Stomatopoda (Crustacea) of the Gulf of Mexico

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2 Reaka, M. L., D. K. Camp, F. Álvarez, A. G. Gracia, M. Ortiz, and A. R. Vázquez-Bader Stomatopoda (Crustacea) of the Gulf of Mexico, Pp in Felder, D.L. and D.K. Camp (eds.), Gulf of Mexico Origins, Waters, and Biota. Biodiversity. Texas A&M University Press, College Station, Texas. 51 Stomatopoda (Crustacea) of the Gulf of Mexico Marjorie L. Reaka, David K. Camp, Fernando Álvarez,* Adolfo G. Gracia,* Manuel Ortiz,* and Ana Rosa Vázquez- Bader* The order Stomatopoda (mantis shrimps) is represented by an ecologically and behaviorally diverse group of actively predatory crustaceans that nevertheless exhibit only moderate morphological diversity compared to the decapods. All are shrimplike in form, with triramous antennules, a large abdomen supported by stiltlike uropods, full pleopods adorned with plumose gills, 3 pairs of walking legs, and 5 pairs of maxillipeds (of which the second pair is modified equally into unique hammering and spearing raptorial claws with a deadly strike mechanism; Patek, Korff, and Caldwell 2004, Patek and Caldwell 2005). The group has a long fossil history. The extinct tyrontophontids (suborder Archeostomatopodea) are known since the Carboniferous. The other suborder, the Unipeltata, includes the extinct sculdids and pseudosculdids from the Triassic and Jurassic as well as the 5 Recent superfamilies (Bathysquilloidea, Erythrosquilloidea, Gonodactyloidea, Lysiosquilloidea, Squilloidea; Schram and Müller 2004). The order Stomatopoda currently comprises 482 species (449 living, 33 fossil), with 122 genera (111 living, 11 extinct), 22 families (4 extinct), 5 living superfamilies (+ uncertain superfamily status of extinct groups), and 2 suborders (1 extinct; Schram and Müller 2004). In this report, we document 10% of the world s living stomatopod species (45 species, 2 of them as species complexes ), 22% of the world s genera (24 genera), 61% of the world s families (11), and 80% of the world s superfamilies (4) in the Gulf of Mexico (GMx). Stomatopoda. After Tavares Stomatopods are predominantly a tropical, shallowwater benthic group, but a few species extend into temperate and boreal waters and a considerable number inhabit deep, cold environments, including the outer continental shelf (some squilloids and lysiosquilloids) and the continental slopes (bathysquilloids; Manning 1969, 1995, Reaka and Manning 1987, Manning and Chace 1990). Although some stomatopods reach hypersaline and estuarine waters, most are restricted to marine habitats. The bathysquilloids, erythrosquilloids, most lysiosquilloids, and most squilloids inhabit level- bottom environments. The lysiosquilloids and squilloids are known to excavate burrows of varying complexity (Caldwell and Dingle 1975, Reaka and Manning 1981). The gonodactyloids generally * Order of authors marked with asterisks determined alphabetically. 901

3 902 ~ Stomatopoda (Crustacea) don 1911, Pearse 1929, Archer 1948, Gunter 1950, Hedgpeth 1950, 1953, 1954, Hildebrand 1954). After the publication of Bulletin 89 (1954), Springer and Bullis (1956) provided important early information on Gulf stomatopods (7 species) that were collected on the R / V Oregon expeditions, and Bullis and Thompson (1965) documented 12 species of stomatopods that were collected by the U.S. Fish and Wildlife Service on the exploratory fishing vessels Oregon, Silver Bay, Combat, and Pelican between 1956 and Raymond B. Manning (1959) provided an early important checklist of the stomatopods of Florida and nearby Gulf of Mexico waters (17 species), and Manning s (1969) monograph on the stomatopods of the Western Atlantic is the most comprehensive and influential treatment of the biology and distribution of stomatopods in the Gulf of Mexico to date (providing 33 of the present 45 species in the Gulf). Other notable studies included the Hourglass Cruises on the Central West Florida Shelf (Camp 1973, 12 species) and work in the northern Gulf (Adkison, Heard, and Hopkins 1983, Adkison and Hopkins 1984, Foster, Thoma, and Heard 2004; new genera and distributional information for 6 species). Studies of stomatopods in the far southeastern Gulf near northwestern Cuba have a long and venerable history, beginning with early exploratory investigations by Guérin- Méneville (1855, 1857), von Martens (1872, 1881), the voyages of the Albatross (Bigelow 1893, 1894), Torralbas (1917), Bouvier (1918), de Boury (1918), Parisi (1922), and the Atlantis and Oregon expeditions (Chace 1939, Springer and Bullis 1956). More recently, the stomatopod fauna of Cuba has been documented by Gómez and Ortiz (1985) and Ortiz and Lalana (2001), the latter providing information on the biology, size, abundance, habitat, and distribution of 16 species in Cuban waters. Of these, we report 7 species here (2 of these are included within species complexes of unknown taxonomic status) from the northwestern part of Cuba (south- southeast [sse] sector of the Gulf). Stomatopod research in Mexican waters of the Gulf has accelerated, with many important contributions having been made during the latter decades of the 20th century (Hernández- Aguilera and Villalobos 1980, Villalobos- Hiriart, Hernández- Aguilera, and Sosa- Hernández 1981, Hernández- Aguilera and Sosa- Hernández 1982, Morales- García 1986, 1987, Gamiño- Cruz et al. 1987, Hernández- Aguilera 1988, Hernández- Aguilera and Salazar 1988, Alvarado- Vásquez et al. 1989, Martínez- Guzmán and Hernández- Aguilera 1993, Hernández- Aguilera, Toral- Almazán, and Ruizoccupy coarse substrates. The larger- bodied hemisquillids and odontodactylids are often found on more open, sometimes moderately deep rubble plains (sometimes coarse sand / mud / shell; odontodactylids have been found in cavities in deep coral rubble; Reaka pers. obs.). The pseudosquillids usually are found in relatively shallow reef habitats, where the juveniles occupy bioeroded holes in coral while larger individuals excavate burrows in sand under coral and rubble. Most of the gonodactylids and protosquillids occupy bioeroded cavities in relatively shallow coral, rubble, and hard reef substrate. Species of smaller body size have significantly smaller geographic ranges, and, in each of the major superfamilies, some taxa of small body size exhibit accelerated evolution and major habitat shifts relative to their ancestral lineage (Reaka 1980a, Reaka and Manning 1987). All