Family CHAERILIDAE Pocock, 1893
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1 Family CHAERILIDAE Pocock, 1893 VICTOR FET This monotypic family includes 21 species and four subspecies (including two nominotypical forms), all belonging to the genus Chaerilus Simon, This genus was originally described in Chactidae; it was separated into a subfamily wihin Iuridae by Pocock (1893b), placed as a subfamily in Buthidae by Laurie (1896b), and elevated to a separate family by Kraepelin (1899). Vachon (1974) demonstrated a peculiar trichobothrial pattern of pedipalps in Chaerilidae ("type B") which differs from both Buthidae ("type A") and all other families ("type C"). The family is found only in South and Southeast Asia. Chaerilini Pocock, 1893b: 306, 309 (subfamily); type genus Chaerilus Simon, Chaerilini: Kraepelin, 1894: 140; Laurie, l 896b: 131 (subfamily). Chaerilidae: Kraepelin, 1899: 157; Pocock, 1900d: 53; Kraepelin, 190la: 273; Pavlovsky, 1924: 81; Pavlovsky, 1925: ; Vachon, 1963b: 161; Biicherl, 1967: 114; Vachon, 1972: 2002; Lamora!, 1980a: 440; Tikader & Bastawade, 1983: 317; Francke, 1982d: 73; Francke, 1985: 16; Sissom, 1990a: 113; Nenilin & Fet, 1992: 19; Kovarik, 1998c: 129. Chaerilinae: Kraepelin, 1905: 340; Birula, 1917a: 164; Birula, 1917b: 56; Werner, 1934a: 288 (part); Kastner, 1941: 235 (part); Mello-Leitao, 1942a: 126 (part); Millot & Vachon, 1949: 429 (part). Chaerilaria ("tribus"): Birula, 1917a: 164 (part). Cherilidae (ISS): Pavlovsky, 1925: 195. DISTRIBUTION: ASIA (south and southwest; see Chaerilus). Genus CHAERILUS Simon, 1877 Chaerilus Simon, 1877b: ; type species by monotypy Chaerilus variegatus Simon, SYNONYMS Chelomachus Thorell, 1889: ; type species by original designation Chelomachus birmanicus Thorell, 1889 [= Chaerilus birmanicus (Thorell, 1889)] (synonymized by Kraepelin, 1894: 141). Uromachus Pocock, 1890b: 250; type species by monotypy Uromachus pictus Pocock, 1890 [= Chaerilus pictus (Pocock, 1890)] (synonymized by Kraepelin, 1894: 141). Chaerilus: Karsch, 1879b: 101, 107; Thorell, 1889: 584; Kraepelin, 1894: 141; Laurie, 1896b: 131; Lonnberg, 1897b: 199; Kraepelin, 1899: 157; Pocock, 1900d: 53-54; Kraepelin, 1913: , fig. 2; Birula, 1917a: 164; Birula, 1917b: 56; Werner, 1934a: 288; Kastner, 1941: 235; Vachon, 1953b: 9, fig. 2; Vachon, 1963b: 162, fig. 3; Vachon, 1974: ; Soleglad, 1976b: 298; Tikader & Bastawade, 1983: 318; Francke, 1985: 7, 16; Sissom, 1990a: 113, fig. 3.lD, 3.4, 3.7A; Nenilin & Fet, 1992: 19-20; Kovarik, 1998c: 129. Uromachus: Pocock, 1894c: 91. DISTRIBUTION: ASIA. Bangladesh, India, Indonesia, Malaysia, Nepal, Singapore, Sri Lanka, Vietnam. NOTES: There was no recent revision of this enigmatic Asian genus which includes 21 species and four subspecies (including two nominotypical forms). Redescriptions and keys were published for Sri Lanka by Vachon (1982), for India and Bangladesh by Tikader & Bastawade (1983), and for several species by Fage (1946a). Additional species were recently described from Malaysia (Vachon & Louren90, 1985) and Indonesia (Louren90, l 995j). 323
