DISTRIBUTION AND RECORDS OF COLORADO TRICHOPTERA

Transcription

1 DISTRIBUTION AND RECORDS OF COLORADO TRICHOPTERA ABSTRACT-Previous published records, new records, dramage, habitat preference, altitudinal range, and adult collection dates are presented for 176 species in 15 families of Colorado Trichoptera. Of this listing 53 species are new records for Colorado. A comprehensive publication on the systematics, distribution, ecology, and records of Colorado Trichoptera is nonexistent. No major caddisfly monographs or species listings similar to that of Ross (1944) for Illinois, Anderson (1976) for Oregon, Roemhild (1982) for Montana or Denning (1956) for California have been published for Colorado. Only one statewide family listing, that for the Hydroptilidae, is available (Nerrmann, 1982). Most records of Colorado caddisflies are scattered throughout the literature and represent collections made from small geographic regions of the State. The purpose of our work is fourfold: 1) to compile the list of literature sources necessary for regional trichopterists to conduct future studies in Colorado, 2) to provide a view on the validity of the Colorado caddisfly records reported in earlier publications, 3) to report new records for Colorado caddisflies, and 4) to provide distributional, altitudinal, and ecological data for Colorado Trichoptera. Colorado is a large state, 103,483 miz (268,021 km2), and topographically diverse. The lowest point in the state, 3350 ft (1020 m), is in the bed of the Arkansas River in eastern Prowers County. Mt. Elbert in Lake County is the highest peak in the state at ft (4399 m) and second highest in the 48 contiguous states. Colorado with a mean altitude of 6800 ft (2073 m) is also the highest state of all 50 states. These topographic extremes along with great climatic variations, steep stream gradients, intermittent stream flows, hot and warm springs, closed drainage basins, and diverse geological chemistries, combine to create many diverse aquatic habitats. The headwaters of four river systems, the Colorado, Rio Grande, Platte, and Arkansas, are located in the mountains of Colorado (Fig 1). MATERIALS AND METHODS-In reviewing previous publication records, only the studies l~sting a collection locality which could be located by county were included. The exception was the case where the record (without a collection locality) was the only Colorado record available in the literature. Publications that cited previous Colorado records but did not supply additional collection data were excluded. A special effort was made to locate specimens of published records for which no specif~c locality data were given, and of species that had not been collerted since being reported in the literature. If specimens of a questionable record could not be located but the species was reported from any of the states adjoining Colorado, the species was retained in our listing and its status documented. Special efforts were also made to assign all the published records to a county. Many of the older locality records were vague, for example, southwest Colorado or Colorado mountains, or simply, Colorado; so county assignment was not always

2 122 The Southwestern Naturalrst bol 31, no 1 poss~ble. \Ve made the assumption that South Park, the locality for several old reco~dr, ii cntirel\ within Park County. Dodds and Hisaw's (1925b) localitirs were divided into two counties basrd on 2438 ~n altitude Every record above 2438 m was cons~dered Gilpin Ciounty and rvcr) record below 2438 m considered Boulder County. The actual Boulder-Cilpin County line rises from 2271 m to 2463 m. Ward's 11976) records from the South Platte River were listed in both Douglar and Jefferson countirs as the river in the collection area is the county line 'I'housands of Colorado speclmenr, primarily adults, were examined by us or other cadd~sfl) spec~alists prlor to preparation of this publicatron. Thrsr specimens came prirnar~ly from collections of Herrmann, who rollected extensively throughout Colorado since Ruiter. M-ho rolle(ted northcentral and northwestern Coloratlo since 1975; the Illino~s Natural H~story Survr\ (INHSJ. Champaign, IL; the Llriited State5 Natlorlal Museum (L'SNM), Smithson~an Institution, tvashirigton. D.C ; and the Peabody Museum, Yale I!n~versity, New Haven, C:X In add~tiot~. man! other rpecimens from individual and institutional collections were examined. L'nitrd State5 Geological Survey (L1.S.G.S.) topographic quadrangle maps (standard 7% nlinute. 15 minute, and county series) wrrr usrd to establish collection site coordinatrr, alt~tudes, and drsrriptions. Pre(ise drlin~tion of aquatic habitats arr neither easily grnerated nor pra<-tl(allt dppllrd. l.ot~c habitats range from large rivers with ~onsiderable volume of flowlr~g surfacr wa:t~ to \mall sreps w~th only enough d~srharge to wet the surrounding soil. Springs. In contrast to seeps, ar' usually chalarteri7ed b) relative consrancy of watel (hem~stry, discharge, and teniperature. The trim "stream" tefers to an intermrd~ate $Ire bod) of running water In a surfacr rhannrl, and includes the terms "creek" and "brook." I.akes and ponds are the p~~nlnry tbprs of Ient~c hab~tats; the former are characteri~ed by large limnet~c and profundal rones and a small Iitto~alzone. and the later by the conbersr set ot zonal proportiorls. In dercribing a sprcles hah~tat rvr grnerally llsted the lotir or lentic descriptor that appears as part of a collection site's proper name on I..S.G.S maps. For rxample, the Colorado River descrlpu)~ for habitat tspe is listed a5 "r~rer", Texas Crrrk ar "stream", and Grand Lake ar "lake." Reference ir made to Pennak' ) Iirnnological 7or1rs of northcentral Colorado; they are plains 1020 m to 1700 m, foothills I700 m to 2500 rn. montane 2500 In to 3200 m. and alpinr (>3200 m). D~scuasion of a rperlrs alutudlnal distribution 1s farilrtatrd by makirlg rrferrr~(r lo these zones. The range ot aluuidr appropriate to each zonr for a giben rrgion of Colorado ma) ~ar) contiderably due to man\ Intpractlng lactors: th~refore, this ronation srhrme 18 appltrd quite RESULTS AND DISCUSSION-In the literature we found Colorado records for 146 species of Trichoptera. Three species, Apatania stigmatella (Zetterstedt) 1840, Limnephilus ectus Ross 1941, and Pseudosteraophylax edwardsi (Banks) 1920 have been reported to be Colorado records that are in error (Nimno, 1971; Anderson, 1976; Anderson, 1974). Three species, Lzmnephilus concolor Banks 1899, Lepidostoma stigma (Banks) 1907 and Nectopsyche coloradensis (Banks) 1899 are reported a5 either unassociated females or damaged, and of questionable species validity (Flint, 1966; Ross, 1946; Haddock, 1977). Another limnephilid, Limnephilus clausus Banks 1924, was considered by Ross and Merkley (1952) to be a synonym of I>. picturatus MacLachlan Hydropsyche frisoni Ross 1938 and Khyacophila bifila Banks 1914 were reported from Colorado by Harris and Lawrence (1978). Personal communication with Harris (1981) has confirmed the error in these two records. Dodds and Hisaw (1925b) reported Rhyacophila atrata Banks 1911 and Hydropsyche.scalari~(Hagen) 1861 from Colorado; these two records are probably a result of incorrect identifications. Ross (1944) cited the known distribution of R. atrata as eastern North America. Flint et al. (1979) discussed the identification problem of H. scalaris and did not include Colorado in the known diatribution. Ward ( 1975) reported Lepzdostoma rayneri Ross 1941 and Agapetus taho Ross I947 as occurring in Boulder County. Colorado; ar

3 November I986 Herrmann et a].-distribution and Records of Trichoptera 423 these species can be confused with Lepidostoma ormea Ross 1946 and Agapetus boulderensis Milne 1936 respectively, and Ward's specimens cannot be located at this time, we will delete these two species from the Colorado list until such time as specimens can be located. Hagen (1875) reported Pycnopsyche divergens (Walker) 1852 from Colorado. Hagen stated that the specimens were fragments and were possibly two species. Until specimens can be located we have deleted this species from the Colorado list. Dodds and Hisaw (1925b) cited Limnephilus sansoni Banks 1918 from one collection at about 2700 m elevation. Since this collection could not be located and substantiation not possible, and since this species has not been reported from Utah (Baumann and Unzicker, 1981), Oregon (Anderson, 1976), or Montana (Roemhild, 1982), we deleted the record of L. sansoni. Rhyacophila tucula Ross 1950 was reported from Boulder County by Ward (1981) and Larimer County by Short and Ward (1980a) from larval identifications. Adults are not available for positive identification, therefore this species has been deleted from our Colorado list. Two species, Brachycentrus occidentalis Banks 1911 and Rhyacophila oreta Ross 1941 were identified as larvae (Canton and Chadwick, 1983a; Canton, 1984). Some uncertainty exists, therefore, for these two records. Brachycentrus occidentalis is commonly collected by Ruiter and Herrmann; R. oreta adults from Colorado have not been collected by us. Colorado records of Ochrotrichia unio (Ross) 1941 in the INHS collection and Nectopsyche albida (Walker) 1852 in the USNM collection were deleted from our listing because the former species' record was based on seven larvae and the latter on three unassociated females. Nimmo (1971) cited Colorado records for Apatania shoshone Banks 1924 and Limnephilus rhombicus (Linnaeus) 1758; however, they were deleted from our list because they could not be substantiated. Searches for Hesperophylax oreades Saether 1970 at the type locality and elsewhere in Boulder County and Colorado have proven fruitless. Parker (1984) felt that this species is actually Psychoronia costalis (Banks) Schmid (1954) reported Asynarchus montanus (Banks) 1907 from Colorado without giving a collection locality; both A. montanus and A. circopa (Ross and Merkley) 1952 were considered A. nigriculus (Banks) 1908 by us. These considerations reduce the number of what we feel are valid published species records from Colorado to 123 species. Distribution Records for Colorado Trichoptera-In the following list families, genera, and species are listed alphabetically using the classification scheme of Wiggins (1984). Each species, with type locality, is cited first, followed in order by previous published county records, new county records, river drainage basins, habitat, altitudinal range in meters above mean sea level, extremr dates of adult collection, and for most species, additional commentary. If a species has been collected from only one site or on only one date, one value appears under the heading of altitudinal range or adult collection date. Species reported as new Colorado records are preceded by an asterisk. BRACHYCENTRIDAE Amrocentrus aspilus (Ross) Type I ocality (T.L.) Wyoming, Pinedale. Previous county records: Jackson (Denning, 1948c), Larimer (Denning, 1948~). New county records: Routt.

4 The Southwestern Naturaizsl

5 November I986 Herrmann et al.-distribution and Records of Trichoptera Drainages: North Platte, South Platte, Yampa. Habitat: rivers, streams. Altitudinal range: 2042 to 2662 m. Adult collection dates: 19 July to 17 August. Commentary: In Colorado this species appears to be restricted to the rivers and streams bordering Wyoming. Brachycentrus americanus (Banks) T. L. New Hampshire, Franconia. Previous records: Boulder (Banks, 1907a; Dodds and Hisaw, 1925b; Flint, 1984; Mecom, 1970, 1972a and b; Ward, 1975 and 1981), Custrr (Flint, 1984), Dolores (Canton and Chadwick, 1983), Douglas (Ward, 1976), Eagle (Flint, 1984). Grand (Banks, 1907a), Gunnison (Stanford and Ward, 1981), Jefferson (Fllnt, 1984; Ward, 1976), Larimer (Flint, 1984), Rio Blanco (Gray and Ward, 1977), Routt (Canton and Ward, 1978; 1981a and c), San Miguel (Flint, 1984), Teller (Short et al., 1978). New records: Chaffee, Clear Creek, Conejos, Costilla, El Paso, Fremont, Garfield, Gilpin, Hinsdale, Huerfano, Jackson, Lake, La Plata, Las Animas, Park, Pitkin, Saguache, Mineral, Montrose, Ouray, Summit. Drainages: Arkansas, Colorado, Gunnison, North Platre, Rio Grande, South Platte, White. Habitat: rivers, streams. Alt~tudinal range: 1606 to 3048 m. Adult collection dates: 17 April to 6 September. Commentary: Denning (1983) compared the male and female of this species with B. occ~dentalts Banks 1911 making future incorrect identifications of adults of the two species unlikely. Flint (1984) wrote keys to adult males and larvae for the species of Brachycentrus in North America. Brachycentrus occidentalzs Banks T. L. British Columbia, Bon Accord. Prev~ous records: Boulder (Flint, 1984), Montrzuma (Canton and Chadwick, 1983a), Montrose (Flint, 1984), Rio Grande (Flint, 1984), and Routt (Flint, 1984). New records: Chaffee, Eagle, Fremont, Garfield, Gunnison, Larimer, Mineral, Park. Rio Blanco. Dra~nages:Arkansas, Colorado, Eagle, Rio Grand?, South Platte, Yampa. Habitat: rlvers, streams. Altitudinal range: 1615 to 2926 m. Adult collection dates: 16 April to 6 July. Commentary: In Colorado th~s species is usually morr common below 1829 rn wh~le B. amerzcanus more common above 1829 m; B. occcdentalzs also appears to emerge earlier. *Mzcrasema bactro Ross T. L. Oregon, Strawberry Camp. New records: Chaffee, Eagle, Garfield, Pitkin. Dra~nages: Arkansas. Colorado, White. Habitat: streams. Altitudinal range: 2103 to 2896 m. Adult collection dates: 24 July to 26 July. Commentary: Anderson (1976) reported Oregon emergence from early May through June. GLOSSOSOMATIDAE Agapetus boulderenszs Milne T. L. Colorado, Boulder. Previous records: Boulder (Milne, 1936; Ward, 1981), Routt (Canton and Ward, 1981a and c). New records: Chaffee, Conejos, Custer, Douglas, Grand, Gunnison, Hlnsdale, Jefferson, Larimer, Mineral, Montezuma, Rio Grande, Teller. Drainages: Arkansas, Colorado, Dolores, Gunnison, Rio Grande, South Platte. Altitudinal range: 1975 to 2749 m. Adult collection dates: 3 August to 2 October. Commentary: Our data show this species to be more widespread in Colorado than previously reported. Anagapetus debalzs (Ross) T. L. Utah, Logan Canyon. Previous records: Douglas (Ross, 1951), El Paso (Ross, 19511, Larimer (Ross, 1951). New records: Chaffee, Clear Creek, Garfield, Mineral, Rio Blanco. Drainages: Arkansas, Rio Grande. South Platte, White. Habitat: springs, streams, rivers.