stomatopods are aggressive carnivores that exhibit strong intra- and interspecific competition and cannibalism of smaller individuals that can affect species distribution in local habitats (Dingle et al. 1973, Reaka 1985, 1987, Steger 1987). Stomatopods show unusual behavioral complexity, including learning (Reaka 1980b), individual recognition (Caldwell 1979, 1985), and complex territorial and reproductive communications that employ tactile, chemical, visual, and sound stimuli (Dingle and Caldwell 1969, 1972, Caldwell and Dingle 1975, Reaka and Manning 1981, Caldwell 1982, 1990, Hatziolos and Caldwell 1983, Patek and Caldwell 2006). The eyes of stomatopods exhibit remarkable complexity of structure and function, and are capable of seeing more colors than do humans, including ultraviolet, polarized, and fluoresced light (Schiff, Manning, and Abbott 1985, Cronin and Marshall 1989, Marshall and Land 1993, Cronin et al. 1994, Cronin, Caldwell, and Marshall 2001, Mazel et al. 2004, Chiao et al. 2005). Review of GMx Literature The stomatopods of the Gulf of Mexico were first comprehensively summarized in Bulletin 89 by Chace (1954), who listed 12 species and discussed a thirteenth species, Lysiosquillina glabriuscula (Lamarck, 1818), in the text. Chace cited 15 papers from the literature prior to We provide herein an additional 41 annotated citations from the literature prior to 1954 because of their significance to researchers interested in definitive identifications of species in the Gulf (many of these papers represent original species descriptions or taxonomic revisions of taxa in the Gulf, although a few are ecological, providing information on species that is otherwise likely to be lost; e.g., McClen-

4 Reaka et al. ~ 903 Nuño 1996). The latter paper reported 9 species of stomatopods. Vázquez- Bader and Gracia (1994) documented the benthic fauna (including 6 species of squilloid and lysiosquilloid stomatopods) from offshore level- bottom habitats of the Bay of Campeche, an important region for shrimp fisheries. We report here 18 species from the west- southwest (wsw) and / or south- southwest (ssw) and 7 additional species from the west- northwest (wnw) sectors of the Gulf, making a total of 25 species (including 2 species complexes of unknown taxonomic status, plus several uncounted intergrading forms) from the region (see chapter 1, fig. 1). We also are aware of research being conducted by Illescas- Monterroso, Salgado- Barragán, and Hernández- Ugalde that documents the stomatopod fauna of México, including the Caribbean coast. When published, that work may alter our list or expand the ranges into the western and southern Gulf of species that we report elsewhere in the Gulf. Systematic Revisions Major systematic developments that have affected the taxonomy of stomatopods in the Gulf of Mexico since Bulletin 89 include Manning s early papers (1963, 1967, 1968) that identified a new family and a number of new genera. These were followed by his monumental Stomatopod Crustacea of the Western Atlantic (1969), which remains the classic taxonomic reference for West Atlantic and Gulf stomatopods. Manning s treatments of East Atlantic (1977) and Indo- West Pacific (1995) taxa revised many genera and families and updated nomenclature for West Atlantic and Gulf of Mexico species. Revisionary works by Ahyong and Harling (2000) and Ahyong (2001) also made important contributions to our knowledge of the systematics and taxonomy of many genera and thus affected the taxonomy of species in the Gulf. The most important current publication for researchers who wish to sort out the taxonomy and associated literature for Gulf stomatopods is Schram and Müller s (2004) Catalog and Bibliography of the Fossil and Recent Stomatopoda, which provides synonomies and references for all species that occur in the Gulf and elsewhere. A number of other taxonomic works have revised taxonomic relationships within genera and erected new genera that affect Gulf species (e.g., Manning and Camp 1981, 1983, Manning and Chace 1990, Cappola and Manning 1995, Holthuis 2000). Manning and Schotte (1993) provide a particularly informative discussion of progress in taxonomy for different regions of the West Atlantic. Molecular studies are just beginning to provide valuable information on the relationships among taxa of stomatopods (Barber and Erdmann 2000, Barber, Moosa, and Palumbi 2002, Barber et al. 2002), and this route of investigation undoubtedly will play an important part in our future understanding of the taxonomic relationships among different species and lineages of stomatopods in the Gulf of Mexico and elsewhere. Comparative Assessment of the Group Of the 45 species reported here (2 of them as species complexes), 20 (44.4%) have been described since Chace (1954), and 7 (15.6%) have been described since Manning s (1969) monograph on West Atlantic stomatopods. One of the two most serious taxonomic problems we encountered was intergrading morphological features between Neogonodactylus bredini and N. wennerae. Discussions with Paul Barber, who is investigating molecular relationships among populations of these species, have also suggested that the genetic relationships among the populations are complex (Barber, pers. comm.). Thus, we treat them here only as the N. bredini / wennerae complex, and we are continuing to investigate the taxonomic and distributional relationships of this group. Another problem we have encountered was indefinite morphological relationships among specimens of Odontodactylus from the Gulf, Florida Keys, and Cuba. Thus, we categorized these records here only as the O. brevirostris / havanensis complex, and we are continuing to examine this issue. Eight and probably 10 ( %) of the 45 species, and 2 and probably 3 ( %) of the 24 genera reported here are apparently endemic to the Gulf of Mexico, although the difficulty of collecting small- bodied, cryptic species suggests that some of these could be found elsewhere in the future. This high apparent endemicity exceeds Briggs (1974, 1995) criterion (10% endemism of species) for establishing separate biogeographic provinces, and suggests that, if this pattern holds for other taxa, the Gulf of Mexico should be regarded as an independent biogeographic province that is separate from the other West Atlantic provinces. Surprisingly, all of the endemic taxa belong to the Lysiosquilloidea, a group that is generally considered to have rather widely distributed species and larvae with lengthy pelagic phases; clearly, extensive dispersal, however, does not hold for some of the groups of smaller body size such as Acoridon, Tectasquilla, and Nannosquilla (Reaka 1980a, Reaka and Manning 1987). Gulf endemics include Acoridon manningi, a genus and species known from only one site off southwestern Florida; it rep-

