s)Takagi, Sadao; Howell, James O. Issue Date Doc URL. Type. File Information (HOMOPTERA : COCCOIDEA)

Title. Author(s)Takagi, Sadao; Howell, James O. Issue Date Doc URL. Type. File Information (HOMOPTERA : COCCOIDEA) Title THE GENUS QUERNASPIS, A POSSIBLE ASIO-AMERICAN ELEME (HOMOPTERA : COCCOIDEA) Author(s)Takagi, Sadao; Howell, James O. Insecta matsumurana. New series : journal of the Fac Citation31-59 Issue Date

More information

COMMON FRESHWATER PLANKTONIC ANIMALS. By J.D. Green* INTRODUCTION

COMMON FRESHWATER PLANKTONIC ANIMALS. By J.D. Green* INTRODUCTION 13 COMMON FRESHWATER PLANKTONIC ANIMALS By J.D. Green* INTRODUCTION Although the occurrence of zooplankton in Auckland lakes and reservoirs has been discussed by Bayly (1962), no elementary descriptions

More information

Aquatic Animal Diversity Background

Aquatic Animal Diversity Background Aquatic Animal Diversity Background Background information for The Water Pixies: The Life Cycle of Aquatic Animals and Aquatic Animal Adaptations. Animals have unique strategies for development that also

More information

Mississippi River Life

Mississippi River Life Mississippi River Life National Mississippi River Museum and Aquarium Conservation Education Curriculum Target Grades: Pre K 4 th grade Key Words: Habitat, diversity Subject Areas: science, biology Duration:

More information

Endangered Crab Found In West Africa s Shrinking Forests

Endangered Crab Found In West Africa s Shrinking Forests Endangered Crab Found In West Africa s Shrinking Forests Neil Cumberlidge and Piotr Naskrecki We were camped in West Africa, on the westernmost fringe of the Upper Guinea forest, a vast area stretching

More information

Internal Anatomy of Fish

Internal Anatomy of Fish Internal Anatomy of Fish The Systems of a Fish Skeletal System Muscular System Respiratory System Digestive System Circulatory System Nervous System Reproductive System Special Organs Skeletal System

More information

Chapter- 9. Multiple Choice Questions

Chapter- 9. Multiple Choice Questions Chapter- 9 Multiple Choice Questions 1. The low-latitude climates: a. are located along the equator b. are located between the tropics of cancer and capricorn c. do not include the ITCZ d. are restricted

More information

How to Hide in the Ocean

How to Hide in the Ocean SEA How to Hide in the Ocean

More information

Choroterpes (Choroterpes) prati n.sp., a new species of mayfly from North-East of Spain (Ephemeroptera : Leptophlebiidae)

Choroterpes (Choroterpes) prati n.sp., a new species of mayfly from North-East of Spain (Ephemeroptera : Leptophlebiidae) Annls Limnol. 32 (4) 1996 : 229-233 Choroterpes (Choroterpes) prati n.sp., a new species of mayfly from North-East of Spain (Ephemeroptera : Leptophlebiidae) M.A. Puig 1 E. Gaino 2 Keywords : Ephemeroptera,

More information

Fantastic Fish. Lesson 10. Leafy Sea Dragon

Fantastic Fish. Lesson 10. Leafy Sea Dragon Fantastic Fish Lesson 10 Fish are cold-blooded vertebrates that live in streams, lakes and oceans. They are abundant in both fresh and salt water. There are over 28,000 known species of fish on the planet.

More information

Life at its Peak of Exuberance: Coral Reefs and Tropical Rainforests

Life at its Peak of Exuberance: Coral Reefs and Tropical Rainforests Life at its Peak of Exuberance: Coral Reefs and Tropical Rainforests Note: These links do not work. Use the links within the outline to access the images in the popup windows. This text is the same as

More information

Levels of the Savannah. Guide Book

Levels of the Savannah. Guide Book Levels of the Savannah Guide Book K-12 Levels of the Savannah What: 60 90 min tour of the Savannah building discussing the niches of the African savannah When: can be done year-round Backpack contents:

More information

SHORT COMMUNICATION ON THE ENDOPARASITIC FAUNA OF SOME PADDY-FIELD FISHES FROM KEDAH, PENINSULAR MALAYSIA

SHORT COMMUNICATION ON THE ENDOPARASITIC FAUNA OF SOME PADDY-FIELD FISHES FROM KEDAH, PENINSULAR MALAYSIA Journal of Bioscience, 19(2), 107 112, 2008 SHORT COMMUNICATION ON THE ENDOPARASITIC FAUNA OF SOME PADDY-FIELD FISHES FROM KEDAH, PENINSULAR MALAYSIA Wahab A Rahman and Mutaqin Bakri School of Biological

More information

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi

NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi NATIONAL BIORESOURCE DEVELOPMENT BOARD Dept. of Biotechnology Government of India, New Delhi For office use: MARINE BIORESOURCES FORMS DATA ENTRY: Form- 1(general) (please answer only relevant fields;

More information

Article. https://doi.org/ /zootaxa

Article. https://doi.org/ /zootaxa Zootaxa 4227 (4): 554 562 http://www.mapress.com/j/zt/ Copyright 2017 Magnolia Press Article https://doi.org/10.11646/zootaxa.4227.4.5 http://zoobank.org/urn:lsid:zoobank.org:pub:a353bed0-fb5c-4af5-8a44-b7b8f8d7f942

More information

A Stream In A Bucket An introduction to aquatic macroinvertebrates and other stream life.