6 426 The Southwestern Natural~sl Altitudinal range: 1896 to 3060 ni. Adult collec tlon dates: 4 June to 1 September. C:ornmrntary: This species IS widespread In Colorado but ieldoni collected in largr numbers. Culoplzla cantlza (Ross) 'L I.. Wyorning. Parco. Preb~ousrecords: unspecrfred localrt) (Fl~nt, 1974a). NI.W rr~ords: MES~, Moffat. I)rairr;~geb: Colorado, Yampa. Habitat: ~iirrs. Altrtudrnal range: 1494 to 1801 In..idult iollert~on dates. 3 August to I I August. Commentary: This specres appear5 to be restrictvd to the west s~de of the continental d~\idr rn Colorado. (:ulopt~/a thoraczca (Ross) I.. Wyomlng, Boulder. Prr\rou$ records: unspec~tird locality (Fl~nt. 1974a). :l'rw rrcords: Clear Creek. Eaglt.. Garrield, Grand, Jefferson. Summit. Ilrainagcs: h~kansas. Colorado. South Plattr Hab~tat:strra~ns,rivrrs..ilt~tud~nal langr: 1615 to idult (ollrc tlori datrs: li July to 10 August. C:~~mmerltary: Nc~rth(entral Colorado countles are the only ones from whrch this speclea h,rs been collectc.d. However, Walt7 and McCafferty (1983) recently repol ted it from New Mex~co. *C;lusrosonla alasten~r Banka, I' L. Alaska, Popof Island. New records: Clear Creek. Ilra~nages: 'i(~rllti Plattc. Itahitat: itrt.anis. :\ltitudinal range: 2817 m. :Adult iollett~on date?: 21 June. (.~)mrnentdr\:nr~nmo(1974) found this hpecles In a w~de rarlrt) of lotii trab~tats rn hlbrrta. (;losrosu~na paruulurn Banks T. L Nrw Mexico, Petoh. Prc\rou\ rrcords. Boulder (Ward, 1981). Ncl\ record,:.ir~huleta, Chatfee. Clrar Creek, Ckjnejos, Douglas, Eagle, Frernont, Garfield. (.rand, Gunn~son. Jackson, Jrfferson, L.akr, La Plata, Larlrne~, Mlneral, Llontrose, Park, Pitl~n,Rio Rlanco, Saguarhe, Surnm~t, rt.llr~ Ilra~nages: An~mas, :irkansac. Colorado, Correjos. Dolorrs, Gunnison, Rro Grandr, San Juan. 5i1uth Piattr. Ll'hitr Habltdt: rlwri. :\ltitudrnal rangr: ~ I I291 1 In. t\dult collec (Ion ddtrs: 6 April to 20 Octohcr. C:onirnentar\: This species 1s ~(~despread ar~d common in Colorado. (;loa~osorna tielorla Roas L. Washington, Centralla PIC\ ious records: Grand (Swegrnan and Ferrington, 1980). h-cw ~rc~~rds, none. 1)ralnagt.c C;olorado. Habrtat, li\er\..\ltitudinal mnge: 2590 m. Adult collrction datec: 2 August. (;ommentar): This record was rnade on the basis of one male collected at Wind~ng R~va Campground just west of the west entrance to Rocky Mountarn National Park. (;lorsosoma uentrale Banks T. L. New Mexrco, East Las r'egas. Preb~ous records: Boulder (Dodds and Hisaw. 1925h), Gilpin (Dodds and Hlsaw, l925h). RIO Blarico (Gra) and Ward, 1977), Ruutt (Carrton and Chadwirk, 1983b; Canton and Ward, l981a arid cj. Nrw rrcords: rlrchuleta, Chaffee, Clear Creek, Custer, Douglas, Eagle. El Paso, Frrrnont, Garf~eld, Grand, Huerfano, Jefferson, Larimer. Park, Pueblo. Drainages Arkansas, Colorado, San Juan, South Plattr. Habitat: rtlrams. :\Itirud~r~al rang(, to 2691 m. i\dt~lt cullvct~on dates: 26 February to 3 September (:t~mmentarv:thrs specie? ha\ a narrower alt~rudrnal rangr than G,pan,ulum.

7 November I986 Herrmann et a1.-distribution and Records of Trichoptera 427 Glossosoma uerdona Ross T. L. Wyoming, Pinedale. New records: Clear Creek, Custer, El Paso, Garfield, Grand, Jefferson, Lake, La Plata, Larimer, Pitkin, Saguache, Summit. Drainages: Arkansas, Colorado, San Juan, South Platte. Habitat: streams. Altitudinal range: 1875 to 3094 m. Adult collection dates: 8 May to 23 August. Commentary: This species is not unexpected in Colorado since Ross' (1938b) holotype male 1s from Wyoming and five of his paratype males are from Utah. Protoptila coloma Ross T. L. California, Hot Creek. Previous records: Grand (Ross, 1941 ). New records: Larimer. Drainages: Colorado, South Platte. Habitat: rivers. Altitudinal range: 1615 to 2423 m. Adult collection dates: 16 July to 4 August. Commentary: Roemhild (1982) found this species commonly in Montana intermountain streams and rivers. It is uncommon in Colorado. *Protoptila erottca Ross T. L. Wyoming, Parco. New records: Garfield, Moffat. Drainages: Colorado, Yampa. Habitat: rivers. Altitudinal range: 1615 m. Adult collection dates: 3 to 20 August. Commentary: This species has been collected only from sites west of the continental divide in Colorado. HELICOPSYCHIDAE Heltcogsyche borealis (Hagen) L. Canada, St. Lawrence River. Previous records: Boulder (Dodds and Hisaw, 1925b; Mecom, 1972a and b; Ward, 1981) New records: Archuleta, Fremont, Garfield, Huerfano, Larimer, Moffat, Park, Pueblo. Drainages: Arkansas, Colorado, San Juan, South Platte, Yampa. Habitat: streams, rivers, springs. Altitudinal range: 1494 to 3109 m. Adult collection dates: 28 February to 7 September. Commentary: This species is more common in Colorado than previously reported HYDROPSYCHIDAE Arctopsyche grandis (Banks) T. L, southwest Colorado. Previous records: Boulder (Mecom, 1972a and b; Wallace, 1975; Ward, 1981), Douglas (Ward, 19761, Grand (Short and Ward, 1980b), Gunnison (Allan, 1975; Stanford and Ward, 1981), Huerfano (Wallace, 1975), Jefferson (Ward, 19761, Pitkin (Wallace, 1975). Routt (Canton and Ward, 1977; 1978; 1981a and c), Summit (Milne and Milne, 1938; Banks, 1943). New records: Chaffee, Clear Creek, Conejos, Eagle, El Paso, Fremont, Garf~eld, Gilpin, Larimer, Las Animas, Ouray, Park, Rio Grande, San Juan, Teller. Drainages: Animas, Arkansas, Colorado, Gunnison, San Juan, South Platte, Yampa. Habitat: rivers, streams. Altitudinal range: 1913 to 3036 m. Adult collection dates: 10 May to 10 October. Commentary: This large hydropsychid is widespread and easily collected with an ultraviolet 11ght. "Cheumatopsyche campyla Ross T. L. Illinois, Momence. New records: Garfield, Grand, Jefferson, Moffat. Drainages: Colorado, South Platte, Yampa. Habitat: rivers. Altitudinal range: 1615 to 1920 m. Adult collection dates: 1 July to 7 August. Commentary: Ross (1944) cited this species as being "...most abundant through the Corn Belt states but is found over almost all other pa ts of the continent...." In Colorado this species appears to be restricted in altitude.

8 *Cheurnatopsychr enonzs Rosa 'r 1.. \Yyom~ng, Parco. New records: Archulcta, C:onrjos, Costilla, Del [a, Faglr. Garfield, Grand, Mesa. Moffat 1)rainages: Colorado, Conejr~s, C.unnisor~, RIO Gtandr, San Juan, Yampa Habitat: rivers Altrtud~rralange: to 2333 In..Adulr collrct~un dates: 13 Julv to 20.August. (:orntnentar): In (.olorado this q)ccirs is rvidesprrad wtsst of the ~ontlnental dir~de but spot[) ~n occurrrnre. Cheunzalopcychr graczl~s (Banks) T. I.. Colorado, Fort Colllr~r. Pr~.\~ous records: Boulder (Dodds and Hlsaw. 1925b1 Ncw records Jackson. Park. I)minagr\: North Platre, South Plattr Habitat: $trt.arn,. tlrrrs..alt~tud~nal range: to 270i rrl..adult collectlorl datcs. 3 to 8 Jul). C.ornrncntar): Th~r speclrs 15 corntnon in thr cen~~al and ea5terrr IT.S. but no1 common In Colorado or other parts of the west and southwest (Ciordon. 1974). Chzumnfopsyihe prtllll (Banks) r. L. Mich~gan, Agricultural C:ollrgc. Prrv~oiie records..arapahoe (Gordon, 1974), Boulder (Ward, 19811, Clrar Creek igordon. 197l). New rl-cnrds Berit. Crowle). I)cn\er, 1)ouplas. El Paso. Jefferson. K~owa. I.arlmer, Motga~~. Par k, Prv\\.crs, Pueblo, Rorltt, Sedgwick, Weld, Yampa. Ilra~nages:.\rkdns;is, Colorado, Rrpubllcan, South Platlc. Hab~tat. ~trr'uns, rl\rrc..alt~tud~nal range to 2138 m. ;\dult collrct~on date\: 21.April to 3 October. (.(~mrnrnta~\: Illst~nction could not br made betweell specimens ~dentified as C. stntthz Gordon and (:, prtl~ti, hrncr, all specimens were considered C. pettllz. 'I'hle sptarles re IJ;IIUI ul,lrl\ (olnrnon In the :\I-hansas and South Platte R~vers of eastern Colorado. '(.heumalopsyrhr waba\ha Drnrllrig 'r L. Minnesota, \Vabasha. Nc\* 1rcr~rd5:Moftat. Dldlnagri: Yampa Hd111tdt. r~\rr.alt~rildlnal Iangr: 1789 rn. Adult cc~llcctio~l dates: 'i Jul) C.onrn~rrrt,tr). This 1s a slgnif~cant southerl) range extenslon. Gordon (1974) Itctrd th~ sprclrs iron^ Olrgon and M~nnesota. *Hydrop.i)"hr bronta Ross TI I.. Michigan. Bronsori. Nr\r records: Pueblc]. Drdlnagec. Arkansai. H;lbttat. rivcr Altrtud~nal range: 1431 In. Adult collection dates: 3 October C:otnmcntar\ This speclee was previously reported from IYyomlng lro\\, 1944) and Arkansas i I'rtrlcker et al., 1970). Hydropsychr rockerell1 Banks I. New Mexico, Pecos. I'rev~ous records. Routt (Canton and Ward, 1981a and c) \lr.u recu~ds: Arapahoe, Archulrta, Boulder, Chaffee, Conejos. Custer, Delta, Douglas, Eagle, F~rrnor~~, Garfield, Grand, H~nsdale, Jackson, Jefferson, I.akt>, Larlmrr, Las Animas, Mesa. Moffat, Montrose. Morgan. Park, Purblo, Rio Rlanco, Saguache. Drainages: Animas, Atkanaas, Colorado, Gunn~son, North Plattr, Rio Grarrde. San Juan, South Platte Habrtat: ctrrarns, rlvers..alt~tudinal range to 3109 m. Adult collrction datrs: 26 May to 7 September. Cixnmentar): In Colorado thls spec-ies is common and widespread in the foothills and montane 7;~nr5 both east and west of the contlriental div~de. 'Hvdropiychr confusa (Walker T 1.. Canada, Northwest T'erritorlt,\ Nrr\ rrc(jrds, hloffdr. Dranagrs. IYhtte

9 November I986 Herrmann et al.-distribution and Records of Trichoptera Habitat: rivers, streams. Altitudinal range: 1828 m. Adult collection dates: 12 July. Commentary: Nimmo (1981) synonymized H. separata Banks 1936 under H. confusa. Our only Colorado record of this species is from a site 2.5 miles (4 km) east of Dinosaur; the male and female specimens of this record were identified by D. G. Denning. Baumann and IJnzicker (1981) reported it from Utah. Hydropsyche morosa Hagen T.L. Canada, St. Lawrence River. Previous records: Larimer (Banks, 1905). New records: Garfield, Grand. Drainages: Colorado, South Platte. Habitat: rivers. Altitudinal range: 1615 to 2550 m. Adult collection dates: 24 July to 8 August. 0 Commentary: Until recently, western populations of this species were named Hydropsyche bifida Banks Schefter and Unzicker (1984) synonyniized H. bifida Banks with H. morosa Hagen. Hydropsyche occidentalis Banks T. L. Washington, Pullman. Previous records: Boulder (Dodds and Hisaw, 1925b; Mecom, 1970, 1972a and b), Gilpin (Dodds and Hisaw, 3925bj, Routt (Canton and Ward, 1981a and c). New records: Adams. Arapahoe, Archuleta, Bent, Crowley, Delta, Denver, Douglas, Eagle, El Paso, Fremont, Garfield, Gunnison, Huerfano, Jefferson, Larimer, Las Animas, Mesa, Moffat, Montezuma, Montrose, Otero, Park, Pitkin, Prowers, Pueblo, Teller, Weld, Yampa. Drainages: Arkansas, Colorado, Dolores, Gunnison, Republican, San Juan, South Platte, Yampa. Altitudinal range: 1085 to 2941 m. Adult collection dares: 25 May to 18 October. Commentary: This species' altitudinal range overlaps that of H. cockerellr, although H. occ~dentalrsoccurs in the lowest portions of the plains zone in eastern Colorado. The lowest altitude reported for H. occidentalis outside Colorado hy Harris and Lawrence (1978) was 805 m. Hydropsyche oslarr Banks T. L. southwestern Colorado. Previous records: Boulder (Ward, 1981), Chaffee (Flint and Herrmann, 1976), Rio Blanco (Gray and Ward, 1977), Routt (Canton and Ward, 1981~). New records: Custer, Douglas, Eagle, El Paso, Fremont, Garfield, Grand, Gunnison, Jefferson, La Plata, Larimer, Moffat, Montezuma, Park, Teller. Drainages: Arkansas, Colorado, Dolores, Gunnison, San Juan, South Platte, Yampa. Altitudinal range: 1615 to 2895 m. Adult collection dates: 10 June to 1 September. Commentary: Fischer ( ) reported the geographical range from British Columbia to California and east to New Mexico and Utah. Hydropsyche simulans Ross T. L. Illinois, Mount Carmel. Previous records: unspecified locality (Ross, 1944). New records: Logan, Washington, Weld. Drainages: South Platte. Habitat: river. Altitudinal range: 1053 to 1406 m. Adult collection dates: 7 July to 17 August. Commentary: Our Colorado records are from the South Platte River in northeastern Colorado; it is common in the Corn Belt (Ross, 1944) of the midwest. Parapsyche almola Ross T L. Washington, Almota. Previous records: unspecified locality (Smith 1968a). New records: Clear Creek, Douglas, El Paso, Garfield. Drainages: Colorado, Rio Grande, White. Altitudinal range: 1981 to 2804 m. Adult collection dates: 18 to 26 July.