5 904 ~ Stomatopoda (Crustacea) resents the only member of the Coronididae in the West Atlantic, although the related genus Coronida occurs in the East Pacific (2 species) and East Atlantic (one species). The large- bodied Lysiosquilla campechiensis in contrast to its more widespread West Atlantic congener L. scabricauda is known only from the Bay of Campeche, while other members of the genus are known from the East Pacific (2 species), East Atlantic (2 species), and Indo- West Pacific / Indian Ocean (4 species). Five of the 6 small- bodied species of Nannosquilla present in the Gulf are apparently endemic, most being known from only a single or a few sites off Florida or Yucatán; N. schmitti is the only moderately widespread species of Nannosquilla that occurs in the Gulf (from off Florida, Yucatán, and Barbuda). The small- bodied monospecific genus Mexisquilla, allied to Nannosquilla, is found only in the Gulf of Mexico and is known from only one site off western Florida (Camp 1971). Two additional species merit special attention. Tectasquilla lutzae likely is also endemic to the Gulf of Mexico. This species is known definitively only from northwest Florida, but a fragment from northeastern Florida / Georgia possibly belongs to this species (Adkison and Hopkins 1984, Manning and Camp 1993). Schotte and Manning (1993) list T. lutzae as a species only from the Gulf of Mexico. Additionally, although many squillids have long- lived larvae and fairly wide distributions, Squilla chydaea is nearly endemic to the Gulf of Mexico, with collections outside the Gulf being limited to only one individual off Cape Canaveral and one individual off Palm Beach, Florida (Manning 1969); however, this species is widespread and very abundant in the Gulf of Mexico. Thirteen (28.9%) other stomatopods have distributions restricted to one or two sectors of the Gulf, although their geographic ranges extend beyond the Gulf. The distribution patterns of this group are especially illuminating biogeographically because most indicate a center of abundance and diversity that extends toward the south and east. The distributions of 9 Gulf species Neogonodactylus caribbaeus (Florida Keys), Pullosquilla litoralis (Florida Keys), Tectasquilla lutzae (northwestern Florida, but see the previous discussion), Tetrasquilla mccullochae (Florida Keys), Eurysquilla plumata (southwestern Florida, Florida Keys, Dry Tortugas), Cloridopsis dubia (northwestern Florida, questionably the Florida Keys, Manning 1967, 1969), Meiosquilla randalli (Florida Keys), Squilla grenadensis (western Florida), and S. intermedia (off Havana) penetrate the Gulf only in one eastern or southeastern sector. All species extend elsewhere in the West Atlantic except for P. litoralis, which occurs only in the Gulf and the Indo- West Pacific. Two other species (T. mccullochae, C. dubia) also occur in other oceans as well as the West Atlantic. Four other species are known in the Gulf from both southeastern and southwestern sectors, although they all extend more widely to the south and east: Neogonodactylus spinulosus (Florida Keys, Campeche Bay), Lysiosquillina glabriuscula (Key West, Dry Tortugas, Veracruz), Nannosquilla schmitti (Florida Keys, western Yucatán), Alima neptuni (Florida Keys, Campeche Bay). Nannosquilla schmitti is reported from a third widely separated West Atlantic site, N. spinulosus and L. glabriuscula are widely distributed in the West Atlantic, and A. neptuni is known from patchy locations in the West Atlantic as well as the Central Atlantic, East Atlantic, and Indo- West Pacific. Supporting the suggestion of a diversity gradient extending from the Gulf toward the south and east, examination of the taxon summary shows that the greatest diversity of stomatopods occurs in the east- southeast (ese) sector, which includes the Florida Keys and northwestern Cuba (61.8% of Gulf species occur there). The next most diverse sector is the east- northeast (ene) part of the Gulf, although this may represent more extensive research off the west coast of Florida in the 1970s and 1980s than is generally true for other sectors. The relative paucity of species we report in the south- southeast (sse) sector of the Gulf (Yucatán) compared to the better studied south- southwest (ssw) sector suggests that more work is needed for the former region. The region north of Veracruz (west- southwest, wsw) also appears to need more investigation, although reef habitat may be less available there than in other areas, accounting for the relatively low numbers of species. The area off the Texas coast (northnorthwest, nnw) also has fewer species than the adjacent west- northwest (wnw), the latter being enriched by the Flower Garden Banks. Three species of stomatopods are documented in the Gulf for the first time in the present study. One of them, Neogonodactylus lacunatus, is reported in the southeastern and western parts of the Gulf, although this species had been known from Southeast Florida and the Caribbean side of Yucatán previously. Another interesting new record is Pullosquilla litoralis, an Indo- West Pacific nannosquillid. A single specimen has been taken in the Florida Keys. Because this family is characterized by high endemism, we feel that this record must represent an introduced species that may have arrived by ballast water from ships or from the aquarium industry. Whether a