2 324 CATALOG OF THE SCORPIONS OF THE WORLD Chaerilus agilis Pocock, 1899 Chaerilus agilis Pocock, 1899a: HoLOTYPE: 9 (BMNH), Batu Caves, Selangor, Malaysia. Chaerilus agilis: Flower, 1901: 34; Kraepelin, 1913: 142; Bristowe, 1952: 698, fig. 1-2; Kovarik, 1998c: 129. DISTRIBUTION: ASIA. Malaysia. Chaerilus anthracinus Pocock, 1900 Chaerilus anthracinus Pocock, 1900d: 57. SYNTYPES: o 9 (BMNH), Dalhousie, Punjab, India. Chaerilus anthracinus: Takashima, 1945: 101; Mani, 1959: 12; Perez, 1974: 31; Tikader & Bastawade, 1983: , fig ; Kovaffk, 1998c: 129. DISTRIBUTION: ASIA. India (Punjab). Chaerilus anthracinus anthracinus Pocock, 1900 Chaerilus anthracinus Pocock, 1900d: 57. DISTRIBUTION: ASIA. India (Punjab). Chaerilus anthracinus rufescens Pocock, 1900 Chaerilus anthracinus rufescens Pocock, 1900d: HOLOTYPE: 9 (BMNH), Dharmsala, Punjab, India. Chaerilus anthracinus rufescens: Mani, 1959: 12; Perez, 1974: 31; Tikader & Bastawade, 1983: , fig ; Kovarik, 1998c: 129. DISTRIBUTION: ASIA. India (Punjab). Chaerilus assamensis Kraepelin, 1913 Chaerilus assamensis Kraepelin, 1913: , fig. 3a, b. SYNTYPES: 1 o, 1 juv 9 (?ZSI), Assam, India. Chaerilus assamensis: Takashima, 1945: 101; Perez, 1974: 31; Kovaifk, 1998c: 129. DISTRIBUTION: ASIA. India (Assam). Chaerilus birmanicus Thorell, 1889 Chelomachus birmanicus Thorell, 1889: HOLOTYPE: juv (MCSNG), Rangoon (Yangon), Burma, now Myanmar. Chaerilus birmanicus: Pocock, 1900d: 62-63; Takashima, 1945: 100. DISTRIBUTION: ASIA. Myanmar. Norns: Kovaffk (1998c: 129) listed this species as a junior synonym of C. variegatus Simon, 1877, without a justification. Chaerilus borneensis Simon, 1880 Chaerilus borneensis Simon, 1880b: HOLOTYPE: juv (MNHN), northern Kalimantan (Borneo), now Indonesia. Chaerilus borneensis: Thorell, 1888: 337; Kraepelin, 1901a: 273; Kopstein, 1921: 143 (species unclear).
3 SCORPIONES: CHAERILIDAE 325 DISTRIBUTION: ASIA. Indonesia (Kalimantan). NOTES: K.raepelin (1894: 144) and Kovarfk (l 998c: 129) considered this species a synonym of C. variegatus Simon. Chaerilus cavernicola Pocock, 1894 Chaerilus cavernicola Pocock, l 894c: 91-93, pl. VI, fig. 5-5a. SYNTYPES: 2 ad (? o S? ), I juv (? o) (BMNH), Ngalau Cavern, near Pajacombo, Sumatra, now Indonesia. Chaeriluscavernicola: Kraepelin, 1899: 160; Kraepelin, 1913: 153; Kopstein. 