10 430 The Southwestern Naturalnct \<>I '31, no 4 Commentary: In Montana Roemhild 11982) reported th~s sprcirs from cold streams and rivers of thr alp~rre and intermountain zones. 'Parapsv<he eisis Milne : L British Columb~a, Cultus Lake. Srw records. Garfield, Pitkin. Rio Blanco, Rio Grande. Ilrainages: C;olorado. Rio Grande, M'hitr. Hab~tat:strrams, rivers. Altitndinal rangc: 2408 to 3200 m..4dult collection dates: 23 June. (.ornrneritar). Thls specirs is uncommon ~n C:olorado. Srrilcrldra fasclalelln McLachlar~ T I.. Tkxas. P~e\iousrecords: Adams (Flint. 1971bl. New records: none. llra~nagrs:5011th Platte. Habitat: ptrrarn.\ltitud~nal larrge: 1670 m. :\dult collrction dates: 4 tu 5 Juli (.ommentar): Ihr on], rtrord of thls species from C:olorado IS that of one fenlale from LVatk~ns ifllnr. 1974b). Boxelder Crcek is an intrrmittent stream passing through the town of Uktk~ns on thc Colorado Piedmont of the Great Plains. Habitat conditions 5imilar 11, thosr described lor this spec~es by Flint are absent in Boxelder Creek at Wakins. Flint (I97lbj rrportrd ih~s sprc~eslrorn Oklahoma, Arizona, and Texas..Crnicrldea stgnata (Banks) : L. Colorado, Fort Coll~nr. Prrbious rerords. Garf~eld (Flint, 1974b). 1.arimer (Bank,, 1903). Nru records: 5ioftat. I)rdinagt.5: C.olr~radu, South Platir, Yarnpa. Habitat. strcains. ~ ~retr..iltiiud~nal rang?: to 1801 ni..adult collr~ tion dates 3 to 7 Aup~st HYDROPTII>IDAE Agrnvlea tnult~punclala Curtis : L. England. Prerlous ~ecords: Gilpin (Doddi and Hisaw. 1925b). Larimer (Banks, 1907b). Routt (Canton and Ward, 1981~). Weld (Herrmann, 1977). Nrw records. Boulder, Chaffre, C:uster. Eagle. El Paso, Garfield, Grand, Huerfano. Jeffrrson. Ida Plata, Park, Rlo Blanto. Ilrainages: Anmas, Arkansas, Colorado, South Platte. Whlte. Habitat. lakes, rlvrrs. ~ilt~tudinal range: 1615 to m..adult collrction dates: 18 May to 1 September. *&raylea saltesea Ross T L. Montana, Saltrse. Nen records: Boulder Drainages: South Plattr. Hab~tat:strram. Altitudinal range: 2505 m. Adult <ollect~on dates: 6 August 'Hydropl~la ajax Ross 1938.?: 1.. Illiriots, Oakwood. New records. Alairiosa, Arapahoe, Delta, Garfield, La Plata, Moffat, Park, Pueblo. Drainages: r\rkansas, Colorado, Gunnison, Rio Grande, San Juan, South Platte, Yampa. Hab~tat: strearnr, rivers. ;\ltiiudinal range: 1441 to 1643 m. Adult collect~on dates: 3 July to 7 November. Commentary: Hydropltla alax appears to be restricted altitudinally to below 5577 ft (1700m). Hydropltin arclta Ross T. L. Idaho, Bear Rher Narrows. Previous records. Chaffee (Flint and Herrmann, 1976) New rrcords: Adams, Arrhuleta, Custrr, Douglas, El Paso, Fremont, Carf~eld, Huerfano, L.arime~, Park, Pueblo Dralndgr5: Arkansas, Colorado, 5an Juan. South Plattr. Hab~tat.strearns, rlrers hltitudinal range to 2749 nl. :idult c~~llect~on dates: 2 Jul\ to I Septrmbrr.

11 November I986 Herrmann et al.-distribution and Records of Trichoptera 431 'Hydroptila argosa Ross T. L. Wyoming, Parco. New records: Archuleta, Delta, Eagle, Garfield, Huerfano, Ouray, Park, Pueblo, Rio Blanco, Routt. Drainages: Arkansas, Colorado, Gunnison, San Juan, South Platte, White, Yampa. Altitudinal range: 1515 to 2633 m. Adult collection dates: 15 June to 10 September. Commentary: This species occurs commonly in the warmer lotic waters of the State. Hydroptila callia Denning T. L. North Carolina, Raleigh. Previous records: unspecified locality (Blickle, 1979). New records: none. Drainages: data not available. Habitat: data not available. Altitudinal range: data not available. Adult collection dates: data not available. Commentary: The only Colorado record of this species is that of Blickle (1979), which does not include collection locality and date information. Neither Denning (1984) nor Blickle (1984) could locate the specimens or collection record for this species from Colorado. Blickle (1979) also reported this species from Wyoming. *Hydroptila conszmilis Morton T. L. New York, Ithaca. New records: Garfield, Larimer,.Park. Drainages: Colorado, South Platte. Habitat: rivers. Altitudinal range: 1615 to 2579 m. Adult collection dates: 13 February to 31 August. Hydroptzla hamata Morton T. L. New York, Ithaca. Previous records: unspecified locality (Blickle, 1979). New records: none. Drainages: data not available. Habitat: data not available. Altitudinal range: data not available. Adult collection dates: data not available. Commentary: The only Colorado citation of this species is that of Blickle (1979), which does not include collection locality and date information. According to Denning (1984) the original vial label for this Colorado record was apparently lost; only a handwritten label of "Colo" exists. This species was reported from Utah (Baumann and Unzicker, 1981) and Wyoming (Blickle, 1979)..Hydroptila icona Mosely T. L. Mexico, Chiapas. New records: Garfield, Larimer. Drainages: Colorado, South Platte. Habitat: rivers. Altitudinal range: 1615 m. Adult collection dates: 10 July to 10 September. Commentary: Confusion between this species and H. ajax can be avoided by using Ross (1944), Figures 525B and 526B. Hydroptila pecos Ross T. L. New Mexico, Carlsbad. Previous records: Boulder (Denning, 1947a). New records: Larimer, Pueblo, Weld. Drainages: Arkansas, South Platte. Altitudinal range: 1388 to 1477 m. Adult collect~on dates: 23 April to 29 September. Commentary: This hydroptilid was only collected east of the continental divide on the plains of Colorado. Its range extends from New Mexico to Wyoming (Denning, 1947a). Hydroptila rono Ross T. L. Utah, Huntsville. Previous records: Grand (Ross, 1941), Larimer (Ross, 1941). New records: El Paso, Garfield, Jefferson, Pitkin, Teller. Drainages: Arkansas, Colorado, South Platte. Habitat: streams.

12

13 November I986 Herrmann et a].-distribution and Records of Trichoptera 433 Ochrotrichia logana (Ross) T. L. Utah, Logan County. Previous records: Chaffee (Flint and Herrmann, 1976), Routt (Canton and Ward, 1981~). New records: Archuleta, Boulder, Douglas, Eagle, El Paso, Fremont, Garfield, Gunnison, Hinsdale, Huerfano, Jackson, Jefferson, Lake, La Plata, Montezuma, Montrose, Ouray, Park, Rio Blanco, Teller. Drainages: Animas, Arkansas, Colorado, Dolores, Gunnison, North Platte, San Juan, South Platte, White. Habitat: streams, rivers, springs. Altitudinal range: 1369 to 2579 m. Adult collection dates: 18 June to 18 October. Commentary: This hydroptilid is abundant in Colorado throughout its altitudinal range on both sides of the continental divide. Ochrotr~chia lometa (Ross) T. L. New Mexico, High Rolls. Previous records: unspecified locality (Blickle and Denning, 1977). New records: none. Drainages: data not available. Habitat: data not available. Altitudinal range: data not available. Adult collection dates: data not available. Commentary: The only Colorado record is that of Blickle and Denning (1977), which does not indicate collection locality and date. The specimens constituting this record could not be located (Denning, 1984). This species was reported from Utah (Baumann and Unzicker, 1981). Ochrotr~ch~a oregona (Ross) T. L. Oregon, La Grande. Previous records: Grand (Denning, 1947a). New records: none. Drainages: Colorado. Habitat: rivers. Altitudinal range: 2423 m. Adult collection dates: 27 September. Commentary: Denning (1947a) also reported this species from Wyoming near the Colorado border. Ochrotrich~a stylata (Ross) T. L. Wyoming, Farson. Previous records: Boulder (Denning, 1947a). New records: Arapahoe, Archuleta, Delta, Denver, Garfield, Huerfano, Jefferson, Moffat, Park, Pueblo, Rio Blanco, Routt. Drainages: Arkansas, Colorado, Gunnison, San Juan, South Platte, White, Yampa. Altitudinal range: 1369 to 2579 m. Adult collection dates: 18 June to 18 October. Commentary: This common species is often collected with 0. logana. Ochrotrich~asusanae Flint and Herrmann T. L. Colorado, Trout Creek Spring. Previous records: Chaffee (Flint and Herrmann, 1976). New records: none. Drainages: Arkansas. Habitat: spring. Altitudinal range: 2749 m. Adult collection dates: 18 April to 20 July. Commentary: This species is known only from the type locality. Ochrotrichia trapoiur Ross T. L. Colorado, Yale Lake. Previous records: Chaffee (Ross, 1947). New records: none. Drainages: Arkansas. Habitat: lake. Altitudinal range: 2807 m. Adult collection dates: 5 August. Commentary: Herbert H. Ross probably collected this site only once, according to his son, C. A. Ross (1984). Attempts to collect more specimens of this species at Yale Lake were not successful.

14 434 The Southwestern Naluralzrt Oxyrthzra dualzs Morton T I.. New Mexico, Las Vegas. Prrviouc records: Chaffee (Flint and Herrmann, 1976). Nrxv records. Boulder, Garfield, Pueblo. Drainage?: Arkansas, Colorado, South Plattr..Altitudlnal range: 1445 to 3098 m..adult collection dates: I July to 4 Octobrr. <:~>mmrntar): Ross 11944) reportcd th~species to be widely distributed ovel thr continent, but not often collected. *Startobzrlla hrustza (Ross) T. 1.. Wyoming, Parco. New rrcords: Archuleta, Eagle, Garfirld, Larlmer, Park Drainages: Colorado, San Juan, South Platte. Habitat. ri\er$, xreams..altitudinal range: 1519 to 2579 m. \dult collection dates: 25 July to 10 September. Cornmentar?: Ross (1938b) and Denning (1947a) reported this species from thr North Plattr R~rer111 \\'yoming. Thr same river in C:olorado has not yirlded this species..s'tactobzella dellra (Ross) T L.. Wisconsin, Spoonrr. Previous records: Grand (Swrgman and Ferrinton, 19801, Jarkson idenning, 1947a), Lailmer t Dcnn~ng. 1947a) New iecords: nolir. Drainagt.5 Colorado, North Plattr, South Platte Habit.it 5trrarn5, rivcrs..altitudiridl rang? to 2590 m..adult collrct~on dates: 26 Jul) to 2 August. (.oirirncntarv. Dtxnning (1947a) reported this species from Wyoming 1.LPIDOST OMATIDAE Lrpzdostoma castadensr (Milne) T. L. Bitish Columbia, Cultus Lake. Pleriour records: Chaffre (Ross, 19461, El Paso (Ross, 1946), Larimer (Ross, 1946). New records: Boulder, Custer, Gunn~son, Jackson, Saguache, San Juan, Teller. 1)rainagrs: Animas, Arkansas, Gunnison, North Plattr, South Plattr. flabitat: stream\, rlvrrs..\lt~tudinal range: 2408 to 3124 m. Adult collect~on dates: 4 July to 21 August. C.omrnentar\:.As ~tspecies rlarne indicatrc, this species inhabits rapld, cascading streams and 11\e1s. "L.rp~dostoma grzsea (Banks) r I>,New York, LVoodworth's I.ake. New ~ecords: C;arfield. 1)ralriages: C:oloiado. Habitat: 5prings.\ltitudinal range: 2743 m. \dult collection dates: 25 August. I.ep~do,toma knou'lton~ Ross T. L. Utah, Clinton. Prrv~ousrecords: Montezuma (Ross, 1946), Routt (Canton and M'ard, 1981~). New ietords: C.onejos, Grand, Gunnison, Montrose, Park. Drainages: (hlorado, Conejos, Dolor?$, Gunnison, San Juan, South Platte. Habitat: rivers.,\ltitud~nal range: 1814 to 2707 m. Adult collection dates: 8 Juy to 30 August. C:oinmentar): Colorado sit?$ closely match Anderson's (1976) site description for Oregon-fast, silt) rtieam flowing over rocks, gravel and stones. *Lep~dostomamzra Denning T L. California. Strawberry. New records. Roulder, Clear Creek, Custer, Gilpin, Huerfano. Dralnagrs:.Arkansas, South Plattr. Hahirat: \creams. rlvers. Altit~~d~nal range: 2316 to 3024 rn.\duit collection date\: 3 Jul) to I8 Augu~t. C:oinmentarv. I hi5 sprrirs 1s seldom collected in great numbrrs.

15 November 1986 Herrmann et al.-distribution and Records of Trichoptera Lepidostorna tnoneka Denning T. L. Arizona, Portal. Previous records: Routt (Canton and Ward, 1981a and c). New records: Archuleta. Drainages: San Juan, Yampa. Habitat: rivers. Altitudinal range: 2006 to 2260 m. Adult collection dates: 14 July to 9 September. Commentary: In Colorado this species has been collected only west of the continental divide. The specimens for both county records were identified by D. G. Denning; those from Archuleta County are located at Brigham Young University (R. W. Baumann), those from Routt County could not be located. Lepidostorna ormea Ross T. L. Utah, Heber City. Previous records: Jackson (Denning, 1948a and 1949). New records: Conejos, Eagle, Garfield, Grand, La Plata, Larimer, Pitkin. Drainages: Animas, Colorado, Rio Grande, South Platte. Altitudinal range: 1615 to 2393 m. Adult collection dates: 14 July to 17 August. Commentary: This species appears confined to the foothills zone in Colorado. Lepadostoma pluviale (Milne) T. L. Colorado, Creede. Previous records: Chaffee (Flint and Herrmann, 1976), Grand (Ross, 1946), Larimer (Ross, 1946; Short et a]., 1980), Mineral (Milne, 1936). New records: Boulder, Custer, Douglas, El Paso, Fremont, Jefferson. Drainages: Arkansas, Colorado, Rio Grande, South Platte. Altitudinal range: 2057 to 2749 m. Adult collection dates: 27 June to 10 August. Commentary: This species was reported from the western montane region (Colorado, Montana, Wyoming, and Utah) and Oregon (Anderson, 1976). Lepidostoma roafi (Milne) T. L. Colorado, Sheephorn. Previous records: Boulder (Ross, 1946), Chaffee (Ross, 1946), Eagle (Milne, 19361, El Paso (Ross, 19461, Grand (Milne, 19361, Larimer (Ross, 1946). New records: Garfield, Gunnison, Jackson, Las Animas, Saguache. Drainages: Arkansas, Colorado, Gunnison, North Platte, South Platte. Altitudinal range: 1926 to 3109 m. Adult collection dates: 19 July to 26 August. Commentary: This species has a great altitudinal range. Leptdoslorna strophts Ross T. L. Michigan, Beulah. Previous records: Jackson (Ross, 1946). New records: none. Drainages: North Platte. I-Iabitat: streams, rivers. Altitudinal range: 2716 m. Adult collection dates: data not available. Commentary: Ross (1946) reported this species from Gould but gave no collection date. Denning (1954) considered this species transcontinental and well distributed throughout the West. Lepadostorna unicolor (Banks) T. L. California, San Gabriel Mountains. Previous records: Chaffee (Ross, 1946), El Paso (Ross, 1946), Larimer (Ross, 1946). New records: Boulder, Custer, Garfield, Jefferson, La Plata, Saguache. Drainages: Animas, Arkansas, Colorado, South Platte. Altitudinal range: 1878 to 2536 m. Adult collectiori dates: 21 May to 5 September. Commentary: This species is widespread in the foothills and montane zones of Colorado. Lep~dostorna ueleda Denning T. L. Wyoming, near Centennial. Previous records: Boulder (Ward, 1981), Larimer (Denning, 1948a). New records: Fremont, Hinsdale, Jefferson, Mineral, Park.