6 Reaka et al. ~ 905 permanent population has been established in the West Atlantic is currently unknown. Additionally, we provide the first definitive record of Cloridopsis dubia for the Gulf of Mexico. A questionable record of this species in the Marquesas Keys was discussed by Manning (1967, 1969), and a dry, fragmented specimen from Mexico with no locality data was cited by Manning (1969) and Schram and Müller (2004). The species was collected in Cuba by Gundlach, but the absence of locality data precludes its inclusion in Gulf waters. Additionally, this study yields a number of new range extensions and enlarged depth distributions. The geographic range is extended into 25 new Gulf sectors for 13 species (28.9% of all Gulf species sport new sector records; each of these species has had its range extended by an average of 1.9 sectors). We definitively document Neogonodactylus bredini / wennerae in the west- northwestern Gulf for the first time, although Manning (1969) cited a questionable record from Texas. Formerly known only from southern Florida (including the Keys), the Bahamas, and the Caribbean Sea, the distribution of N. curacaoensis is expanded here into both the northeastern and northwestern Gulf. The range of N. torus is enlarged in the northeastern and northwestern Gulf with this study. The distribution of Odontodactylus brevirostris / havanensis complex and Pseudosquilla ciliata now includes new sectors in both the northeastern and the northwestern parts of the Gulf. Rare in the West Atlantic (Reaka, pers. obs.), Pseudosquillisma oculata had been known only from northwestern Cuba (Pinar del Rio Province), the Florida Keys, and Caribbean Yucatán (Manning 1969). A postlarva in this study extends the distribution of this species to the Flower Garden Bank off Texas. Pullosquilla litoralis and Parasquilla coccinea have now been documented in the sse sector of the Gulf, and Gibbesia prasinolineata is recorded in the ssw sector for the first time. The range of Meiosquilla schmitti now is enlarged to include the westnorthwestern Gulf and the Florida Keys. The distribution of Squilla edentata edentata is expanded into both the eastern and the southwestern parts of the Gulf. This study also extends the depth distributions of 9 species (20% of all Gulf species). The known depth ranges are expanded into deeper water in 8 cases (N. bredini / wennerae, N. curacaoensis, Lysiosquillina glabriuscula, Meiosquilla randalli, M. schmitti, Squilla chydaea, S. edentata edentata, S. rugosa) and shallower water in 2 cases (Neogonodactylus caribbaeus and Squilla edentata edentata). A number of additional species likely will be found in the Gulf of Mexico in the future because a number Stomatopoda. After Bigelow in Bowers of West Atlantic species have been taken from regions close to the Gulf. For example, Alachosquilla digueti and A. floridensis have been reported from near Miami, and from Lake Worth and Key Biscayne, respectively, as well as more widespread locations. Nannosquilla antillensis and N. yucatanica occur on the Caribbean side of Yucatán; N. whitingi is known from the east coast of Florida and Heterosquilloides insolita from the northern straits of Florida (the latter also possibly occurs in the Galapagos). Eurysquilla chacei, Squilla discors, and Meiosquilla lebouri are recorded only from the Bahamas, while M. tricarinata has been collected in the Bahamas and off eastern Florida. The amphi- Atlantic Alima hildebrandi is known from an unspecified locality in Cuba (from Gundlach s collection) as well as Panamá and ; thus, this species may someday be collected in the Gulf of Mexico as well. None of the 12 families of stomatopods now found in the West Atlantic (WA) are endemic to the WA; all but one (Hemisquillidae) of these families (91.7%) occur in the Gulf. Thirty genera are now present in the West Atlantic (including Pullosquilla, apparently introduced but perhaps not established). Of these, 24 (80%) are found in the Gulf of Mexico. Five of these 30 genera (16.7%) are endemic to the West Atlantic, and 2 (possibly 3, %) genera are endemic to the Gulf. Of the 86 species now known to occur in the West Atlantic (including P. litoralis and counting the Neogonodactylus bredini / wennerae and Odontodactylus brevirostris / havanensis complexes as one species each), 45 (52.3%) occur in the Gulf. Of these 45 Gulf species, 8 (possibly 10, %) are endemic or nearly endemic to the Gulf. The distributions of all of the remaining species extend to the West Atlantic or beyond. Twenty- six (57.8%) of the Gulf species occur only in the West Atlantic, and 9 (20%) of the Gulf species are additionally distributed in other oceans:

7 906 ~ Stomatopoda (Crustacea) 2 (4.4%) are shared only with the East Pacific (Bigelowina biminiensis, Cloridopsis dubia), 3 (6.6%) are shared only with the Indo- Pacific (Bathysquilla microps, Odontodactylus brevirostris / havanensis complex (?), Pullosquilla litoralis), none are shared only with the East Atlantic, 3 (6.6%) are shared only with the Indo- West Pacific and the East Atlantic (Pseudosquilla ciliata, Pseudosquillisma oculata, Alima neptuni), and one (2.2%) is shared only with the Indo- West Pacific and the East Pacific (Tetrasquilla mccullochae). Except for Pullosquilla litoralis, which we believe is introduced, no species occurs only in the Gulf of Mexico and another ocean. Thus, the Gulf of Mexico does not appear to be a refuge from West Atlantic extinction in the Tertiary. Plan of the Checklist and Abbreviations Used The attached checklist summarizes the current status of the stomatopods of the Gulf of Mexico. Superfamilies are arranged alphabetically; families, genera, and species are nested alphabetically within each group. Abbreviations for habitat include: ben = benthic; bur = burrower; crr = coral reef; dps = deep sea; hsb = hard substrate; inf = infaunal; ocs = outer continental shelf; lvb = level bottom; rbl = rubble; sft = soft substrate (mud, sands, clays); sgr = sand and gravel; shl = shell; slp = continental slope; and snd = sand. Abbreviations for endemic and nonindigenous to Gulf of Mexico = end and nid. Depth records in the GMx are based on our data and the literature and are indicated first in roman type, followed by the global depth distribution previously known for the species in parentheses and italic type; sh = shallow; lit = littoral / intertidal; sublit = sublittoral / subtidal. For the Overall Geographic Range, GMx = Gulf of Mexico; WA = Western Atlantic; EA = Eastern Atlantic; CA = Mid- Atlantic; EP = Eastern Pacific; CP = Central Pacific; IWP = Indo- West Pacific; IO = Indian Ocean; and E, W, N, S = eastern, western, northern, southern. Both overall geographic range and GMx range include our data plus reports in the literature; our new distribution records and depth records are documented in endnotes. Abbreviations for the institutions who hold the specimens we cite are identified in our acknowledgments, except for specimens from the Florida Fish and Wildlife Research Institute, whose specimens are prefaced by FSBC I, and specimens from Texas A&M University, whose specimens reside in the Texas Cooperative Wildlife Collection at TAMU College