1921: 143; Kopstein, 1923: 186; Giltay, 1931: 18; Wolf. 1937: 534; Kovaffk, 1998c: 129. DISTRIBUTION: ASIA. Indonesia. Chaerilus celebensis Pocock Chaerilus celebensis Pocock, l 894c: HOLOTYPE: juv (BMNH), Luwu, Sulawesi. now Indonesia. SYNONYMS Chaerilus rectimanus Pocock, I 899a: (synonymized by Fage, I 946a: 7'2). SYNTYPES: o S? (BMNH), Singapore. Chaerilus celebensis: Kraepelin, 1894: , pl. II, fig. 56: Thorell, 1894: 15-16; Kraepelin, 1899: 158; Borelli, 1904b: 4; Kraepelin, 1913: 142, ; Kopstein, 1921: 142; Kopstein, 1923: 186; Banks, 1928: 505; Giltay, 1931: 18; Fage, 1933: 27, 29-30; Fage, 1936: 181; Pelt, 1936: ; Werner, I 934a: 288, fig. 378; Takashima, l 945: 99: Fage, I 946a: 71. fig. I; Takashima, 1952: 29; Sissom, 1990a: 114, fig. 3.20; Kovai'fk, 1998c: 129. Chaerilus rectimanus: Flowers, 1901: 34; Kraepelin, 1913: 142; Fage, 1933: 27, 29; Fage. 1936: 180; Takashima, 1945: 100. DISTRIBUTION: ASIA. Indonesia (Billiton, Kalimantan, Sulawesi), Malaysia (Sarawak), Singapore, Philippines, Vietnam. NOTES: Kovarfk (l 998c: 129) listed C. rectimanus as a valid species, without a justification. Chaerilus ceylonensis Pocock, 1894 Chaerilus ceylonensis Pocock, I 894a: HoLOTYPE: S? (BMNH ), Trincomalee, Sri Lanka. Chaerilus ceylonensis: Kraepelin, 1899: 159; Pocock, l900d: 62; Kraepelin. 1913: 143; Takashima, 1945: 99; Vachon, 1982: 102. fig , 76, 78; Tikader & Bastawade, 1983: 326, fig ; Nenilin & Fet, 1992: 19; Kovaffk, I 998c: 129. DISTRIBUTION: ASIA. Sri Lanka. Chaerilus chapmani Vachon & Lourern;o, 1985 Chaerilus chapmani Vachon & Lourern;o. 1985: 10-17, pl. I, fig. 1-16, tab. I. HOLOTYPE: 'i.> (MNHN RS 8479), Leopard Cave, Gunong Api, Gunong Mulu National Park, Sarawak, Kalimantan (Borneo) Island, Malaysia. PARATYPES: IS? (BMNH RS 8477), same locality as holotype; I '.? (MNHN RS 8522; allotype), Cave of the Winds (Lohang Angin); I juv S? (MNHN RS 8476). IS? <BMNH RS 8478), Clearwater Cave, GuaAir Jernih, Gunong Api, Gunong Mulu National Park; Sarawak, Kalimantan (Borneo) Island, Malaysia. Chaerilus chapmarzi: Lourenc;o & Francke, 1985: 5; Lourenc;o, l994a: 183; Kovarik, I998c: 129. DISTRIBUTION: ASIA. Malaysia (Sarawak).
4 326 CATALOG OF THE SCORPIONS OF THE WORLD Chaerilus gemmifer Pocock, 1894 Chaerilus gemmifer Pocock, 1894a: 81. SYNTYPES: 2 2 (BMNH), Sylhet, now Bangladesh. Chaerilus gemmifer: Kraepelin, 1899: ; Pocock, 1900d: 60-61, fig. 18B; Takashima, 1945: 100; Perez, 1974: 31; Tikader & Bastawade, 1983: , fig ; Kovaffk, 1998c: 129. Chaerilus pictus gemmifer: Kraepelin, 1913: DISTRIBUTION: ASIA. Bangladesh. Chaerilus granifrons Kraepelin, 1913 Chaerilus granifrons Kraepelin, 1913: 143, HOLOTYPE:? 2 (?ZSI), type locality unknown,?india. Chaerilus granifrons: Takashima, 1945: 101; Perez, 1974: 31; Kovaffk, 1998c: 129. DISTRIBUTION: ASIA.?India (exact distribution unknown). Chaerilus granosus Pocock, 1900 Chaerilus granosus Pocock, 1900d: SYNTYPES: 4 (?) 