16 Dralnages: Arkansas. Gunnison, Rio Grarlde. South Platte. Habitat: streams, river,. Alt~tudinal range: 1700 to 2835 m...2dult collrction dates: 24 June to 19 August. Commeritary: Frorn Ward's (1981) altitudinal data on Middle St. Vrain Creek, Boulder County. the lower limit lor this sprcies was adjusted downward to 1700 m. LEPTOCERIDAE Ceraclea annullcornts (Stephens) T. L. England Previous records: Grand (Ross. 1944). Nrw records: Garfield. Dralnages: Colorado. Habitat: rlvrrs. Altitudinal range: 2420 m. Adult collection dates: 24 July. Commentary: Ross (1944) cited this species as being holarrt~c. *Ceraclea coohus Ross T L.. Wyoming, Pinedale. New records: Grand, Park. Dralnagrs: Colorado, South Plattr. Habitat: rlvrra. Alt~tudinal range: 2420 to 2649 m. Adult rollettiorr dates: 17 June to 10 July. Commentary: Roemhild (1982) recorded this species in Montana as being rarr. 'Cerarlea marulata (Banks) T L. District of Columbia. Washington. New records: Larimrr. Dra~nages: South Plattr. Hab~tat: liver. Altitud~nal range: 1890 m. Adult collection dates: 20 July. Commentary: Only one collection site, the Rig Thompson Canyon near Drake, was found for this specles In Colorado. Mystaczdes alafzmbrtata Hill-Griffin T: L. Oregon, Mt. Jefferson. Prevlous records: C;ilpin (Dodds and Hisaw, L925b). Nru, records: none. 1)raindges: South Platte. Habitat: data not available. Altitudinal range: 2591 to 2896 m. Adult rollecuon dates: data not available. Commrntarl-: 'The altitudinal range was derived from Fig. 3 of Dodds and Hisaw (1925b). Mystaczdes lnterlecta (Banks) T. L. Ontario, Georgian Bay. Previous re(.ords: Grlpin (Dodds and Hisaw, 1925b), Larimer (Yamamoto and Wiggins, 1964). New records. Boulder, Jackson. Drainages: North Platte, South Platte Habitat: lake5 hltitudirlal range: 1646 to 3097 m. Adult collection dates: 3 July to 27 July. Cornrnmtary: Molanna flavicornls is frequently collected with this species. Yamamoto and Ross 11966) statrd all previous North American recoreds cited under the name M. longicornis (Linnaeus) 1758 should be considered interjects. Neclupsyche d~arana (Ross) T. L. Illinois, Havana. Previous records: Boulder (Ward, 1981). New records: Jackson. Drainages. North Platte, South Platte. Altitudinal range: 1544 to 2438 m. Adult collection dates: data not available. Commentary: Roemhild (1982) reported this species as common in Montana and inhabiting streams and rivers.

17 November I986 Herrmann et a1.-distribution and Records of Trichoptera Nectopsyche gracilts (Banks) T. L. Mexico, Cuernavaca. Previous records: Boulder (Ward, 1981). New records: none. Drainages: South Platte. Habitat: stream. Altitudinal range: 1544 m. Adult collection dates: data not available. Commentary: Ward's (1981) record of this species from Middle St. Vrain Creek is outside the expected distribution described by Haddock (1977). *Nectopsyche lahontanensis Haddock T L. Utah, Sevier River. New records: Crowley, Garfield, Jackson, Larimer, Logan, Sedgwick, Yuma. Drainages: Arkansas, Colorado, North Platte, Republican, South Platte. Altitudinal range: 1053 to 2384 m. Adult collection dates: 23 April to 19 August. Commentary: Our data extend eastward the published range (Haddock, 1977) of this species. Nectopsyche stzgmatica (Banks) T. L. New Mexico, Jemez Mountains. Previous records: Boulder (Haddock, 1977; Ward, 1981 ). New records: Jackson, Larimer, Moffat. Drainages: North Platte, South Platte, Yampa. Altitudinal range: 1544 to 2438 m. Adult collection dates: 20 March to 28 August. Commentary: The distribution pattern for N. sttgmatica remains much the same as described by Haddock (1977). +Oecetts arizonica Denning T. L. Arizona, Huachuca. New records: Archuleta, La Plata. Drainages: San Juan. Habitat: rivers. Altitudinal range: 2006 m. Adult collection dates: 28 July. Commentary: This species was previously reported only from central and southern Arizona (Denning, 1951). Oecetis auara (Banks) T. L. Quebec, Sherbrooke. Previous records: Boulder (Ward, 1981 ), Routt (Canton and Ward, 1981 ). New records: Archuleta. Delta, Douglas, Eagle, Garfield. Grand. Jackson, Jefferson. Larimer, Mesa, Moffat, Teller. Drainages: Colorado, Gunnison, North Platte, San Juan, South Platte, Yampa. Habitat: streams, rivers, lakes. Altitudinal r&ge: 1494 to 2454 m. Adult collection dates: 3 July to 21 August. Commentary: The incorrect identification of Oecetrs disjuncta as 0. avara is easily avoided by the use of Smith and Lehmkuhl (1980). 'Oecetts disjuncta (Banks) T. L. California, San Gabriel Mountains. New records: Archuleta, Boulder, Conejos, Custer, Douglas, El Paso, Fremont, Garfield, Huerfano, Jefferson, Larimer, Park, Teller. Drainages: Arkansas, Colorado, Conejos, San Juan, South Platte. Hab~tat: streams, rivers, lakes. Altitudinal range: 1320 to 2720 m. Adult collection dates: 20 June to 1 September. Commentary: Our data indicate that 0. disluncla at altitudes almost 305 m higher and 183 m lower than 0. avara. Oecetrs lnconspacua (Walker) T. L. Georgia. Previous records: Weld (Herrmann, 1977). New records: Arapahoe, Archuleta, Baca, Boulder, El Paso, Grand, Kiowa, Kit Carson, Larimer, Park, Pueblo. Drainages: Arkansas, Colorado, Gunnison, Republican, San Juan, South Platte. Habitat: streams, rivers, lakes, seeps. Altitudinal range: 1295 to 2896 m.

18 438 The Soulhwrslern Natural~st Adult collection dates: 10 June to 1 September Commentary: Th~specles is common in the ar~d regions of Colorado. *O~cetisochrnc~a(Curt~s) T. I.. England. Nrw ~rcords. Denver, Larirnir Drainages: South Plattr. Hab~tat:data riot available..altitudlnal range: 1615 to 1656 rn Adult collection datrc: 27 July to 20 August. Colnrnrntary: Our recordc of 0. ochmcea are from highly populated Denver and Fort Collin,. No aquatic liabltat was apparent at the collertlon sites. Roemhlld (1982) reported this rpr11r.s to ~nhab~t warm-watrl lakrs In Montana. Triaenodrs fro~tlalis Barrks 'r.l. Colorado, Fort C:ollir~s. Prev~oris records: Larilnr~ (Bank\. 1907a~. Keu reco~ds. C-a~ficld, l\.lon<e~urna. Park. Drainages C:olorado, San Juan, South Platte Hahitat: streams, rlxers. lake,.,altltudirral lange: 1615 to 2805 m \dult collecuon dater: 13 Jul, to 15 August. Triarriodes grtsea Banks Colorado, 1)enver. Prev~ourecords: Boulde~ (Dodda and Hisaw, 1925b3, Denver (Banks, 1899). Krw records: Garfield, Gunniaon, Huertano, Lake, I.ar~mrr, Park. llra~nages:arkansac, C:r)lorado, C;unn~son, South Platte. Habltar: itlrams, ri\.er\. ponds, lakes..altitudinal range: to 2920 In. dult collection dater: 2,3 Junc to I I September. C.ommen~a~):rhr altitud~n,~l d~str~bution of thia specles ~ncludea the footh~lls and montane z011es. *Triae~ir>des tarda Milrle I I.. Ontarlo, 1Y1rontn Nrw records: Boulder, F1 Paso, lilt Carson. Drainagrs: Arkansas, Republican. 5outh Platte Hahitat: streams, rtvrrs, lake\. i\ltitudinal range: 1435 to nr. Adult (ollettton datrs: 1 I Jul! to 1 September (lo~nrnerrtar). Thir sprcie\ has the na~rowrst alt~tudinal range of the three specles of Trznrnod~i ~eportrd for C;olorddo, and ur~like the pre\,ious two, occurs on]) on the eart s~de of thr cr~ntlnrntal d~vide. L.lhINEPHII.IDAE Allornyiu gnathos (Ross) : L. Colorado, Tolland. Prevlous records: Chaffer (Ross. 1950h), Gilpin (Ross, 1950b). New records: Clear Creek. C:uster. Gunniton, Park, Pitkin. 1)ralnages:.Arkansas, Coloradc~. C;unnlson, South Plattr. Hah~tat:rtrearns..-\lt~tud~nal rangr: 2713 to 3616 n ~. Adult collection dates: 19 June to 5 August. Commentar~: LTnt~l recently, this speclrs was placed in the genus lmanza (Schm~d, 1980). 7h1s %?nu\ and \peclrs appears restricted to cold rap~d \treams of the montane and alp~r~e?ones of Colorado Allornyza trtpunclala (Banks) 'r. L.. Alaska, Yakutat. Previous records: Boulder (Ward, 1981), Gilpin (Dodds and H~saw, 1925b). New rrcords. Chaffee. Drainagrs:.-\rkansas, South Plattr. Hab~tat:streams. Altitudlnal range: m. Adult collection dates: 7 Julv. Cr~mrnentary: The upper 1i1r11t in the alt~tudlnal range was derived from PIS. 3 of Dodds and H15aw I 1925b). 'Ampi~~~osmo~cus canax (Rob\) 19ii. -1'. L. Utah. L.c>gan Can\oli New rerr~rtih: Chaffer, Frernol~t, C;arf~eld. Jeffersorr. Lake. Ourax Dra~nagrs.Arkansac. Colorado, Glinnlson. 4r)uth Plattr.

19 November I986 Herrmann et al.-distribution and Records of Trichoptera Altitudinal range: 1765 to 2911 m. Adult collection dates: 11 January to 11 October. Commentary: Usually adults appear in September and October in Colorado. Anabolia bzmaculata (Walker) T. L. Canada, Albany River. Previous records: Gilpin (Dodds and Hisaw, 1925b), Grand (Swegman and Ferrington, 1980), Routt (Canton and Ward, 1981~). New records: Archuleta, Boulder, Conejos, Eagle, El Paso, Elbert, Jefferson, Jackson, Larimer, Rio Grande. Drainages: Arkansas, Colorado, Conejos, Gunnison, North Platte, Rio Grande, San Juan, South Platre. Habitat: streams, rivers, lakes. Altitudinal range: 1615 to 3200 m. Adult collection dates: 17 July to 15 October. Commentary: This limnephilid is common in the foothills and montane zones of Colorado. "Anabolia consocia (Walker) T. L. North America. New records: Elbert. Drainages: South Platte. Habitat: stream. Altitudinal range: 2045 m. Adult collection dates: 17 July. Commentary: In Colorado this species 1s known only from K~owa Creek at Elbert. *Apatanta zonella (Zetterstedt) T. L. Lapland. New records: Custer, Huerfano. Drainages: Arkansas. Habitat: lakes. Altitudinal range: 3761 to 3844 m. Adult collection dates: 31 July to 3 August. Commentary: Apatania zonella is rare and only found In the highest of alpine lentic habitats in Colorado. Asynarchus nzgriculus (Banks) T. L. Colorado, Clear Creek. Previous records: Clear Creek (Banks, 1908), Gilpin (Dodds and Hisaw 1925b), Teller (Banks, 1908). New records: Boulder, Custer, Delta, Eagle, Gunnison, Jackson, Lake. Larlmer, Pltkln, Saguache, Summit. Drainages: Arkansas, Colorado, Gunnison, North Platte, South Platte. Habitat: lakes. Altitudinal range: 2621 to 3687 m. Adult collect~?n dates: 3 July to 3 September. Commentary: This species is common from the middle montane into the alpine zone of Colorado. Chyranda cenlral~s (Banks) T. L. Colorado, South Park. Previous records: Boulder (Dodds and Hisaw, 1925b), Clear Creek (Banks, 1900), Gilpin (Dodds and Hisaw, 1925b), Park (Banks, 1900). New records: Garfield, Gunnison, Huertano, Summlt. Drainages: Arkansas, Colorado, Gunnison, South Platte. Habitat: streams. Altitudinal range: 2743 to 3658 m. Adult collection dates: 18 June to 26 August. Commentary: In Colorado this species appears to occur only in the montane and alpine zones. 'Clzstoronia maculata (Banks) T. L. New Mexico, Pecos. New records: Elbert. Drarnages: South Platte. Habitat: stream. Altitudinal range: 2045 m. Adult collection dates: 1 May to 17 July. Commentary: This species is known in Colorado only from Kiowa Creek at Elbert. Ross (1938a) synonymized C, /ormosa (Banks) 1900 with this species. We have followed that conclus~on.