Station and are prefaced by TCWC. References include dis- cussions of Gulf species, even if the study was conducted outside of the Gulf (see notes in the annotated references list). Gulf sectors are east- southeast = ese; east- northeast = ene; north- northeast = nne; north- northwest = nnw; west- southwest = wsw; south- southwest = ssw; and south- southeast = sse. Question marks beside a literature citation indicate that we are not sure of the identity of the species cited in the reference due to subsequent taxonomic work. Acknowledgments We thank Dr. Wes Tunnell and the Harte Research Institute for providing funding that allowed us (MLR, DKC) to visit all of the major collections of stomatopods from the Gulf of Mexico, and Drs. Darryl Felder and Suzanne Fredericq and the National Science Foundation (grant #DEB ), which supported recent cruises in the Gulf of Mexico in which one of us (MLR) participated. We are especially grateful to Drs. Darryl Felder, Nancy Voss, Raphael Lemaitre, Fernando Álvarez, and Ardis Johnston for providing data and graciously hosting some of us (MLR, DKC) in their laboratories at the University of Louisiana; Rosenstiel School of Marine and Atmospheric Sciences, University of Miami (RSMAS); Division of Crustacea at the National Museum of Natural History, Smithsonian Institution (USNM); Department of Zoology and National Crustacean Collection at the Universidad Nacional Autonóma de México (UNAM); and Museum of Comparative Zoology at Harvard University, respectively. We are indebted to Dr. Wes Tunnell of Texas A&M University Corpus Christi and to Ms. Sandra Farrington of the Florida Fish and Wildlife Research Institute for providing access to their collections and records, and we greatly appreciate the assistance of Ms. Sara LeCroy of the University of Southern Mississippi Gulf Coast Research Laboratory (GCRL) and Dr. Mary Wicksten of Texas A&M University College Station (TAMU), who sent us specimens and / or data from their institutions. We thank Dr. Luis Soto, Dr. Carlos Illescas- Monterroso, and Carmen Hernández- Ugalde of the Instituto de Ciencias del Mar y Limnología, UNAM, as well as Dr. Richard Heard of the Gulf Coast Research Laboratory, for useful discussions and sharing information. Special thanks are due Dr. Fred Schram, who shared information from his Catalog and Bibliography of Fossil and Recent Stomatopoda with one of us (MLR) as it was being developed, without which complete compilations of literature for Gulf species

8 Reaka et al. ~ 907 would have been much more difficult and undoubtedly less complete. References 1. Adkison, D. L., R. W. Heard, and T. S. Hopkins Description of a new genus and species, Acoridon manningi (Stomatopoda: Coronididae), from the Gulf of Mexico. Journal of Crustacean Biology 3: [Gulf species: Acoridon manningi: original description (West Florida)] 2. Adkison, D. L., and T. S. Hopkins Tectasquilla lutzae, a new genus and species (Crustacea: Stomatopoda: Lysiosquillidae) from the Gulf of Mexico. Proceedings of the Biological Society of Washington 97: [Gulf species: Tectasquilla lutzae: original description (West Florida)] 3. Ahyong, S. T Revision of the Australian stomatopod Crustacea. Records of the Australian Museum, Supplement 26: [Revisionary work affecting Gulf genera and species] 4. Ahyong, S. T., and C. Harling The phylogeny of the stomatopod Crustacea. Australian Journal of Zoology 48: [Revision] 5. Alvarado- Vázquez, H. N., O. Badillo- Marmolejo, M. R. Calderón- Ramírez, et al Crustáceos, estomatópodos y decápodos de los Arrecifes de Enmedio, Veracruz y Cayos Arcas, Campeche. Rep. Biol. Campo, Fac. Cienc., UNAM. 71 pp. [Gulf species: UNKNOWN; cited in Marínez- Guzmán and Hernández- Aguilera, 1993] 6. Anonymous Annotated List of the Fauna of the Grand Isle Region, Marine Laboratory, Louisiana State University, Baton Rouge. 20 pp. [Gulf species: Lysiosquilla scabricauda (Louisiana); Coronis scolopendra (Louisiana); Squilla empusa (Louisiana)] 7. Archer, A. A The shrimp surveys. Alabama Conservation 20(2): 10, 1 fig. [Gulf species: Squilla rugosa (Alabama: Mississippi Sound)] 8. Barber, P. H., and M. V. Erdmann Molecular systematics of the Gonodactylidae (Stomatopoda) using mitochondrial cytochrome oxidase C (subunit I) DNA sequence data. Journal of Crustacean Biology 20: [Revision] 9. Barber, P. H., M. K. Moosa, and S. R. Palumbi Rapid recovery of genetic diversity of stomatopod populations on Krakatau: temporal and spatial scales of marine larval dispersal. Proceedings of the Royal Society of London 269: Barber, P. H., S. R. Palumbi, M. V. Erdmann, and M. K. Moosa Sharp genetic break among populations of Haptosquilla pulchella (Stomatopoda) indicate limits to larval transport: patterns, causes and consequences. Molecular Ecology 11: Behre, E. H Annotated list of the fauna of the Grand Isle region, Occasional Papers of the Marine Laboratory, Louisiana State University 6: [Gulf species: Lysiosquilla scabricauda (Louisiana); Coronis scolopendra (Louisiana); Squilla empusa (Louisiana)] 12. Bigelow, R. P Preliminary notes on the Stomatopoda of the Albatross collections and on other specimens in the National Museum. Johns Hopkins University Circular 12: [Gulf species: Meiosquilla quadridens: original description (Florida Keys); Squilla rugosa (West Florida); Squilla edentata edentata (Mississippi Delta); Squilla empusa (West Florida); Pseudosquilla ciliata (Florida); Odontodactylus brevirostris / havanensis complex (O. havanensis original description: Cuba, locality unknown);?neogonodactylus oerstedii (Florida Keys)] 13. Bigelow, R. P The Stomatopoda of Bimini. Johns Hopkins University Circular 12(106): [Gulf species: Bigelowina biminiensis: original description] 14. Bigelow, R. P Report on the Crustacea of the Order Stomatopoda collected by the steamer Albatross between 1885 and 1891 and on other specimens in the U.S. National Museum. Proceedings of the U.S. National Museum 17: , pls [Gulf species: Lysiosquilla scabricauda (Texas); Meiosquilla quadridens (Florida Keys); Squilla rugosa: original description (West Florida); Squilla edentata edentata (Mississippi Delta); Pseudosquilla ciliata (Florida); Odontodactylus brevirostris complex (Cuba: no specific locality); Neogonodactylus torus (Cuba: off Havana);?Neogonodactylus oerstedii (Florida Keys)] 15. Bigelow, R. P The Stomatopoda of Porto Rico. Bulletin of the U.S. Fish Commission 20: [Gulf species: Eurysquilla plumata: original description)] 16. Boone, L Crustacea: Stomatopoda and Brachyura. Scientific results of the cruises of the yachts Eagle and Ara, , Wm. K. Vanderbilt, commanding. Bulletin of the Vanderbilt Marine Museum 2: 1 228, 74 pls. [Gulf species:?neogonodactylus oerstedii (Florida Keys)] 17. Bouvier, E. L Sur les arthropodes du Mozambique et de San Thome offerts au Muséum par M. Almada Negreiros. Bulletin du Museum d Histoire Naturelle, Paris 7: [Gulf species:?neogonodactylus oerstedii (Cuba: no specific locality)]