A Stream In A Bucket An introduction to aquatic macroinvertebrates and other stream life. A Stream In A Bucket An introduction to aquatic macroinvertebrates and other stream life. Safety Notes: Stream water is not sterile. Students should be instructed not to allow any water to enter their

More information

Systematics. A Survey of the Necrophilous Diptera on the Island of Dominica. Jonathan A. Cammack

Systematics. A Survey of the Necrophilous Diptera on the Island of Dominica. Jonathan A. Cammack Systematics A Survey of the Necrophilous Diptera on the Island of Dominica Jonathan A. Cammack Department of Entomology Texas A&M University, College Station T, USA 77843 Neotropical Entomology 00(0):000-000

More information

NEW SPECIES OF OLETHREUTINE MOTHS (TORTRICIDAE) FROM TEXAS AND LOUISIANA

NEW SPECIES OF OLETHREUTINE MOTHS (TORTRICIDAE) FROM TEXAS AND LOUISIANA Journal of the Lepidopterists' Society 40(4), 1986, 322-326 NEW SPECIES OF OLETHREUTINE MOTHS (TORTRICIDAE) FROM TEXAS AND LOUISIANA EDW ARD C. KNUDSON 808 Woodstock, Bellaire, Texas 77401 Research Associate,

More information

Notes on Varroa destructor (Acari: Varroidae) parasitic on honeybees in New Zealand

Notes on Varroa destructor (Acari: Varroidae) parasitic on honeybees in New Zealand Systematic & Applied Acarology Special Publications (2000) 5, 9-14 Notes on Varroa destructor (Acari: Varroidae) parasitic on honeybees in New Zealand ZHI-QIANG ZHANG Landcare Research, Private Bag 92170,

More information

[Type the abstract of the document here. The abstract is typically a short summary of the contents of the document. Type the abstract of the document

[Type the abstract of the document here. The abstract is typically a short summary of the contents of the document. Type the abstract of the document [Type the abstract of the document here. The abstract is typically a short summary of the contents of the document. Type the abstract of the document here. The abstract is typically a short summary of

More information

Chapter 7. Marine Animals Without a Backbone

Chapter 7. Marine Animals Without a Backbone Chapter 7 Marine Animals Without a Backbone Molluscs Characteristics of Phylum: More than 200,000 species Name means "soft body" Basic body plan head, muscular foot and visceral mass in most species Mantle-

More information

Long-tailed Weasel. Mustela frenata. Other common names. Introduction. None

Long-tailed Weasel. Mustela frenata. Other common names. Introduction. None Long-tailed Weasel Mustela frenata Other common names None Introduction Like its cousin the short-tailed weasel, the long-tailed weasel develops a white coat in winter to aid with camouflage. This habit

More information

Aquatic Insects of India-A fieldguide

Aquatic Insects of India-A fieldguide 1 Aquatic Insects of India-A fieldguide K.A.Subramanian and K.G.Sivaramakrishnan Ashoka Trust for Research in Ecology and Environment (ATREE) Small Grants Programme 2007 2 Design and layout: R.Shobana

More information

UNIFYING CONCEPTS OF ANIMAL CIRCULATION

UNIFYING CONCEPTS OF ANIMAL CIRCULATION UNIFYING CONCEPTS OF ANIMAL CIRCULATION Every organism must exchange materials with its environment, relying upon diffusion, the spontaneous movement of molecules from an area of higher concentration to

More information

28 2 Groups of Arthropods

28 2 Groups of Arthropods 28 2 Groups of Arthropods 1 of 42 28 2 Groups of Arthropods Arthropods are classified based on the number and structure of their body segments and appendages particularly their mouthparts. The three major

More information

STUDY GUIDE. Physical Features. The Land. Chapter 23, Section 1. Landforms. Rivers. Natural Resources. Terms to Know DRAWING FROM EXPERIENCE

STUDY GUIDE. Physical Features. The Land. Chapter 23, Section 1. Landforms. Rivers. Natural Resources. Terms to Know DRAWING FROM EXPERIENCE For use with textbook pages 569 574. The Land Terms to Know subcontinent A landmass that is large and distinct but joined to a continent (page 569) alluvial plain An area of fertile soil deposited by river

More information

A Survey of the North American Species of Merragata

A Survey of the North American Species of Merragata The Ohio State University Knowledge Bank kb.osu.edu Ohio Journal of Science (Ohio Academy of Science) Ohio Journal of Science: Volume 17, Issue 4 (February, 1917) 1917-02 A Survey of the North American

More information

ABCs & Name the parts!

ABCs & Name the parts! ABCs & Name the parts! Objective: These two activities are used to introduce the students to the variety of marine life living in European waters while learning their ABCs and practicing their letter writing.

More information