20 440 The Southwestern Naturalist vol. 3 1, no. 4 D~cosmoecusatripes (Hagen) T. I.. Colorado, mountains. Previous records: Lake (Hagen, 1877; Banks, 19431, Park (Banks, 1943), Routt (Canton and Ward, l98lc). New records: Alamosa, Boulder, Clear Creek, El Paso, Fremont. Garfield, Gilpin, Grand, Gunnison, Jefferson, Larimer, Las hnimas, RIO Blanco, Saguache, Teller. Drainages: Arkansas, Colorado. Gunnison, Rio Grande, South Platte. White, Yampa. Habitat: streams, rivers, lakes. Xltitudinal range: 1640 to 3688 In. Adult collection dates: I8 July to 7 September. Commentary: Wiggins and Richardson (1982) aummar17ed the taxonomy, anatomy, and b~ology of this species and others in the genus Dicosmoecus. L)zcosmoecu~ jucundus Banks 1943 was synonymized with this specles by Flint (1966). L)zcosmoecus gzlvipes (Hagen) 'T. L. British Columbia, Quesnel Lakc. Previous records: Gilpin (Dodds and Hisaw, 1925b). New records. none. Drainages. South Platte. Habitat: strearn. Alt~tudinal range: 2743 m. Adult rollection dates: data not available. Commentary: The altitudinal value was derlved from Fig. 3 of Dodds and Hisaw i1925bi. Eccl~somy~a conspersa (Banks) T. L. Washington, Olympia. Prcvious records: Gunnison (Allan, 1975). New records: Clear Creek, Custer, Larimer, Park, San Juan, Teller. Drainages: Animas, Arkansas, South Platte Habitat: strramr Altitudinal range: 2971 to 3667 rn. Adult collectior~ dates: 28 May to 17 July. Commentary: Roemhild 0982) found this species abundant in Montana. It is uncommon in Colorado. Ecclz~omt.ia maculosa Banks T. L. Colorado. Boulder. Prebiour records: Boulder (Ross, 1938a and 1950b; Mecom, 1970, 1972a and b; Ward, 1981), Gllpin (Dodds and Hlsaw, I925b), Summit (Ross, 1938a and 1950b). Kew records: Garfield, Grand, Gunnison, Larimrr. Drainages: Colorado, Gunn~son, South Platte Habltat: streams, rivers. Xlt~tud~nal range: 2558 to n. Adult (ollect~orr date5: 12 July to 6 September C.ornrnrntary: Roemhild (1982) reported E. maculo~a only from the alpine zone of Montana. In Colorado ~t was collected from montane and alpine zones. *(;lyphopsyche zrrorata (Fabricius) 1781.?: L.. Unknown. Kew records: Boulder, Larimer, Pitkin. Drainages: Colorado, South Plattr. Habitat: slreams Altitud~nal rangc: 2999 m Adult collrct~on dates. 9 April to 29 September. Commentary: This species was rarely reported in C;olorado. C;rammotaul~us lorpttae Denning, T L. Colorado, Camerson Pass. Previons records: Gunnison (Flint, 1960), Larimer (Denning, 1941 ). New reto~ds: Elbert, Garfield, Pitkln. Drainages: Colorado, Gunnison, South Platte. White. Habitat: ponds, lakes. Alt~tudinal range: 1935 to 3085 m..kdult c~llertlori dates: I1 August. Co~nmentary: I-his species war reported only [rom Colorado. rhere was probably a t)pograph~cal error in the type description, with Cameron Pass being the type locality. Hesperophylax rons~mil~s (Banks) T L. C;olorado, South Park. Prrbious records: Park (Banks, 1900), Rio Blanco ~Cra) and Ward, 19771, Routt (Canton and \Vard New records. Eagle, Garfield. (,rand. <;unn~si)n. La Plata. P~tk~n

21 November I986 Herrmann et al.-distribution and Records of Trichoptera Drainages: Animas, Colorado, White. Altitudinal range: 1615 to 2743 m. Adult collection dates: 13 February to 10 September. Commentary: We appreciate the assistance of Charles R. Parker for providing information allowing the identification of much of our Hesperophylax material. A systematic revision of this genus is forthcoming (Parker and Wiggins, In Press). Hesperophylax designatus (Walker) T. L. Canada, Great Bear Lake. Previous records: Gilpin (Dodds and Hisaw, 1925a and b), Larimer (Ward and Dufford, 1979), Pueblo (Banks, 1904). New records: Chaffee, El Paso, Fremont, Jefferson, Larimer, Rio Blanco. Drainages: Arkansas, South Platte, White. Habitat: streams, rivers, springs. Altitudinal range: 1768 to 2432 m. Adult collection dates: 24 May to 9 September. Commentary: Ross (1938~) recognized this species as being similar to Hesperophylax occidentalis (Banks) 1908, but not a synonym. Parker and Wiggins (In press) consider Hesperophylax incisus (Banks) 1943 a junior synonym of H. designatus Hesperophylax magnus Banks T. L. Arizona, Palmerlee. Previous records: Park (Banks, 1918). New records: El Paso, Larimer, Pueblo. Drainages: Arkansas, South Platte. Habitat: streams, rivers, springs. Altitudinal range: 1826 to 1963 m. Adult collection dates: 20 August to 11 October. Commentary: This species appears confined to the lower foothills zone of eastern Colorado. Hesperophylax occzdentalis (Banks) T.L. Colorado, Florissant. Previous records: Gilpin (Dodds and Hisaw, 1925b), Rio Blanco (Martinson and Ward, 1982), Routt (Canton and Ward, 1981c), Teller (Banks, 1908). New records: Boulder, Chaffee, Clear Creek, Eagle, El Paso, Fremont, Garfield, Grand, Gunnison, Jefferson, Lake, Larimer, Park, Rio Grande. Drainages: Arkansas, Colorado, Rio Grande, South Platte, White. Habitat: streams, rivers, springs, lakes. Altitudinal range: 1756 to 3667 m. Adult collection dates: 2 April to 19 August. Commentary: Martinson and Ward (1982) described portions of the life history of this species from three springs in northwestern Colorado. Hesperophylax occidenlalis occurs from the lower foothills to alpine; in alpine lakes it shows brachyptery, reduced body size, and darker body pigmentat~on. Homophylax ilauipenn~s Banks T. L. Colorado, South Park. Previous records: Boulder (Denning, 1964a), Park (Banks, 1900). New records: Custer, Garfield, Gunnison. Drainages: Arkansas, Colorado, Gunnison, South Platte. Habitat: streams. Altitudinal range: 2743 to 3616 m. Adult collection dates: 6 July to 24 August. Commentary: Roemhild (1982) reported this species to be rare in cold, alpine streams of Montana. Lenarchus breuipennis (Banks) T. L. Colorado. Previous records: Gilpin (Dodds and Hisaw, ). New records: Larimer. Drainages: South Platte. Habitat: ponds, lakes. Altitudinal range: 3036 m. Adult collection dates: 27 July. Commentary: In Montana Roemhild (1982) found this species only in the alpine and rarely so. 'Lenarchus fauttni (Denn~ng)1949. Wyoming, Libby Flats. New records: Grand, Jackson.

22 442 The Southwest~rt~ Naturalrst Drainages: Colorado, North Plattr Hab~tat:lakrs, ponds. Altitudinal lange: 2873 m. Adult collect~on dates: 9 August. C:ommentary: Baurnann and L:nz~cke~ (1981) ~eported this specirr from L'tah. Lirnnephrlus abbreuratrrs Banks T. I.. Colorado, Tabernash. F're\lous ~ecordi. Gilpin 1Dodds and Hisaw. 1925b), Grand (Banks, 1908). Nru rrcords: Boulder. Chaffer. El Paso, Gunnison. Jackson, Lake. Lar~mer, Mrsa, Park, 5umzn1t. 'I'eller. 1)rainages. Arkansas, Colorado. C,unnison, North Platte, South Platte. Hab~lat ponds, lakrs, strrams. retpr..llt~tud~nal rang? in.ldult collrct~orl datei: I I July to 10 September. Commentatv: Ron and Merklr) I 1952) reported thls limneph~lid as occurring only in Colorado and!v\,oni~ng *Ldrrnnephrlus castor Ross and Mrrklry T. I.. Ctah, 5h miles rast of Beaber h'rw rrtords: Curter. Garfield, Gunnison, L.a Plata. Ilrainages: Ar~imas, Arkansas. Colorado. Gunn~sorl Habltat: ponds. lakes. Xltitudlnal lange: 1990 to 3048 ni Adult collection dates: 4 to 25 August. Cornmentar): '1-111s sperles has been reported only from Colorado and (.[ah. Llmnephzlus ~olomdenszs (Bank,) T. I.. C;olorado, Fort Collins. Previous records: Gilpin (Dodds and Hisaw. 1925b), Larimer (Banks, 1899). Srw records: Boulder. Gur~niiorl, Jackson, Park. Rio Blanco, Summ~t. Teller 1)rainagta: Arkansas, C.olorado. Gunnlson, North Plaue, South Platte Hab~tat:lakes, seeps. Altltudlnal lange: 2743 to 3658 m..adult c(~llert~on dates: 10 July to 10 Septrmber C.~,lnrnrntary: Banks rrported the type localitv as Fort C:ollins, Colorado. The altitudr of Fort <:ollins mi is Inconsistent w~th that of our lowest collectiori site (2743 n~). Therefore, \vr doubt thr valldit) ot Fort Coll~rls belrlg the type local~ty for this sperirs. Lzrnnrphzlus drspar McI.achlan '. L. F~nland. Prr\iour rrcordr: urlspecil~rd localit! inlmmo, New rtcords: Boulder. Ttllrr. Drainages:.Arkansas. South Plattr Hab~tar:lakes, pondr.\ltit~ldlnal range: 3094 to 3319 m. Adult tollrct~~m datt.i: 7 July to 25 August. C.~~nirnrntdr\ Rccently Mal~rk) (1979) s)nonym~zed Lzrnnephzlus rnrnusculus (Banks) 1907 under L. dlspar. M'e haw specimens from only two Colorado ~ites-rrd Rock Lakc and P~kes Peak 5r~err Lake, Lrmnrphrltis d~uerrus ibdnki) : 1. Ar17ona. Prescott. Prrr ic,ui records: unspecif~ed localit) (Banks, 1903 and 1907h). New rrcords: Arapahoe, El Paso, Pueblo, Weld. Drainagrs: Arkansas, South Platle. Habitat. strearns, lakrs, springs. Altitudinal lange: 1434 to 1737 ni. Adult collrct~on dates: 2 May to 21 Srptember. Cornmentar): This specir\ is most often collected In May and June, and is apparently restricted to the plains and lower foothills of eastern Colorado. Lrmnephilus externus (Hagen) T. I-. Canada, North Red River. Previous rc~ords. Gilp~n (Dodds arid Hisaw, 1925b), Gland (Banks, 1907a), Park (Banks, I907a: Ktndall et al., 1981). Routt (Canton and Ward, 1981~) New record. Boulder, Delta, Eagle. Garfield, Gunnison. Jackson, Jefferson, Larimer. Mesa. Teller Drarnages..Arkan\,as, C:olo~ado. C,unnlson, North Pl,ttte. Soll~h PIattr Habltar. ponds, lakrs, rlvrls. itrearrl\,altitudrrral rang?: 2096 to m.

23 November 1986 Herrmann et al.-distribution and Records of Trichoptera 443 Adult collection dates: 13 February to 9 October. Commentary: This species is commonly collected from the lower foothills to the alpine zone. Limnephilus frijole Ross T. L. Texas, Frijole. Previous records: Larimer (Ward and Dufford, 1979). New records: Chaffee, Pueblo, Weld. Drainages: Arkansas, South Platte. Hab~tat:springs, streams, rivers. Altitudinal range: 1477 to 1512 m. Adult collection dates: 23 April to 17 June. Commentary: This limnephilid seems restricted to the eastern plains of Colorado. Ltmnephllus hageni Banks T. L. Northwest Territories, Fort Resolution. Previous records: Grand (Swegman and Ferrington, 1980). New records: none. Drainages: Colorado. Habitat: ponds. Altitudinal range: 2590 m. Adult collection dates: 2 August. Commentary: Two males from Winding River Campground near the west entrance of Rocky Mountain National Park constitute the only record for Colorado. L~mnephtlus hyalinus Hagen T. L. Canada, North Red River. Previous records: indeterminable locality (Banks, 1943). New records: Gunnison, Larimer, Moffat, Teller. Drainages: Gunnison, South Platte, White. Habitat: lakes, ponds. Altitudinal range: 1828 to 2736 m. Adult collection dates: 12 July to 2 September. Commentary: Banks' Colorado record is from Great Lake; we are unable to locate this lake. Limnephilus janus Ross T. L. Colorado, Tolland. Previous records: Gilpin (Dodds and Hisaw, 1925b; Ross, 1938a) Grand (Swegman and Ferrington, 1980). New records: Boulder, Larimer. Drainages: Colorado, South Platte. Habitat: ponds, lakes. Altitudinal range: 2420 to 3097 m. Adult collection dates: 24 July to 15 August. Lzmnephilus labus Ross T. L. Colorado, Wood Lake. Previous records: Boulder (Ross, 1941), Gilpin (Ross, 1941). New records: Douglas, Elbert, Fremont. Drainages: Arkansas, South Platte. Habitat: streams, rivers, springs. Altitudinal range: 1911 to 2896 m. Adult collection dates: I May to 26 July. Commfntary: Nimmo (1971) stated the range of this species was from Alberta and Saskatchewan south to Colorado. In Alberta 11 was found only in the southern and central plains and foothills. Limnephtlus lithus (Milne) T. L. Colorado, Boulder. Previous records: Boulder (Milne, 1935); indeterminable locality (Milne 1935). New records: Larimer, Park, Pueblo. Drainages: Arkansas, South Platte. Altitudinal range: 1487 to 2707 m. Adult collection dates: 12 April to 3 October. Commentary: We were unable to locate Milne's collection site of Regnier, Colorado. Limnephzlus moestus Banks T. L. Newfoundland, Grand Lake. Previous records: unspecified locality (Ross, 1944). New records: Boulder, El Paso, Gunnison, Larimer, San Juan, Teller. Drainages: Animas, Arkansas, Gunnison, South Platte. Habitat: ponds, lakes. Altitudinal range: 2438 to 3358 m.

