Hamamelidaceae (& Altingiaceae)

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1 Hamamelidaceae (& Altingiaceae) Hamamelis study weekend IDS Arboretum Kalmthout, Belgium 1 & 2 February 2014

2 Hamamelidaceae (& Altingiaceae) vegetative key to species in cultivation

3 Morphologically very diverse genera. With number of species in this key: Altingia* [3] Corylopsis [10] Disanthus [1] Distylium [4] Exbucklandia [1] Fortunearia [1] Fothergilla [2] Hamamelis [6] Liquidambar* [5] Loropetalum [1] Parrotia [2] Parrotiopsis [1] Rhodoleia [1] Sinowilsonia [1] Sycoparrotia [1] Sycopsis [1] Trichocladus [2] (Altingiaceae*)

4 key to families Altingiaceae Hamamelidaceae

5 Altingiaceae Altingia gracilipes

6 Altingiaceae bruised lamina with sharp scent Altingia gracilipes

7 Altingiaceae bruised lamina with sharp scent leaf arrangement spirally Altingia gracilipes

8 Hamamelidaceae Loropetalum chinense var. rubrum

9 Hamamelidaceae bruised lamina scentless Loropetalum chinense var. rubrum

10 Hamamelidaceae bruised lamina scentless leaf arrangement two-ranked Loropetalum chinense var. rubrum

11 Altingiaceae bruised lamina with sharp scent leaf arrangement spirally Altingia gracilipes

12 key to groups 2 groups each with following diagnostic characters

13 GROUP A: LAMINA VENATION PINNATE (secondary veins scattered along midvein) Altingia poilanei

14 GROUP A: LAMINA VENATION PINNATE (secondary veins scattered along midvein) Altingia poilanei

15 GROUP B: LAMINA VENATION PALMATE (strong veins predominantly radiating from base) Liquidambar orientalis

16 GROUP B: LAMINA VENATION PALMATE (strong veins predominantly radiating from base) Liquidambar orientalis

17 Hamamelidaceae bruised lamina scentless leaf arrangement two-ranked Loropetalum chinense var. rubrum

18 key to groups 3 groups each with following diagnostic characters

19 GROUP A LAMINA VENATION PALMATE strong veins predominantly radiating from base

20 GROUP A LAMINA VENATION PALMATE strong veins predominantly radiating from base GROUP B LAMINA VENATION PINNATE most secondary veins curving + anastomosing before margin

21 GROUP A LAMINA VENATION PALMATE strong veins predominantly radiating from base GROUP B LAMINA VENATION PINNATE most secondary veins curving + anastomosing before margin GROUP C LAMINA VENATION PINNATE most secondary veins ending at margin

22 GROUP A LAMINA VENATION PALMATE strong veins predominantly radiating from base

23 GROUP A: LAMINA VENATION PALMATE Disanthus cercidifolius

24 GROUP A: LAMINA VENATION PALMATE Disanthus cercidifolius

25 GROUP A: LAMINA VENATION PALMATE Lamina unlobed, papery, apex obtuse to acute. Disanthus cercidifolius

26 GROUP A: LAMINA VENATION PALMATE Lamina unlobed, papery, apex obtuse to acute. Stipules free. Disanthus cercidifolius

27 GROUP A: LAMINA VENATION PALMATE Exbucklandia populnea

28 GROUP A: LAMINA VENATION PALMATE Lamina unlobed(rarely lobed), leathery, apex acuminate to caudate. Exbucklandia populnea

29 GROUP A: LAMINA VENATION PALMATE Lamina unlobed (rarely lobed), leathery, apex acuminate to caudate. Stipules united. Exbucklandia populnea

30 GROUP A: LAMINA VENATION PALMATE Lamina unlobed (rarely lobed), leathery, apex acuminate to caudate. Stipules united. Exbucklandia populnea

31 GROUP A LAMINA VENATION PALMATE strong veins predominantly radiating from base GROUP B LAMINA VENATION PINNATE most secondary veins curving + anastomosing before margin

32 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Distylium racemosum

33 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Distylium racemosum

34 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Trichocladus ellipticus

35 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Shoot, petiole and lamina pubescent with long-armed stellate hairs. Trichocladus ellipticus

36 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Shoot, petiole and lamina pubescent with long-armed stellate hairs. Trichocladus ellipticus

37 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Loropetalum sinense

38 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Shoot, petiole and lamina pubescent with long-armed stellate hairs. Loropetalum sinense

39 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Shoot, petiole and lamina pubescent with long-armed stellate hairs. Loropetalum sinense