9 908 ~ Stomatopoda (Crustacea) 18. Briggs, J. C Marine Zoogeography. McGraw Hill, N.Y. 475 pp. 19. Briggs, J. C Global Biogeography. Elsevier Science, St. Louis. 452 pp. 20. Brullé, M Crustacés. In: P. Barker- Webb and S. Berthelot, eds. Histoire Naturelle des Iles Canaries. Zoologie 2(2) Entomologie: 13 18, atlas, pls. [Gulf species: Pseudosquillisma oculata: original description] 21. Bullis, H. R. Jr., and J. R. Thompson Collections by the exploratory fishing vessels Oregon, Silver Bay, Combat, and Pelican made during in the southwestern North Atlantic. U.S. Fish and Wildlife Service, Special Scientific Report 510: [Gulf species: Lysiosquilla scabricauda (Louisiana); Lysiosquillina glabriuscula (Florida); Lysiosquilla campechiensis (Mexico); Bathysquilla microps (SW of Tortugas); Meiosquilla quadridens (Straits of Florida, Gulf of Mexico); Squilla deceptrix (West Florida); Fennerosquilla heptacantha (Gulf of Mexico); Squilla chydaea (West Florida to Mexico); Squilla edentata edentata (Gulf of Mexico); Pseudosquilla ciliata (Gulf of Mexico); Parasquilla coccinea (NW Florida, Mississippi Delta, Mexico); Odontodactylus brevirostris / havanensis complex (Louisiana)] 22. Caldwell, R. L Cavity occupation and defensive behavior in the stomatopod Gonodactylus festae: evidence for chemically mediated individual recognition. Animal Behavior 27: Caldwell, R. L Interspecifically chemically mediated recognition in two competing stomatopods. Marine Behavior and Physiology 8: Caldwell, R. L A test of individual recognition in the stomatopod Gonodactylus festae. Animal Behavior 33: Caldwell, R. L Variation in reproductive behavior in stomatopod Crustacea. Pp in R. T. Bauer and J. W. Martin, eds. Crustacean Sexual Biology. Columbia University Press, N.Y. 26. Caldwell, R. L Recognition, signaling and reduced aggression between former mates in a stomatopod. Animal Behavior 44: Caldwell, R. L., and H. Dingle Ecology and evolution of agonistic behavior in stomatopods. Naturwissenschaften 62: [Review; Gulf species: Neogonodactylus bredini / wennerae complex, Bermuda]. 28. Camp, D. K Platysquilla horologii (Stomatopoda: Lysiosquillidae), a new species from the Gulf of Mexico, with an emendation of the generic definition. Proceedings of the Biological Society of Washington 84: [Gulf species: Mexisquilla horologii: original description (West Florida)] 29. Camp, D. K Stomatopod Crustacea. Memoirs of the Hourglass Cruises 3(2): [Gulf species: Lysiosquilla scabricauda; Bigelowina biminiensis; Meiosquilla quadridens; Meiosquilla schmitti; Gibbesia neglecta; Squilla grenadensis; Squilla rugosa; Squilla deceptrix; Squilla empusa; Eurysquilla plumata; Parasquilla coccinea; Neogonodactylus bredini / wennerae complex (all West Florida)] 30. Camp, D. K Occurrence of Squilla heptacantha (Chace, 1939) (Crustacea: Stomatopoda: Squillidae) in the northeastern Gulf of Mexico. Northeast Gulf Science 6: [Gulf species: Fennerosquilla heptacantha (West Florida)] 31. Camp, D. K Occurrence of Heterosquilloides armata (Smith) in the northwestern Gulf of Mexico, with notes on variation within the species (Crustacea: Stomatopoda: Lysiosquillidae). Journal of Crustacean Biology 5: [Gulf species: Heterosquilloides armata (Texas)] 32. Camp, D. K., and R. B. Manning Five new species of Nannosquilla from the northwestern Atlantic (Crustacea: Stomatopoda). Smithsonian Contributions to Zoology 368: [Gulf species: Nannosquilla adkinsoni: original description (West Florida); Nannosquilla heardi: original description (West Florida)] 33. Camp, D. K., and R. B. Manning Observations on Nannosquilla, with descriptions of three new species from the northwestern Atlantic (Crustacea: Stomatopoda). Smithsonian Contributions to Zoology 444: [Gulf species: Nannosquilla adkisoni (West Florida); Nannosquilla disca: original description (West Florida)] 34. Cappola, V., and R. B. Manning [1994]. Crustacea Stomatopoda. Research on the Coast of Somalia. Tropical Zoology 7(2) (for 1994): [Revision; Pseudosquillisma] 35. Cary, L. R., and H. M. Spalding Further contributions to the marine fauna of the Louisiana coast. Publications of the Gulf Biological Station, Cameron: [Gulf species: Squilla empusa (Louisiana)] 36. Chace, F. A. Jr Preliminary descriptions of one new genus and seventeen new species of decapod and stomatopod Crustacea. Reports on the scientific results of the first Atlantis expedition to the West Indies, under the joint auspices of the University of Havana and Harvard University. Memorias de la Sociedad Cubana de Historia Natural 13: [Gulf species: Fennerosquilla heptacantha: original description (northern coast of Cuba)]

10 Reaka et al. ~ Chace, F. A. Jr Six new species of decapod and stomatopod Crustacea from the Gulf of Mexico. Proceedings of the New England Zoological Club 19: 79 92, pls [Gulf species: Odontodactylus brevirostris / havanensis complex (O. nigricaudatus: original description, Mexico)] 38. Chace, F. A. Jr Stomatopoda. Pp in P. S. Galtsoff, ed. Gulf of Mexico, Its Origin, Waters, and Marine Life. Fishery Bulletin 89. Fishery Bulletin of the Fish and Wildlife Service, Volume 55, Washington, D.C. [Gulf species: Lysiosquilla scabricauda (Florida, Louisiana, Texas); Lysiosquillina glabriuscula (Gulf of Mexico); Coronis scolopendra (Alabama, Louisiana); Gibbesia neglecta (Florida: Sanibel Is.); Gibbesia prasinolineata (Mexico); Squilla rugosa (West Florida); Squilla edentata edentata (northern Gulf of Mexico); Squilla empusa (West Florida, Louisiana, Texas, Mexico); Pseudosquilla ciliata (Florida Keys); Odontodactylus brevirostris / havanensis complex (Florida Keys, Mexico); Neogonodactylus torus (Cuba: off Havana);?Neogonodactylus oerstedii (Key West, Tortugas)] 39. Chiao, C. C., T. W. Cronin, R. L. Caldwell, and J. Marshall Biological polarized light reflectors in stomatopod crustaceans. Polarization Science and Remote Sensing II: Society of Photo- Optical Instrumentation Engineers, San Diego. [Gulf species: Odontodactylus brevirostris / havanensis complex as O. havanensis (Florida)] 40. Cronin, T. W., and N. J. Marshall A retina with at least ten spectral types of photoreceptors in a mantis shrimp. Nature 339: [Gulf species: Pseudosquilla ciliata (collection region not given)] 41. Cronin, T. W., R. L. Caldwell, and N. J. Marshall Tunable color vision in a mantis shrimp. Nature 411: Cronin, T. W., N. J. Marshall, C. A. Quinn, and C. A. King Ultraviolet photoreception in mantis shrimp. Vision Research 34: Cronin, T. W., N. Shashar, R. L. Caldwell, J. Marshall, A. G. Cheroske, and T. H. Chiou Polarization vision and its role in biological signaling. Integrative and Comparative Biology 43: Dana, J. D Crustacea. Part I. United States Exploring Expedition During the Years 1838, 1839, 1840, 1841, 1842 Under the Command of Charles Wilkes, U.S.N. 13: 1 685; atlas, 1855: 1 27, pls C. Sherman, Philadelphia. [Gulf species: Gibbesia prasinolineata: original description] 45. Dawson, C. E Progress Report. Gulf Coast Research Laboratory Museum, Mimeo, Ocean Springs, Mississippi. iii + 34 pp. [Gulf species: Squilla edentata edentata (Alabama)] 46. Dawson, C. E Progress Report. Gulf Coast Research Laboratory Museum, Mimeo, Ocean Springs, Mississippi. iii + 57 pp. [Gulf species: Lysiosquilla scabricauda (Mississippi); Coronis scolopendra (Mississippi); Squilla chydaea (Alabama, Louisiana); Squilla edentata edentata (Alabama); Squilla empusa (Alabama, Mississippi, Louisiana)] 47. de Boury, M. E Quelques observations sur la moeurs et sur l habitat des crustacés à l île de Cuba. Bulletin Museum Histoire Naturelle, Paris 24: [Gulf species:?neogonodactylus oerstedii (Cuba: no specific locality)] 48. de Lamarck, J. B. P. A Histoire naturelle des animaux sans vertèbres, présentent les caractéres généraux et particuliers de ces animaux, leur distribution, leur classes, leurs familles, leurs genres, et la citation des principales espèces qui s y rapportent; précédée d une introduction offrant la détermination des caractères essentials, de l animal, sa distinction du végétal et des autres corps naturelles, enfin, l exposition des principes fondamentaux de la zoologie, 5. Deterville, Paris. 612 pp. [Gulf species: Lysiosquilla scabricauda: original description; Lysiosquillina glabriuscula: original description] 49. Dingle, H., and R. L. Caldwell The aggressive and territorial behavior of the mantis shrimp Gonodactylus bredini Manning (Crustacea; Stomatopoda). Behaviour 33: [Gulf species: Neogonodactylus bredini / wennerae complex (Bermuda)] 50. Dingle, H., and R. L. Caldwell Reproductive and maternal behavior of the mantis shrimp Gonodactylus bredini Manning (Crustacea; Stomatopoda). Biological Bulletin 142: [Gulf species: Neogonodactylus bredini / wennerae complex (Bermuda)] 51. Dingle, H., R. C. Highsmith, K. E. Evans, and R. L. Caldwell Interspecific aggressive behavior in tropical reef stomatopods and its possible ecological significance. Oecologia 13: Fabricius, J. C Mantissa Insectorum Sistens Eorum Species Nuper Detectas Adjectis Characteribus Genericis, Differentiis Specificis, Emendationibus, Observationibus. 1. Christ. Gottl. Proft, Hafniae. xx pp. [Gulf species: Pseudosquilla ciliata: original description] 53. Faxon, W Crustacea. In: A. Agassiz, W. Faxon, and E. L. Mark, eds. Selections from embryological monographs, I. Memoirs of the Museum of Comparative Zoölogy, Harvard College 9(1): pls [Gulf species: Squilla empusa (Mexico)]

11 910 ~ Stomatopoda (Crustacea) 54. Foster, J. F., B. P. Thoma, and R. W. Heard Stomatopoda (Crustacea: Hoplocarida) from the shallow, inshore waters of the northern Gulf of Mexico (Apalachicola River, Florida to Port Aransas, Texas). Gulf and Caribbean Research 16: [Gulf species: Neogonodactylus bredini / wennerae complex (NW Florida); Lysiosquilla scabricauda (NW Florida, Mississippi Sound); Bigelowina biminiensis (NW Florida, Mississippi); Coronis scolopendra (NW Florida, Mississippi); Squilla empusa (NW Florida, Mississippi, Louisiana); Gibbesia neglecta (Mississippi Sound)] 55. Gamiño, M. G., S. G. González, E. González, M. Hermoso, E. Mendoza, L. Quintana, N. Trujillo, C. Zapote, A. Velázquez, J. Miranda, R. Romero, V. Reynoso, I. Islas, M. Cano, and C. Avendaño Comparación de los crustáceos superiores de una zona costera y una insular del Estado de Veracruz, Ver. México. Biología de Campo, Facultad de Ciensias, UNAM. 72 pp. [Gulf species: Neogonodactylus oerstedii (Mexico: Veracruz); Pseudosquilla ciliata (Mexico: Veracruz)] 56. Gibbes, L. R On the carcinological collections of the cabinets of natural history in the United States, with an enumeration of the species contained therein, and descriptions of new species. Proceedings of the American Association for the Advancement of Science, Third Meeting: [Gulf species: Gibbesia neglecta: original description;?neogonodactylus oerstedii (Key West)] 57. Gómez, O., and M. Ortiz Lista de especies y bibliografia de los estomatópodos (Crustacea, Hoplocarida) de aguas cubanas. Revista de Investigaciones Marinas 6 (2-3): [Gulf species: Neogonodactylus bredini / wennerae complex, Neogonodactylus oerstedii, Neogonodactylus torus, Odontodactylus brevirostris / havanensis complex, Bigelowina biminiensis, Pseudosquilla ciliata, Pseudosquilla oculata, Lysiosquilla glabriuscula, Lysiosquilla scabricauda, Cloridopsis dubia, Fennerosquilla heptacantha, Gibbesia prasinolineata, Squilla empusa, Squilla intermedia, Squilla rugosa (all records from Cuba, no specific locality); Alima hieroglyphica: erroneous record] 58. Guérin- Méneville, F. E Crustáceos. Pp. v xxxii; atlas, 8: pls. 1 3 in R. de la Sagra, ed. Historia fisica política y natural de la isla de Cuba, Historia Natural, 7. Arthus Bertrand, Paris. [Gulf species: Lysiosquilla scabricauda (Cuba)] 59. Guérin- Méneville, F. E Crustacés. Pp. xiii lxxxvii, pls. 1-3 in R. Sagra, ed. Histoire physique, politique et naturelle de l ile de Cuba. Arthus Bertrand, Paris. [Gulf species: Lysiosquilla scabricauda (Cuba)] 60. Gunter, G Seasonal population changes and distributions as related to salinity of certain invertebrates of the Texas coast, including the commercial shrimp. Publications of the Institute of Marine Science, University of Texas 1: [Gulf species: Squilla empusa (Texas)] 61. Hansen, H. J Isopoden, Cumaceen und Stomatopoden der Planktonexpedition. Ergbn. Planktonexped. Humbolt- Stiftung 2 (Gc): 1 105, pls [Gulf species: Neogonodactylus oerstedii: original description] 62. Hatziolos, M. E., and R. L. Caldwell