24 444 The Southwestern Naturalzst vol. 31, no. 4 Adult collection dates: 4 July to 25 August. Commentary: In Alberta, Nimmo (1971) collected this species primarily in the mountains. Ltmnephilus perpus~llus Walker T. L. Canada, Hudson's Bay. Previous records: unspecified locality (Ross and Merkley, 1952). New records: Boulder, Chaffee, El Paso, Summit, Teller. Drainages: Arkansas, Colorado, South Platte. Habitat: streams, lakes, springs Altitudinal range: 2438 to 3319 In. Adult collection dates: 30 June to 25 August. Commentary: Nmmo (1971) reported this limnephilid primarily from ponds or small creeks with abundant sedge or bulrushes in the plains and low foothills of Alberta. In Colorado it was collected in comparable habitats from the upper foothills to lower alpine. Ltmrieph~lus plcturatus MacLachlan T. L. Sweden. Previous records: Boulder, (Dodds and Hisaw, 1925b), Gilpin (Dodds and Hisaw, 1925b), unspecified locality (Banks New records: Chaffee, Grand, C;tinnison, Jackson, Larimer, Teller. Drainages: Arkansas, Colorado. C.unnison, North Platte, South Platte. Habitat: ponds, lakes. Altitudinal range: 2438 to 3658 m Adult collection dates: 24 July to 6 September. Commentary: Ross and Merkley (1952) synonymized the following: L. exulans McLachlan 1876, L. clausus Banks 1924, and I.. ktncazdi Banks Banks (1924) described L. clausa from Long Lake; however, there are several Long Lakes in Colorado, and we do not know which one is correct. This species I, abundant in montane and alplne ponds with highly colored browrr water Dodds and Hisaw (1925b) cited an altitudinal range of 1828 m to 3353 m tor L. clausn. 'Limneph~lusproductus Banks % L. Utah, Vineyard. Nrw ~rcords: Garfield, Gunnison, Lincoln. Drainages: Arkansas, Colorado, C;unnison. Habitat: streams. Altitudinal lange: 1597 to 2896 nr. Adult collect~on dates: 3 August to 16 August. C;ommerrtary: Anderson (1976) reported thls species from Utah, Californ~a and Oregon; Roemhild (1982) cited it for Montana. Flint (1966) found the lectotype of L. concolor Banks 1899 to be an unassociated frmale without an abdomen. He stated this species might be L. asslmtlis (Banks) 1908, L. productus, or L. dtvrrsus. Caudell's (1902) specimens from Pikes Peak were ident~fled as 1.. corrcolor by Banks. From our altitudinal data it appears as ~f L. concolor (Ranks) is L, produclus Banks. Lzrnneph~lusrohwert Banks T. L. Colorado, Florissant. Previous records: Teller (Banks, 1908). New records: Chaffer, Eagle, Gunn~son, Jackson, Lake, Park. Ilrainages: Arkansas, Colorado, Gunnison, North Platte, South Platte. Habitat: streams, springs, lakes. Altitudinal range: 2490 to 3177 rn. Adult collection dates: 10 Junr to 20 August. Commentary: This species appears to be restricted to the montane zone of Colorado. L~mnpphtlusecludens Banks T. L. Brltish Columbia, Penticton. Previous records. Grand (Swrgman and Ferrington, 1980), unspecified locality (Nimmo, 1971). New records: Jackson, Larirner. Drainages: North Platte, South Platte. Habitat: data not available. Altitudinal range: 2286 to 2713 m. Adult collection dates: 25 July to 28 August. Commentary: N~mmo (1971) found this species In Alberta near lakes and sloughs with luxuriarrt growth of sedges and cattails; he collected it primarily on the plains but (,ccasionally the foothills. Lzmnephtlus sptrtatus Banks 'T.L. IJtah. Vineyard. Prev~ous records: unspecified lorality (RIISS and Merkley, 1952). Ncw rerords. Chaffer, C.urrn~son. Park

25 November I986 Herrmann et a1.-distribution and Records of Trichoptera Drainages: Arkansas, Gunnison, South Platte. Habitat: streams, rivers, springs. Altitudinal range: to 2859 m. Adult collection dates: 8 July to 1 September. Commentary: Our collections were not from lakes or ponds as reported by Nimmo (1971) and Roemhild (1982). Limnephilus sublunatus Provancher T. L. Canada, Quebec. Previous records: unspecified locality (Ross and Merkley, 1952). New records: Eagle, Grand, Gunnison, Jackson, Larimer. Drainages: Colorado, Gunnison, North Platte, South Platte. Habitat: lakes. Altitudinal range: 2550 to 2896 m. Adult collection dates: 12 June to 28 August. Commentary: Nimmo (1971) reported this species inhabiting pools, sloughs, and lakes fringed with horse-tails or sedges. Limnephilus laloga Ross T. L. Oklahoma, Taloga. Previous records: Weld (Herrmann, 1977; Kumar et al. 1976) New records: Garfield, Kit Carson, Prowers, Pueblo, Rio Blanco. Drainages: Arkansas, Colorado, Republican, South Platte, White. Habitat: streams, springs, ponds, lakes. Altitudinal range: 1091 to 2164 m. Adult collection dates: 28 March to 3 October. Commentary: This caddisfly inhabits some of the more arid regions of Colorado. Limnephilus tarsalis (Banks) T. L. Colorado, Ward. Previous records: Boulder (Banks, 1920), Gilpin (Dodds and Hisaw, 1925b). New records: none. Drainages: South Platte. Habitat: data not available. Altitudinal range: 2743 to 3353 m. Adult collection dates: data not available. Commentary: The altitudinal range of this species is derived from Fig. 3 of Dodds and Hisaw (1925b). *Limnephilus thorus Ross T. L. Utah, Blue Creek. New records: Larimer. Drainages: South Platte. Habitat: data not available. Altitudinal range: data not available. Adult collection dates: 23 August. Commentary: This species was reported only from Utah (Ross, 1938~; Ross and Merkley, 1952; Baumann and Unzicker, 1981). The only Colorado record is that of a male and a female from Rocky Mountain National Park. Identification was made by G. A. Schuster, Snow Museum, University of Kansas. Nemotaulius hostilis (Hagen) T. L. Michigan and New England. Previous records: Gilpin (Dodds and Hisaw, 1925b). New records: Boulder, Gunnison, Larimer. Drainages: Gunnison, South Platte. Habitat: lakes. Altitudinal range: 2475 to 3112 m. Adult collection dates: 2 June to 29 September. Commentary: In Colorado this species commonly occurs in semidrainage lakes described by Pennak ( 1969). 'Neophylax splendens Denning T. L. Wyoming, southwest of Sheridan. New records: Garfield. Drainages: Colorado, White. Habitat: streams. Altitudinal range: 1920 m. Adult collection dates: 23 April to 19 September. Oligophlebodes ardis Ross T. L. Colorado, Berthoud Pass. Previous records: Clear Creek (Ross, 1941), Lake (Ross, 1949).

26 146 The Southwestern Naluralzil New records: Grand, Routt, Summit. Drainages: Arkansas, Colorado, Yampa. Habitat: streams. Altitudinal range: 3048 to 3658 In. Adult collection dates: 19 June to 6 August Commentary: Ross (1941) reported this species occurring in a switt montane creek. 01zgophlebodes minula (Banks) T. L. Colorado. Prrxious records: Boulder (Ross, 1949), Clear Creek (Ross, 19491, El Paso (Ross, 1949), Gilpin rdodds and Hisaw, 1925b1, Grand (Ross, 19491, Lake (Ross, 1949), Larimer (Ross, 1949), Tellrr (Ross, 1949) New records. Archuleta. Chaffer, Custrr. Eagle, Frrmont, Garfield, Gunn~son, Hinsdale, Huerfano, Jrfferson, Park, Pitkin, Routt, San Juan, San Miguel, Summit. Dra~nagea:..\nimas, Arkansas, Coloiado. Gunnison, San Juan, South Platte, Yampa. Hab~tat: streams, rlvcra -\ltitudinal range: 2012 to 3094 m..2dult collecuon dates: 8 Junr to 2 September. Commentary: Clear,high gradient streams characterize the usual habitat of this common species in Colorado Oligophlrbod~i ~irrra Ross 1944 T. L. California. Yosemite National Park. Previous records: unspecified locality (Nimmo, 1971) New records: Eagle, Pitkin. Drainages: C:olorado. Habitat: streams..alutudinal range: 2853 m. Adult collection dates. 26 July. C:oinmentary. As~de from Nimmo's rrcord, only three specimens from two Colorado local~t~es ~rre~ecorded. *Olzgophlrbodes slgma Milne : L. Utah, Parowan Canon. New ~ecords: C:uster, Garfield. Ilialnages: Arkansas, Colorado. Habitat: strearn,.altitudinal range: 2536 to 2871 m Adult collection dates: 7 July to 4 August. Commentary: This species was collected ~nfrequently in Colorado from clear, turbulent. montane creeks and streams. Onocosmoecus un~color (Banks) L. Washington, Snokomish River. Prcx~ouh records: Boulder (Banks, 1943), Routt (Canton and Ward, 1981~). Sex ~ecords: Chaffee, Conejos, Costilla, El Paso, Garfield, Gilpin, Grand, Gunnison, Hinsdale. Iluerfano, Jackson, Jefferson, Lar~mer, Las Animas, Pitkin, San Miguel. Drainages:.An~mas, Arkanras, Colorado, Dolores, Gunnison, North Platte, Rlo Grande, South Platte, Yampa. Habltat: streams, rlvers. i21titudinal lange: 1768 to 3124 m. Adult collection dates: 19 July to 6 September. Cornmental): We were not able to distinguisl~ 0. un~color from 0, coloradenszs (Lllmer) 1905; thciefore, we have reported all records as 0. untcolor. Phtlarrlus quaerts (Milne) T. L. British Columbia, Quesnel Lake. Prexious records. Teller (Milne, 1935). New records: Gunnison, Jackson, Park. Dra~nages. Gunnison. Nortli Platte, South Platte. Habitat: ponds. Altitudinal range to 3034 m. Adult collection dates: 8 July to I September. Commentary: Wiggins (1963) found thir species abundant in shallow ponds and sloughs on the prairies of Canada. Nimmo (1971) observed P. quaerzs s~milarly in Alberta and eastern Britlsh Columbia. We collected this species in Colorado only from the montane zone *P~ychoglyphaorrnzae (Ross) I.. ITtah, Southfield. Nriv records. Larimer, Park. Dralnagc5. South Plattc

27 November I986 Herrmann et al.-distribution and Records of Trichoptera Habitat: streams. Altitudinal range: 2878 to 8475 m. Adult collection dates: 9 October. Commentary: This species was reported from Utah, Oregon, and California (Denning, 1970). Psychoglypha subborealis (Banks) T. L. Alaska. Previous records: Park (Kendall et al., 1981). New records: Custer, Garfield, Gunnison, Larimer, Mesa, Rio Blanco. Drainages: Arkansas, Colorado, Gunnison, South Platte. Habitat: streams, ponds, lakes. Altitudinal range: 2896 to 3616 m. Adult collection dates: 1 May to 9 October. Commentary: Since adult collections were made from September to May in Alberta, Nimmo (1971) concluded that adults overwinter. We have not collected adults during the winter months. On 2 October 1980 large numbers of adults were observed taking wing under freezing temperatures at 3616 m in the Sangre de Cristo Mountains. The wind was calm, and the sun shown brilliantly. Solar insolation possibly warms the adults sufficiently to permit flight activity. Psychoronia coslalzs (Banks) T. L. New Mexico, Las Vegas Range. Previous records: Teller (Wiggins, 1975 and 1977). New records: Boulder, Clear Creek, Eagle, El Paso, Gilpin, Lake, Larimer, Park, Summit. Drainages: Arkansas, Colorado,'South Platte. Habitat: seeps, streams. Altitudinal range: 2536 to 3627 m. Adult collection dates: 2 May to 29 September. Commentary: Females of this species are brachypterous. This limnephilid appears confined to cool to cold montane and alpine seeps and slower reaches and back waters of mountain streams. 'Pycnopsyche subfascaata (Say) T. L. Pennsylvania. New records: Larimer. Drainages: South Platte. Habitat: streams. Altitudinal range: 1524 m. Adult collection dates: 13 September. Commentary: We have only one record of one male from Colorado; it was collected at Fort Collins in MOLANNIDAE Molanna flauicornis Banks T. L. Manitoba, Husavick. Previous records: Gilpin (Dodds and Hisaw, 1925b). New records: Boulder, Garfield, Gunnison, Larimer. Drainages: Gunnison, South Platte, White. Habitat: lakes. Altitudinal range: 2743 to m. Adult collection dates: 26 June to 31 July. Commentary: This species is frequently collected with Mystacides interlecta. PHILOPOTAMIDAE *Chirmarra utahensis Ross T. L. Utah, Gandy. New records: Huerfano, Park. Drainages: Arkansas, South Platte. Habitat: streams, rivers, springs. Altitudinal range: 1957 to 3109 m. Adult collection dates: 2 July to 18 August. Commentary: Since Fischer ( ) listed this species from Idaho, California, South Dakota, and Wyoming, it has been reported from Oregon (Anderson, 1976), New Mexico (Waltz and McCafferty, 19831, and Montana (Roemhild, 1982). We have a record from a plains/foothills stream and a montane warm spring. Dolophilodes aequalis (Banks) T. L. Colorado, Tolland. Previous records: Boulder (Ward, 1981), Gilpin (Banks, 1924; Dodds and Hisaw, 1925b). New records: Chaffee, Clear Creek, Custer, Eagle, El Paso, Garfield, Larimer, Las Animas, Pitkin, Rio Blanco, Routt, Saguache.

28 118 The Southwestern ~Vaturalzst Drainages: Arkansas, Colorado, Rio Grandr, South Platte, Whitr, Yampa. Habitat: streams. Alt~tudlnal range: 1875 to 3048 m. Adult collection datrs: 12 March to 3 September. Commentary: Our records Indicate a peak spring rmergence for th~s species similar to what.anderson (1976) reported in Oregon. We have not observed an obvious difference in size and pigmentation for early versus later emerging adults, as did Denning (1949). It'ormaldza gabrzella (Banks) L. California, San Gabriel Mountains. Previous records: Routt (Canton and Ward, 1981~). New rrcords: Douglas, Garfield, Jackson. Ilralnages: Colorado, North Platte, South Platte. Altltudlnal range: 2103 to 2662 m. Adult collection dates: 27 April to 20 August. Commentary: Our records extend Nimmo's (1974) eastern limit to Colorado. PHRYGANEIDAE *Agrypnza deflata (M~lne)1931. T 1..Saskatchewan, Indian Head. New rrcords. Garfield, Lake, Larirner, Summit, Teller. Drainagrs: Arkansas. Colorado, South Platte. Habitat: ponds, lakes. Altitudinal range: 2736 to 3181 m. Adult collection dates: 20 June to 18 August. Commentary: Adult daytime resting areas were almost exclusively sedges at the water's edge. *Agrypnza glaczalts Hagen L. Labrador. New records: Larimer, Park. Drainages: South Platte. Hab~tat: ponds. Altitudlnal range: 2720 to 2743 m..idult collection dates: I July to 16 July. Commentary: The larvae and pupae are able to tolerate total dissolved solids levels up to 6000 mg/i.. Agrypnta slramtnea Hagen T L. Canada, Saskatchewan. Previous records: unspecified locality (Ross, 1944). New records: Larirner. Drainages: South Platte. Habltat: lake. Alutudinal range: 2958 m. Adult collection dates: 26 July and 27 July. Ciommentary: We have only collected this species on two dates from Nymph Lake, Rocky blountaln Natlonal Park. Phryganea cznerea Walker T. L. Canada, Albany R~ver. Previous records: Routt (Canton, 1982; Canton and Ward, 1979 and 1981b). h'ew records: Boulder, Larimer, Moffat, Pueblo. Drainages: Arkansas, South Platte, Yampa. Habitat: ponds, lakes. Altitudinal range: 1707 to 3112 m. Adult collection dates: 7 June to 25 October. Commentary: Roemhild (1982) and Anderson (1976) reported this species to occur in rivers. 'Plzlostom~s semifasciala (Say) T. L. Ohio. New rerords: El Paso. Drainages: Arkansas. Habitat: streams. Altitudinal range: 2015 m. Adult collection dates: 27 August. Commentary: Our only record of this large species is a male from a site 4 mi (6.4 km) southsouthwest of Rock Creek Park near Colorado Springs. POLYCXNTROPODIDAE *Polycentropus clnereus Hagen L. Canada. St. Lawrence River New records: Boulder, Larlmer, Moffat.