40 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Shoot, petiole and lamina glabrous or pubescent with scale-like to short-armed stellate hairs. Rhodoleia championii

41 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Lamina LS +/- glaucous green, margin darker than surface. Rhodoleia championii

42 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN leaf arrangement spirally Rhodoleia championii

43 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN leaf arrangement spirally Rhodoleia championii

44 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Rhodoleia championii sensu lato lamina rather variable

45 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Sycopsis sinensis

46 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Lamina LS green, margin translucent (10 LENS). Sycopsis sinensis

47 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Lamina LS green, margin translucent (10 LENS). Sycopsis sinensis

48 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Lamina LS green, margin translucent (10 LENS). Lamina apex predominantly gradually acuminate to (sub-)caudate. Sycopsis sinensis

49 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Lamina LS green, margin translucent (10 LENS). Distylium myricoides

50 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Lamina LS green, margin translucent (10 LENS). Distylium myricoides

51 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Lamina LS green, margin translucent (10 LENS). Apex obtuse to (abruptly) acute or abruptly acuminate. Distylium myricoides

52 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Distylium racemosum

53 GROUP B: LAMINA VENATION PINNATE MOST SECONDARY VEINS CURVING AND ANASTOMOSING BEFORE MARGIN Distylium racemosum

54 GROUP A LAMINA VENATION PALMATE strong veins predominantly radiating from base GROUP B LAMINA VENATION PINNATE most secondary veins curving + anastomosing before margin GROUP C LAMINA VENATION PINNATE most secondary veins ending at margin

55 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Hamamelis mollis

56 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Hamamelis mollis

57 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Fortunearia sinensis

58 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lamina margin with >25 teeth or vein endings/side. Fortunearia sinensis

59 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lamina margin with >25 teeth or vein endings/side. Lamina base rounded to cuneate. Fortunearia sinensis

60 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Sinowilsonia henryi

61 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lamina margin with >25 teeth or vein endings/side. Sinowilsonia henryi

62 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lamina margin with >25 teeth or vein endings/side. Lamina base cordate to obliquely cordate. Sinowilsonia henryi

63 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Parrotia persica

64 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lamina margin with 25 teeth or vein endings/side. Parrotia persica

65 GROUP C: LAMINA VENATION PINNATE Lamina margin with 25 teeth or vein endings/side. MOST SECONDARY VEINS ENDING AT MARGIN Corylopsis sinensis

66 Teeth bristly at apex. GROUP C: LAMINA VENATION PINNATE Lamina margin with 25 teeth or vein endings/side. MOST SECONDARY VEINS ENDING AT MARGIN Corylopsis sinensis

67 Teeth bristly at apex. GROUP C: LAMINA VENATION PINNATE Lamina margin with 25 teeth or vein endings/side. MOST SECONDARY VEINS ENDING AT MARGIN Corylopsis sinensis

68 Teeth bristly at apex. GROUP C: LAMINA VENATION PINNATE Lamina margin with 25 teeth or vein endings/side. Lamina base cordate. MOST SECONDARY VEINS ENDING AT MARGIN Corylopsis sinensis

69 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Hamamelis mollis

70 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lamina LS with lowermost secondary vein completely surrounded by leaf tissue. Hamamelis mollis

71 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Fothergilla major

72 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lowermost secondary vein OFTEN marginal for at least basal 2-10 mm. Fothergilla major

73 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lowermost secondary vein OFTEN marginal for at least basal 2-10 mm. Fothergilla major

74 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Hamamelis japonica

75 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lowermost secondary vein OFTEN marginal for at least basal 2-10 mm. Hamamelis japonica

76 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lowermost secondary vein OFTEN marginal for at least basal 2-10 mm. Hamamelis japonica

77 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Parrotia persica

78 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lamina LS secondary vein axils with tiny membranous domatia. Parrotia persica

79 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Parrotiopsis jacquemontiana

80 GROUP C: LAMINA VENATION PINNATE MOST SECONDARY VEINS ENDING AT MARGIN Lamina often suborbicular. Parrotiopsis jacquemontiana

81 JDL vegetative identification keys

82 JDL vegetative identification keys

83 thanks to IDS Wolfgang Bopp Tom Hudson Peter Brownless Jaime Morin Koen Camelbeke Jacky Pousse Tom Clark Abraham Rammeloo Francisco Garin Paul Reader Paul Goetghebeur Rolf Zumbrunn John Grimshaw Arboretum Kalmthout Arboretum Wespelaar Ghent University Botanical Garden Hillier Gardens and Arboretum Iturraran Botanical Garden Polly Hill Arboretum Royal Botanic Gardens Edinburgh Royal Botanic Gardens Kew

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