Role reversal in courtship in the stomatopod Pseudosquilla ciliata (Stomatopoda). Animal Behavior 31: [Gulf species: Pseudosquilla ciliata (Jamaica)] 63. Hedgpeth, J. W Annotated list of certain marine invertebrates found on Texas jetties. Pp in H. L. Whitten, H. F. Rosene, and J. W. Hedgpeth, ed. The Invertebrate Fauna of Texas Coast Jetties; A Preliminary Survey. Publications of the Institute of Marine Science, University of Texas 1: [Gulf species: Squilla empusa (Texas)] 64. Hedgpeth, J. W An introduction to the zoogeography of the northwestern Gulf of Mexico with reference to the invertebrate fauna. Publications of the Institute of Marine Science, University of Texas 3(1): , figs [Gulf species: Squilla empusa (Gulf of Mexico, Texas)] 65. Hedgpeth, J. W Bottom communities of the Gulf of Mexico. Pp in P. S. Galtsoff, ed. Gulf of Mexico, Its Origin, Waters, and Marine Life. Fishery Bulletin 89. Fishery Bulletin of the Fish and Wildlife Service, Volume 55, Washington, D.C. [Gulf species: Squilla empusa (Gulf of Mexico, Texas)] 66. Hernández- Aguilera, J. L Comparación de la fauna carcinológica (Estomatopoda y Decapoda) de cinco arrecifes del Golfo de México. In: Reunión Indicativa de Actividades Regionales Relacionadas con la Oceanografía (Golfo de México y Mar Caribe mexicanos). Veracruz, Ver., de Noviembre de [Gulf species: Neogonodactylus bredini / wennerae complex (Mexico: Veracruz); Neogonodactylus oerstedii (Mexico: Veracruz, Campeche); Pseudosquilla ciliata (Mexico: Veracruz)] 67. Hernández- Aguilera, J. L., and A. M. Hermoso- Salazar Algunos estomatópodos de la costa este de México con la descripción de una nueva especie (Crustacea: Stomatopoda). Investigaciones Oceanográficas, serie B, 4(2): [Gulf species: Nannosquilla candidensis:

12 Reaka et al. ~ 911 and varieties of Crustacea Stomatopoda in the Indian Museum. Records of the Indian Museum 6: [Gulf species: Alima hildebrandi; original description as A. hieroglyphica, Gundlach s Cuban record)] 75. Latreille, P. A Squille, Squilla. Encyclopédie Methodique Histoire Naturelle..., 10: [Gulf species: Coronis scolopendra: original description] 76. Lee, B. D., and W. N. McFarland Osmotic and ionic concentrations in the mantis shrimp Squilla empusa Say. Publications of the Institute of Marine Science, University of Texas 8: [Gulf species: Squilla empusa (Texas)] 77. Lemos de Castro, A Contribuição ao conhecimento dos crustáceos do ordem Stomatopoda do litoral brasileiro: (Crustacea, Hoplocarida). Boletim do Museu Nacional de Rio de Janeiro, nova serie, Zoologia, 128: 1 68, pls. I XVIII. [Gulf species: Meiosquilla schmitti: original description] 78. Linnaeus, C Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis Locis. Edition 12. Stockholm, pp. [Gulf species: Alima neptuni: original description] 79. Lunz, G. R. Jr The stomatopods (mantis shrimps) of the Carolinas. Journal of the Elisha Mitchell Scientific Society 51(1): , figs [Gulf species: Coronis scolopendra (Alabama)] 80. Lunz, G. R. Jr Stomatopoda of the Bingham Oceanographic Collection. Bulletin of the Bingham Oceanographic Collection 5(5): [Gulf species: Gibbesia neglecta (Florida: Sanibel Is.); Squilla edentata edentata: original description (West Florida nne); Squilla empusa (West Florida nne); Odontodactylus brevirostris complex (Florida Keys)] 81. Manning, R. B A checklist of the stomatopod crustaceans of the Florida- Gulf of Mexico area. Quarterly Journal of the Florida Academy of Sciences 22: [Gulf species: Lysiosquilla scabricauda (Florida, Louisiana, Texas, Mexico); Lysiosquillina glabriuscula (Florida); Coronis scolopendra (Mississippi); Meiosquilla quadridens (Gulf of Mexico); Meiosquilla schmitti (West Florida); Gibbesia prasinolineata (West Florida, Cuba: locality not specified); Gibbesia neglecta (West Florida); Squilla rugosa (Tortugas shrimp grounds, Mexico: Gulf of Campeche); Squilla chydaea (West Florida [nne] to Mexico); Squilla edentata edentata (northern Gulf of Mexico off NW Florida & Mississippi Delta, Texas); Squilla empusa (Gulf of Mexico, Louisiana, Texas, Mexico); Eurysquilla plumata (Florida Keys); Pseudosquilla ciliata (Florida); Odontodacoriginal description; Squilla chydaea; Squilla empusa (all Mexico: Gulf of Campeche)] 68. Hernández- Aguilera, J. L., and P. Sosa- Hernández Crustáceos decápodos y estomatópodos en las costas de Tabasco y Campeche. Secr. Mar. Dir. Gral. Ocean. México. Investigaciones Oceanográficas, serie B, 1(8): [Gulf species: Neogonodactylus bredini / wennerae complex (Mexico: Campeche); Squilla chydaea (Mexico: Tabasco, Campeche); Squilla empusa (Mexico: Tabasco, Campeche)] 69. Hernández- Aguilera, J. L., R. E. Toral- Almazán, and J. A. Ruiz- Nuño Especies catalogadas de crustáceos estomatópodos y decápodos para el Golfo de México, Río Bravo, Tamps. a Progresso, Yuc. Dirección de Oceanografía, Dirección General de Oceanografía Naval, Secretaría de Marina, México. 98 pp. [Gulf species: Neogonodactylus bredini / wennerae complex (Mexico: Veracruz, Campeche, Yucatán); Neogonodactylus oerstedii (Mexico: Veracruz, Campeche, Yucatán); Neogonodactylus spinulosus (Mexico: Campeche); Pseudosquilla ciliata (Mexico: Veracruz); Squilla chydaea (Mexico: Tamaulipas, Veracruz, Tabasco, Campeche); Squilla empusa (Mexico: Tamaulipas, Veracruz, Campeche); Lysiosquillina glabriuscula (Mexico: Veracruz); Nannosquilla candidensis (Mexico: Yucatán); Nannosquilla schmitti (Mexico: Yucatán)] 70. Hernández- Aguilera, J. L., and J. L. Villalobos- Hiriart Contribución al conocimento de los crustáceos decápodos y estomatópodos de la Sonda de Campeche. Investigaciones Oceanográficas, serie B, 80-07: [Gulf species: Squilla empusa (Mexico: Campeche)] 71. Hildebrand, H. H A study of the fauna of the brown shrimp (Penaeus aztecus Ives) grounds in the western Gulf of Mexico. Publications of the Institute of Marine Science, University of Texas 3: [Gulf species: Lysiosquilla scabricauda (Texas, Mexico); Coronis scolopendra (Texas); Squilla empusa (Louisiana, Texas)] 72. Hildebrand, H. H A study of the fauna of the pink shrimp (Penaeus duorarum Burkenroad) grounds in the Gulf of Campeche. Publications of the Institute of Marine Science, University of Texas 4: [Gulf species: Lysiosquilla scabricauda (Mexico); Squilla empusa (Mexico)] 73. Holthuis, L. B Nomenclatural notes on eighteenth century Stomatopoda (Hoplocarida). Journal of Crustacean Biology 20(Special number 2): [Revision; Alima neptuni] 74. Kemp, S Preliminary descriptions of new species