29 November 1986 Herrmann et a].-distribution and Records of Trichoptera Drainages: South Platte, Yampa. Altitudinal range: 1707 to 3112 m. Adult collection dates: 7 June to 25 October. *Polycentropur gertschi Denning T. L. Arizona, Oak Creek Canyon. New records: Conejos. Drainages: Rio Grande. Habitat: river. Altitudinal range: 2475 m. Adult collection dates: 30 July to 17 August. Commentary: This species was collected twice from the Conejos River in the lower San Luis Valley in Polycentropus halrdus Milne T. L. New Mexico, Hot Springs. New records: Delta. Drainages: Gunnison. Habitat: river. Altitudinal range: 1515 m Adult collection dates: 18 August. Commentary: Our only Colorado record is of two males collected from the Gunnison River near Delta in Polycentropus interruptus (Banks) T. L. New Hampshire, Hampton. Previous records: Jefferson (Banks, 1914). New records: none. Drainages: South Platte. Habitat: data not available. Altitudinal range: data not available. Adult collection dates: data nor available. Commentary: This species has not been reported for Colorado with the exception of Banks' original listing and Ross (1944). ROSS probably repeated Banks' (1914) report in his 1944 listing of five eastern states and Colorado. *Polycentropus uariegalw Banks T. L. Washington, Pullman. New records: El Paso. Drainages: Arkansas. Habitat: stream. Altitudinal range: 1884 m. Adult collection dates: 1 September. Commentary: Fischer ( ) reported this species from British Columbia, Washington, California, and Wyoming. Since Fischer's report, Anderson (1976) cited it from Oregon and Roemhild (1982) from Montana. We have only one Colorado record, a single male collected from Rock Creek in PSYCHOMYIIDAE Psychomyia flauida Hagen T. L. Canada, St. Lawrence River. Previous records: Boulder (Dodds and Hisaw, 1925b; Ward, 1975), Larimer (Banks, 1899), Mesa (Stanford and Ward, 1981). New records: Archuleta, Chaffee, Custer, Delta, Douglas, Eagle, Fremont, Garfield, Grand, Huerfano, Jackson, Jefferson, Lake, Moffat, Park, Pueblo, Rio Blanco. Drainages: Arkansas, Colorado, Gunnison, North Platte, South Platte, White, Yampa. Altitudinal range: 1444 to 2920 m. Adult collection dates: 6 June to 16 September. Commentary: This species is common in most major rivers of Colorado from the upper plains zone to the lower montane. We did not observe the unexpected sex ratio reported by Nimmo (1966) when using ultraviolet light traps. RHYACOPHILIDAE Rhyacophila alberta Banks T. L. Alberta, Banff. Previous records: Boulder (Ward, 1981), Gilpin (Dodds and Hisaw, 1925b; Ross, 1950a), Gunnison (Allan, 1975), San Juan (Ross, 1950a). New records: Clear Creek, Grand, Lake, Larimer, Mineral, San Miguel.

30 ti0 The Southwestern Naturaltst 1)rainages: i\n~mas, Arkansas, Colorado. Gunnison, Rio Grande, South Platte..%lt~tudinal range: 2536 to 3295 m. Adult collection dates: 4 August to 9 October Rhyacophila artgelzta Banks L. California, Pasadena. Previous records: Boulder (Ward, 1981), Clear C~eek (Denning, 1948b), Dolores (Canton and Chadwick, 1983a), Gunnison (Allen, 1975), Larimer (Denning, 1948b; Short and Ward, 1980a1, Rmtt (Canton and Ward, 1977 and 1978), Teller (Short et a]., 1978). Nrw records: Eagle, Garfield, Gilpin, Grand, Ja~kson, Jefferson, Lake, Pitkin. Drainages: Arkansas, Colorado, Gunnison, North Platte, South Platte, Yampa. Alt~tudinal range: 1975 to 3036 nl. Adult collecuon dates 19 July to 19 October. Commentary: Smith (1968b) found this speciea to occur in cold. clear streams in Idaho; this rpecies 1s found under similar condit~ons in Colorado. Rhya~ophzla brurtrzea Banks T.L. New Mexico, Beulah. Pievlous lecorda: Boulder (Mecorn, 1972a and b; Ward, 1981); Dolores (Canton and Chadw~ck, IY83a), Gilpln (Dodds and Hlsaw. 1925b). Gland (Short and Ward, 1980b), Gunnison (Allan, 197.5), Lar~mrr (I)enning, 1948b; Short et al., 1980; Short and Ward, 1980a), Tellel (Short et a]., l978~, Weld (Herrmann, 1977; Kumar et al., 1976). New records: C:haffee. Clear Creek. Custel, Eagle. El Paso, Garfield, Hinsdale, Huerfano, Jefferson, Lake, Las An~nias, Mlneral, Park. Pitkin, Pueblo, Rio Grande, Routt, San Juan, San M~gurl. Summit, Tellrr. Dra~nages.Arkansas, Colo~ado. Gunnison, Rio C;rande, South Platte, Yampa. Habitat: steams, rlrers. Alt~tudlnal range: 2012 to 3261 In Adult collectiori dateq: 2 April to 6 September. Commentary: Thia rhyarophilid IS abundant in Colorado throughout its altitudinal range. Smith and Manuel (1984) synonymized R. arcopedes Banks 1914 with R. brunnea,.4ll Colorddo records before 1984 would be listed as R. acropedes. Rhyarophtla roloradensts Banks T L. New Mexico, Las Vegas. Prrbious records: Boulder (Banks, 1924; Dodds and Hlsaw, 1925b; Ward, 1975 and 1981), Dolores (Canton and Chadwick, 1983a), Douglas (Ward, 1976), Gilpin (Dodds and Hisaw, 1925b), Gunnison (Allan, 1975), Jackson (Denning 1965), Jefferson (Ward, 1976), Larimer (Banks, 1904; Short and Ward, 1980a). New records: Archuleta, Chaffer, Clear Creek, Costilla, Custer, Delta, Eagle, El Paso, Fremont, C>arf~eld,Grand, Hinsdale. Huerfano, La Plata, Mineral, Park, Pitkin, Pueblo, Rio Blanco, Rio Grande, Saguache, San Miguel, Summit, Teller. Dra~riages: An~maa, Arkansas. Colorado, Dolores, Gunnison, Rio Grande, San Juan, South Platte, Wh~te. Habitat: streams, r~vers. Altitudinal range: 1658 to 3319 m. Adult collect~on dates: 22 March to 20 September. C;ommentar): Smith (1968b) reported the immature stages from "...the margins and in the deeper portion of rapids." In Colorado the majority of the pupae overwinter and emerge in thr spring. Perk and Smith (1978) cited this rhyacophilid from Arizona, New Mex~co, Colorado, Utah, Wyoming, Montana, and Idaho. Rhyacoph~la harmstoni Ross T L. Utah, Strawberry Valley. Previous records: Larimer (Denning, 1948b). New records Clea~ Creek, Eagle, Gilpin, Grand, Gunnison, Summit. Dra~nages: C:olorado, Gunnison, South Platte. Altitudinal range: 2713 lo 3024 m. Adult rollec tion dates: 18 April to 18 August. Comnirnta~y: To date we know of reports of this species only from Utah (Ross 1944). Califorma (Drnnlng, 1956), and Colorado. Rhyaroph~la hyalinata Banks T. I.. southwestern Colorado. Prr\~ous rrcords Boulder (Dodds and Hisaw, 1925b; Ward, 1981), Gunn~son (Allrn, 1975), Lar~rner (I)rnn~n~. 1948b, Short and Ward, 1980a1.

31 November 1986 Herrmann et al.-distribution and Records of Trichoptera New records: Custer, Grand, Ouray, San Miguel, Summit, Teller. Drainages: Animas, Arkansas, Colorado, Gunnison, South Platte. Altitudinal range: 2804 to 3542 m. Adult collection dates: 20 July to 8 September. Rhyacophila nevadens~s Banks T. L. Nevada, Reno. Previous records: Gilpin (Dodds and Hisaw, 1925b). New records: none. Drainages: South Platte. Habitat: streams. Altitudinal range: 2743 to 3353 m. Adult collection dates: data not available. Commentary: We have not collected this species even though we made repeated visits to South Boulder Creek where Dodds and Hisaw (1925b) made their collections. The data for altitudinal range were derived from Dodds and Hisaw (1925b). Rhyacophila oreta Ross T. L. Utah, Logan Canyon. Previous records: Dolores (Canton and Chadwick, 1983a). New records: none. Drainages: Dolores. Altitudinal range: 2688 to 2761 m. Adult collection dates: data not available. Commentary: Canton and Chadwick (1983a) reported this species from larval identification. This rhyacophilid was not unexpected in Colorado since it was reported from Utah (Baumann and Unzicker, 1981) and Wyoming (Smith, 1968b). Rhyacophila pellisa Ross T. L. Colorado, Rocky Mountain National Park. Previous records: Boulder (Ward, 1981 ), Larimer (ROSS, 1938b; Denning, 1948b). New records: Chaffee, Clear Creek, Custer, Eagle, El Paso, Garfield, Gilpin, Grand, Gunnison, Jefferson, Lake, Las Animas, Park, Routt, Summit, Teller. Drainages: Arkansas, Colorado, Gunnison, South Platte, Yampa. Altitudinal range: 2286 to 3194 m. Adult collection dates: 8 May to 24 August. Rhyacophila rotunda Banks T. L. Nevada, Reno. Previous records: unspecified locality (Smith, 1968b). New records: Clear Creek, Garfield, Larimer, Ouray. Drainages: Colorado, Gunnison, South Platte. Habitat: streams. Altitudinal range: 2012 to 2950 m. Adult collection dates: I3 February to 4 November. Commentary: This species appears to be limited in distribution to California, Nevada, Utah, and Colorado (Smith, 1968b). Rhyacophila valuma Milne T. L. British Columbia, Cultus Lake. Previous records: Gunnison (Allan, 1975), Lake (Milne, 1936), Pitkin (Denning, 1948b). New records: Fremont. Drainages: Arkansas, Colorado, Gunnison. Altitudinal range: 2195 m. Adult collection dates: 13 July. Commentary: Anderson (1976) reported this species as having a flight period from late March through September in Oregon. Rwmhild (1982) found this species rare in alpine streams of Montana. 'Rhyacophila vao Milne T. L. British Columbia, Cultus Lake. New records: Summit. Drainages: Colorado. Habitat: stream. Altitudinal range: data not available. Adult collection dates: 8 August. Commentary: Anderson (1976) stated R. vao was more tolerant of siltier and warmer streams than R. brunnea (syn. R. acropedes).

32 452 The Soulhwestern,Vat~~ralrst ~(>l.3 I. 110.,+ Rhyacoph~ln uerrula Milne T. L. British Columbia, Cultus Lake. Prfvious records: Boulder (Ward, 1981), Gunnison (Allan, 1975), Larimer (Short and Ward, 1980a), Teller (Short et al., 1978). New records: Clear Creek, Custer, Eagle, Garfield, Gilpin, Grand, Lake, Pitkin. Drainages: Arkansas, Colorado, Gunnison, South Platte. Habitat. streams, rivers. Altitudinal range: 1707 to 3048 m. Adult collection dates: 13 July to 19 October. Commentary: Smith (1968b) reported this specips from Colorado but gave no collection data. He also polnted out that, in Idaho, R. uen-ula was always found in association with a green alga, Praszola sp., which it ate. *Rhyacoph~lauocala Milne T L. Brit~sh Columbia, Cultus Lake. New records: P~tkln. Drainages. Colorado. Habitat: river\. Altitudinal range: 2853 m. Adult collection dates: 26 July. Commentary: Smith (1968b) found this species to be similar to R. hyalznata; he stated it may be a bariable form of R. hyalznata. This species was reported from Utah (Baumann and L'nzickfr, 1981). 'Rhyacoph~lavofzxa Milne T. L. Alberta, Edmonton. New records: Eagle, Lake. Dra~nages: Arkansas, Colorado. Habitat. streams. Alt~tudinal range: 3121 to 3177 m. Adult collert~on dates: 16 to 17 July. Commentary: Nimmo (1971) recorded the flight season in Alberta to be from 15 July to 31 August GENOIDAE Neothremma alzcla Dodds and Hisaw T. L. Colorado, Tolland. Prevlous records: Boulder (Mecom, 1972a and b; Ward, 1981), Gilpin (Dodds and Hisaw, 1925a; Banks, 19301, Gunnison (Allan, 1975), Routt (Canton and Ward, 1978). New records: Clear Creek, Conejos, Custer, Eagle, El Paso, Garfield, Grand. Drainages: Arkansas, Colorado, Rio Grande. South Platte, White. Habitat: steams, seeps. Altitudinal range: 2536 to 3170 m. Adult collection dates: 24 July to 25 August. Commentary: The two western Nearct~c genera ~Veothremmaand Farula were removed from the famil) Limnephilidae and assigned to the family Uenoidae (Wiggins et al., In press) The total number of species of Trichoptera reported from each Colorado county is shown in Fig. 1. The counties in the eastern Great Plains contain the fewest records. No species have been reported for Phillips County or Cheyenne County. The low number of records from eastern Colorado is a result of both low collection effort in this area and lack of permanent surface water throughout much of the region. Those counties with a range in elevation from the plains to the alpine zone appear to have the greatest trichopteran species diversity. Previous valid Colorado records total 123 species; with the addition of 53 new records from our work the total now stands at 176 species in 15 families. Undoubtedly other new records for Colorado will be published in the future. We are particularly interested in seeing specimens and new records from Colorado counties bordering adjoining states.