2 (BMNH), Mussooree, Uttar Pradesh, India. Chaerilus granosus: Kraepelin, 1913: 143, fig. 4a; Takashima, 1945: 101; Mani, 1959: 12; Perez, 1974: 31; Tikader & Bastawade, 1983: , fig ; Kovaffk, 1998c: 129. DISTRIBUTION: ASIA. India (Uttar Pradesh). Chaerilus hirsti Kraepelin, 1913 Chaerilus hirsti Kraepelin, 1913: 144, SYNTYPES: 1 specimen (?ZSI), Simla, Himachal Pradesh, India; 1 specimen (?ZSI), Dehra Dun, Uttar Pradesh, India; 1 specimen (?ZSI), Nepal. Chaerilus hirsti: Takashima, 1945: 101; Perez, 1974: 31; Kovarik, 1998c: 129. DISTRIBUTION: ASIA. India (Himachal Pradesh, Uttar Pradesh), Nepal. Chaerilus insignis Pocock, 1894 Chaerilus insignis Pocock, 1894a: HoLOTYPE: 2 (BMNH ), Ladak, Kashmir, India. Chaerilus insignis: Pocock, 1900d: 58-59, fig. 17C; Kraepelin, 1913: 144, ; Takashima, 1945: 101; Mani, 1959: 12; Perez, 1974: 31; Tikader & Bastawade, 1983: , fig ; Kovaffk, 1998c: 129. DISTRIBUTION: ASIA. India (Jammu and Kashmir, Uttar Pradesh). Chaerilus laevimanus Pocock, 1899 Chaerilus laevimanus Pocock, 1899a: HOLOTYPE: (sex unknown) (BMNH), Pulo Gaya, British North Borneo, now Sarawak, Malaysia. Chaerilus laevimanus: Kraepelin, 1913: 145, fig. 3c; Kopstein, 1921: 141; Kopstein, 1923: 186; Giltay, 1931: 18; Takashima, 1945: 101; Takashima, 1952: 29; Kovaffk, 1998c: 129. DISTRIBUTION: ASIA. Indonesia (Billiton, Kalimantan), Malaysia (Sarawak).
5 SCORPIONES: CHAERILIDAE 327 Chaerilus pictus (Pocock, 1890) Uromachus pictus Pocock, 1890b: , pl. XIB, fig. 1-lc. HOLOTYPE:?o (BMNH), Sylhet, now Bangladesh. PARATYPE:? '? (BMNH), same locality as holotype. Chaerilus pictus: Pocock, 1894c: 91; Kraepelin, 1894: , pl. II, fig. 42; Kraepelin, 1899: 159; Pocock, 1900d: 59-60, fig. 17B, 18C; Kraepelin, 1913: ; Takashima, 1945: 100; Perez, 1974: 31; Tikader & Bastawade, 1983: , fig ; Kovarik, 1998c: 129. DISTRIBUTION: ASIA. Bangladesh, India (Meghalaya). Chaerilus robinsoni Hirst, 1911 Chaerilus robinsoni Hirst, 1911b: 47~71. HoLOTYPE: '? (BMNH), Bukit Besar, Malay Peninsula, now Malaysia. Chaerilus robinsoni: Takashima, 1945: 101; Kovarik, 1998c: 129. DISTRIBUTION: ASIA. Malaysia. Chaerilus sabinae Lourern~o, 1995 Chaerilus sabinae Lourern;o, 1995j: , fig HOLOTYPE: juv '? (MNHN), ll9 38'E, 4 48'44"S, cave ofguaatas, hills ofmatampa, near Pangkajene, region of Maros, Sulawesi Selatan Province, Sulawesi, Indonesia. Chaerilus sabinae: Louren90, 1996h: 1; Kovarik, 1998c: 129. DISTRIBUTION: ASIA. Indonesia (Sulawesi). NOTES: The originally published (Lourengo, 1995j) type locality (in India) is incorrect; it was corrected by Lourengo (1996h). Chaerilus tricostatus Pocock, 1899 Chaerilus tricostatus Pocock, 1899d: 266. HoLOTYPE: '? (BMNH ), Sadi (Sadiya), northeastern Assam, India. Chaerilus tricostatus: Pocock, 1900d: 59-60, fig. 17 A; Henderson, 1913: , fig. 2; Kraepelin, 1913: 142; Takashima, 1945: 101; Perez, 1974: 31; Tikader & Bastawade, 1983: , fig ; Bastawade, 1985: 260; Kovarik, 1998c: 129. DISTRIBUTION: ASIA. India (Arunachal Pradesh, Assam, Meghalaya). Chaerilus truncatus Karsch, 1879 Chaerilus truncatus Karsch, 1879b: SYNTYPES: 2 specimens (ZMB 3055), "Himalaya", India. SYNONYMS Chaerilus margaritatus Pocock, 1894a: (synonymized by Kraepelin, 1913: 143). HOLOTYPE: '? (BMNH), Dehra Dun, Uttar Pradesh, India. PARATYPE: 0 (BMNH), same locality as holotype. Chaerilus truncatus: Simon, 1880b: 381; Kraepelin, 1894: , pl. II, fig. 54; Kraepelin, 1899: 160; Pocock, 1900d: 63; Simon, 1901: 79; Kraepelin, 1913: ; Fage, 1936: 180; Vachon, 1940b: 248, fig. 11; Takashima, 1945: 100; Page, 1946a: 71; Perez, 1974: 31; Moritz & Fischer, 1980: 325; Tikader & Bastawade, 1983: , fig ; Kovarik, 1998c: 129. Chaerilus margaritatus: Pocock, 1900d: 58, fig. 18A; Takashima, 1945: 100; Perez, 1974: 31. DISTRIBUTION: ASIA. India (Himachal Pradesh, Uttar Pradesh), Vietnam.
6 328 CATALOG OF THE SCORPIONS OF THE WORLD Chaerilus variegatus Simon, 1877 Chaerilus variegatus Simon, 1877b: , pl. III, fig. 13. HoLOTYPE: 1 specimen (MNHN), Java, Indonesia. Chaerilus variegatus: Karsch, 1879b: ; Simon, 1880b: 38 1; Thorell, 1888: 337; Kraepelin, 1894: , pl. II, fig. 53, 55, 57-58; Laurie, 1896b: 122; Kraepelin, 1899: 160; Pocock, 1894c: 90-91, pl. VI, fig. 4-4a; Kraepelin, 190la: 273; Werner, 1902: 604; Kraepelin, 1913: 143, ; Kopstein, 1921: ; Kopstein, 1923: 186; Pavlovsky, 1924: 80-81; Pavlovsky, 1925: 149; Kopstein, 1926: 109; Banks, 1928: 505; Giltay, 1931: 18; Fage, 1933: 27; Werner, 1934a: 288, fig. 377; Kopstein, 1935: ; Kastner, 1941: 235; Takashima, 1945: ; Page, 1946a: 71; Dammerman, 1948: 71 ; Takashima, 1952: 29 ; Kovai'fk, 1992a: 185; Kovai'fk, 1998c: 129(part). DISTRIBUTION: ASIA. Indonesia (Banka, Java, Kalimantan, Krakatau, Nias, Sebesi), Indochina. NOTE: This species is possibly listed in Herbst (1800: 57, fig. 1) as Scorpio australasiae (nee Fabricius, 1775). Chaerilus variegatus variegatus Simon, 1877 Chaerilus variegatus Simon, 1877b: Chaerilus variegatus variegatus: Kopstein, 1935: 209. DISTRIBUTION: ASIA. Indonesia, Indochina. Chaerilus variegatus nigricolor Pocock, 1899 Chaerilus variegatus var. nigricolor Pocock, 1899a: 419. HOLOTYPE: (sex unknown) (BMNH), Protjat, eastern Java, Indonesia. PARATYPES: o <:i? (BMNH), Kogok, western Java, Indonesia. Chaerilus variegatus nigricolor: Kopstein, 1921: 143; Giltay, 1931 : 18; Kopstein, 1935: ; Vachon & Abe, 1988: 26; Kovai'fk, 1998c: 129. DISTRIBUTION: ASIA. Indonesia (Java, Krakatau).
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