33 November 1986 Herrmann et al.-distribution and Records of Trichoptera 453 We thank the following individuals for the loan of specimens and/or the identification of specimens: 0. S. Flint (U.S. National Museum); C. L. Remington (Peabody Museum); G. A. Schuster (Snow Museum, University of Kansas); C. R. Parker (U.S. National Museum); U. N. Lanham and S. K. Wu (University of Colorado Museum); W. D. Fronk, J. V. Ward, L. Cline, and H. Evans (Colorado State University); F. M. Brown (Wright-Ingraham Institute); S. Canton (Chadwick & Associates, Littleton, CO); B. Wahle (Longmont, CO); G. B. Wiggins (Royal Ontario Museum); W. G. Downs (Ginger Quill Ranch, Cowdrey, CO); D. G. Denning (Moraga, CA); R. W. Baumann (Brigham Young University); and the late H. H. Ross. We are also indebted to the following for assisting with field work: R. A. Donato (University of Southern Colorado); P. Zichterman (National Park Service, Rocky Mountain National Park); J. Laxson (Colorado Springs School District); J. Fabian (Pueblo County School District 70); J. Butler and C. Rains (City of Colorado Springs); H. W. Hobbs (Lamar Community College); J. N. Herrmann, J. M. Herrmann, B. Brooks, J. Marvin, B. Harvey, S. Arthur, T. Ruiter, and D. Garrison. Robert E. Gregg (University of Colorado) kindly permitted use of his Colorado base map, which was modified to generate Fig. 1. Financial support for field work by the senior author for 1981, 1982 and 1983 was provided by the Office of the Vice President for Academic Affairs, University of Southern Colorado. LITERATURE CITED ALLANJ. D The distributional ecology and diversity of benthic insects in Cement Creek, Colorado. Ecology 56: ANDERSON, N.H Observations on the biology and laboratory rearing of Pseudoslenophylax edwardsi (Trichoptera: Limnephilidae). Can. J. Zool The distribution and biology of the Oregon Trichoptera. Agric. Exper. Sta. Tech. Bull. 134, Oreg. State Univ., Corvallis. BANKS,N New North American neuropteroid insects. Trans. Amer. Entomol. Soc. 24: Descriptions of new North American neuropteroid insects. Trans. Amer. Entomol. Soc. 25: New genera and species of nearctic neuropteroid insects. Trans. Amer. Entomol. SOC. 26: Some new neuropteroid insects. J. N. Y. Entomol. Soc. 11: Neuropteroid insects from New Mexico. Trans. Amer. Entomol. Soc. 30: Descriptions of new nearctic neuropteroid insects. Trans. Amer. Entomol. Soc. 32:1-20, a. Descriptions of new Trichoptera. Proc. Wash. Entomol. Soc. 8: b. New Trichoptera and Psocidae. J. N. Y. Entomol. Soc. 40: Neuropteroid insects-notes and descriptions. Trans. Amer. Entomol. Soc American Trichoptera-notes and descriptions. Can. Entomol. 46: , , , New neuropteroid insects. Bull. Harv. Univ. Mus. Comp. Zool. 62:s New neuropteroid insects. Bull. Harv. Univ. Mus. Comp. Zool Descriptions of new neuropteroid insects. Bull. Harv. Univ. Mus. Comp. Zool. 65: New neuropteroid insects from the United States. Psyche 37: Notes and descriptions of nearctic Trichoptera. Bull. Harv. Univ. Mus. Comp. Zool BAUMANN, R. W. AND J. D. UNZICKER Preliminary checklist of Utah caddisflies (Trichoptera). Encyclia 58: BLICKLE, R. L Hydroptilidae (Trichoptera) of America north of Mexico. New Hamp. Agr. Exp. Sta. Bull. #509, Univ. New Hamp., Durham Personal communication. BLICKLE, R. L. AND D. G. DENNING New species and a new genus of Hydroptilidae. J. Kans. Entomol. Soc. 50: CANTON, S. P Comparative limnology and biota of mine spoils ponds in Colorado. Southwest. Nat. 27: Personal communication.

34 45.1 The Southwestern Naturaltst rol. 31. no. 4 CANTON, S. P. AND J. CV. CHADWICK. 1983a. Seasonal and longitudinal changes in invertebrate functional groups in the Dolores River, Colorado. Freshwat. Invertebr. Biol AND- 1983b. Aquatic insect co~nmunities of natural and artificial substrates in a montane stream J. Freshwat. Ecol <,AUTON, 5. P AND J. V. WARD Effects of coal mine drainage on macrolncertebrates of Trout Creek, C:olorado. Environmental Research Papers, f9, Colo. State Univ., Fort Colllns. -A ND Env~ronmental effects of western coal aurface mining. Part 11-Thr aquatlc macroinvertebrates of Trout Creek, Colorado. Envlron. Protect. Agency Report , Environmental Research Laborator), Duluth, MN. - 4hu Enrlronmental rffects of westrln coal rurlacr minlng. Part I--rhr limnology and blota of mlne rpo~ls ponds in northwest Colorado. Environ. Proterr...\gencj Report 600i Environmental Research Laboratory, Duluth, MN ---- AXD a. Thr aquatic insects, with emphasis on the Tr~choptera, of a Colorado stream affected b) coal strip-mine dralnage. Southwest. Nat. 24: AND- 198lb. Benthos and zooplankton of coal strip mine ponds ~n the mountains (11 northwestern Colorado, I..S.A. Hydrobiologia 85: AX[) ~. Emergencr of 'Trirhoptr~a from Trout Creek, Colorado, I'SA. 171 Proc. Third Int. Simp, on'tr~choptera,c;. P. Moretti, ed. Dr. \V. Junk, Thr Hagur. CAUDLLI.. A. N Somr Insectr from thr surnmit ot Pike's Peak, found on snow Proc Entomol. Soc. LVash DENYING, D. G The genus (;rammotaul~us In North Amerlca, with a desrriptlon of a new spears i?r~rhoptera, 1.imnephilidae). Can. Entomol. 73: a. New species and rrcords of nrarctic Hydroptilidac ftrichoptera). Bull. Brooklyn Entomol. Soc. 42: b. New sperle5 and rrcordi of North..\meritan Hydroptilidae (T~~rho:)- teraj. Psyche ITi a. New and l~ttle known specles of nearctic Trlchoprela. Psyche 55: b...\ review of the Rhyacophilidar. Can. E~itomol. 80: ~. Drscriptlons ol t~ghtnrw specirs of Trichoptera. Bull. Brooklyn Entomol. o c 13: New and little known species of caddisflies. Amer. ~Midl. Natur. 42: Records and drscriptions of nearctlc caddisflle5. Part 111. J. Kansas Entomol. Soc. 24: New aperirs of I.rpldostoma (Trichoptera: 1.rpldostomaudae). Pac-Pac. Entomoi. 30: Trlchoptera. In.quati( ~rirects of Califomla, R. L. LTsinger, cd. 1Tn1r. Calif. Press, Brrkelr) a. The genus Homophylax (Trichoptela: L~mnephilldari. Ann. Entomol. Sor. Arner. 57: b. Trlchoptera of Baja California. Pan-Par. Entomol. 40: Sew rhyacophilids and l~mnephilids (Trithoptrra: Rhyacophil~dae and I.lmnephil~dae). Can. Entomol , The genus Psychoglypha (Trichoptera: 1.1mnephilidae). <:an. F.ntomol. 102: New and interesting Ttlchoptera from the western L'nltrd States. Pan-Pac Eritomol Personal connurnlcation. DENYING, D. G. AND R. I.. BL.ICKLE A review of the genus Ochrotrirhia (Trlrhoptrra: Hydroptilidaej. Ann. Entomol. Soc. hmer. 65: DODDSG. S. AND F. S. HISAW. 1925a. Ecological studies of aquatic Inserts. 111 Adaptations ol caddlsfly larvae to swift streams. Ecology 6: AND -- I925b. Ecological studies on aquatic Insects. IV. Altitudinal range and zonatlon of mayfllcs, stonellies and caddisflles In the Colorado Rockies. Ecology FISCHER, F. C. J 'I'r~chopterorum catalogus. 15 vol. Nederlandbr F.ntomolc)glschr Vrrernlglng, Amsterdam. FLINT, JR 'Taxonomy and biology of nearctlc Ilrnnephll~d larva? ftrlctioptrral, wlch special referencr to 5pecle5 111 ra\tern ITnlted States. Entomol. Am?] 40:l-I 17

35 November 1986 Herrmann et a].-distribution and Records of Trichoptera Notes on certain nearctic Trichoptera in the Museum of Comparative -. Zoology. Proc. U.S. Nat. Mus. 118: a. The genus Culoptila Mosely in the United States, with two new combinations. Proc. Entomol. Soc. Wash b. The genus Smtcridea from North and Central America. Smithsonian Contrib. Zool. 167:l The genus Brachycentrus in North America, with a proposed phylogeny of the genera of Brachycentridae (Trichoptera). Smithsonian Contrib. Zool. 398:l-58. FLINT, 0. S., JR., AND S.J. HERRMANN The description of, and environmental characterization for, a new species of Ochrotrichta from Colorado, USA. Ann. Entomol. Soc. Amer. 69: FLINT,0. S., JR., J. R. VOSHELL, JR. AND C. R. PARKER The Hydropsyche scalaris group in Virginia, with the description of two new species (Trlchoptera: Hydropsychidae). Proc. Biol. Soc. Wash GORDON, A. E A synopsis and phylogenetic outline of the nearctic members of Cheumatopsyche. Proc. Acad. Natur. Sci. Phil. 126: GRAY, L.J. AND J. V. WARD Potential effects of oil shale extraction and processing activities on macroinvertebrates of Piceance and Black Sulfur Creeks, Colorado. Environmental Research Papers, #lo, Colo. State Univ., Fort Collins. HADDOCK, J. D The biosystematics of the caddisfly genus Nectopsyche in North Amer~ca with emphasis on the aquatic stages. Amer. Midl. Nat. 98: HAGEN,H. A Report of the Pseudo-Neuroptera collected by Lirut. W. L. Carpenter in 1873 in Colorado. U.S. Geol. Survey Terr. Report for 1873, pp Report upon the collections of Neuroptera and Pseudo-Neuroptera made in portions of Colorado, New Mexico and Arizona during the years 1872, 1873, and Report upon geographic and geological explorations and surveys west of the one hundredth mer~dian Wheeler Survey, U.S. Army 5: HARRIS, T. L Personal communication. HARRIS, T. L. AND T. M. LAWRENCE Environmental requirements and pollution tolerance of Trichoptera. Environ. Protect. Agency Report 600/ , Environmental Monitoring and Support Laboratory, Cincinnati, OH. HERRMANN, S. J Analysis and structure of four lentic communities on the Colorado (U.S.A.) shortgrass pralrle. Int. Rev. Gesamten Hydrobiol Hydroptilidae (Trichoptera) of Colorado. J. Colo.-Wyo. Acad. Sci. 14:15. (Abstr.) KENDALL, D. M., P. G. KEVAN AND J. D. LAFONTAINE Nocturnal flight activity of moths (Lepidoptera) in alpine tundra. Can. Entomol KUMAR,R., R. J. LAVICNE, J. E. LLOYD AND R. E. PFADT Insects of the Central Plains Experiment Range, Pawnee National Grassland. Science Monogr. #32, Agr. Exp. Sta., Univ. pf Wyo., Laramie. MALICKY, H Notes on some caddisflies (Trichoptera) from Europe and Iran. Aquatic Insects 1:3-16. MARTINSON, R. J. AND J. V. WARD Life history and ecology of Hesperophylax occidenlalis (Banks) (Trichoptera: Limnephilidae) from three springs in the Piceance Basln, Colorado. Freshwat. Invertebr. Biol. 1: MECOM,, J Unusual case building behavior of Hydropsyche occtdentalzs larvae (Trichoptera: Hydropsychidae). Entomol. News 81: a. Feeding habits of Trichoptera in a mountain stream. Oikos 23: b. Productivity and distribution of Trichoptera larvae in a Colorado mountain stream. Hydrobiologia 40: MILNE,L. J Studies in North American Trichoptera. Part Cambridge, Mass Studies in North American Trichoptrra. Part 3. 3: Cambridge, Mass. MILNE, L. J. AND M. J. MILNE The Arctopsychidae of continental America north of Mexico (Trichoptera). Bull. Brooklyn Entomol. Soc. 33: NIMMO, A. P The arrival pattern of Trichoptera at artificial light near Montreal, Quebec. Quaest. Entomol The adult Rhyacophilidae and Limnephilidae of Alberta and eastern British Columbia, and their post-glacial origin. Quaest. Entomol The adult Trichoptera (Insecta) of Alberta and eastern British Columbia, and their post-glacial origins. 11. The families Glossosomatidae and Philopotamldae. Quaest. Entomol. 30:

36 456 The Southwestern Naturalzst vol. 31, no Franc~s Walker types of, and new synonymies for, North Amerrcan Hydropsyche species (Trichoptera, Hydropsychidae). Psyche PARKER, C. R Personal communication. PARKER, C. R. AND G. B. WIGGINS. In Press. The Nearctic caddisfly genus Hesperophylax Banks (Trichoptera: Limnephilidae). Can. J. Zool. PECK,D. L. AND S. D. SMITH. 1978, A revision of the Rhyacophzla coloradensrs complex (Trichoptera: Rhyacophilidae). Melanderla 27: PENNAK, R. U' Regional lake typology in northern Colorado, USA. Verh. Int. Vrr. Lrmnol. l3: Colorado semldrainage mountain lakes. Limnol. Oceanog. 14: ROEMHILD,G The Trichoptera of Montana with drstrrbutronal and ecologrcal notes. Northwest Sci. 56:8-13. Ross, C. A Personal rommunication. Ross, H. H. 1938a. Lectotypes of North American caddisflies in the Museum of Comparatrve Zoology. Psyche 45:l b. Descriptions of new North American Trlchoptera. Proc. Entomol. Soc. Wash. 40:lli ~. Descriptions of nearctlc caddrsflies. Bull. Ill. Nat. Hist. Surv. 21: Descrrptions and records of North American Trichoptera. Trans. Amer. Entomol. Soc. 67: The caddisflies, or Trichoptera, of Illinois. Bull. Ill. Nat. Hist. Surv. 23:l A review of thf nearctic Lepidostomatidae (Trichoptera). Ann. Entomol. Soc...her. 39: Descriptions and records of North Amerlcan Trichoptera with synoptic notes Trans. Amer. Entomol. Soc. 73: Notes and descriptions of nearctic Hydroptilidae. J. Wash. Acad. Sri. 38: Descriptions of some western L.imnephilidae. Pan-Pac. Entomol. 25: a. New species of nearctic Rhyacophrla. J. Wash. Acad. Sci. 40: b. Synoptrc notes on some nearctic limnephilid caddisflies (Trichoptera: 1.rmnephilidae). Amer. Midl. Nat. 43: The caddisfly genus Anagapetus (Trichoptera: Rhyacophilrdae). Pan-Pac. Entomol. 27: Ross, H. H. AND D. R. MERKLEY An annotated key to the nearctrc males ot Lzmtzephzlus. Amer. Midl. Nat. 47: SAETHER.,0. A Chironomids and other invertebrates from North Boulder Creek, C,olorado, linrv. Colo. Studies. Series rn Biology, No. 31, Cont. No. 49. Boulder. SCHEFTER, P LV. AND J. D. UHZLCKER A review of the Hydropsyche morosa-bzfzda complex in North America (Trichoptera: Hydropsychidae). In Proc. Fourth Int. Svrnp. 'Trirhoptera. J. C. Morse, rd. Dr. W. Junk, The Hague. SCHMID, F L.e genre Asynarchus McL. (Trichopt., Limnoph.). Mitt. Schweiz. Entomol. Ges Les insects et arachnrdes du Canada. Part 7. Genera des Trichopteres du Canada et des rtats adjacents. Agr. Can. Publ SCHUSTER.G. A AVD 5. LV. HAMILTON Records of the trichopteran famllres Hydroptrlidae, Philopotamidae, and Psvchomyiidae from Kansas. Tech. Publ. St. Biol Surv. Kan. No. 6: SHORT, R.?L., S. P. CANTONAND J. V. WARD Detrital processing and associated macroinvertebrates in a Colorado mountain stream. Ecology 61: SHORT, R. A. AND J. V. WARD. 1980a. Macrornvertebrates of a Colorado high moun~ain stream. Southwest. Nat A ~ D b. Leaf litter processing in a regulated Rocky Mountain stream. Can. J. Fish. Aquat. Sci. 37: SHORT,R. A,, J. V. WARD, H. L. GARY ASD P. 0. CURRIE. 1978, Aquatic biota of Trout Creek, Man~tou Experimental Forest, Colorado Gen. Tech. Report RM-54, Rocky Mountain Forest and Range Experiment Station, V.S. Dept. Agr., Forest Servrce, Fort Collins, CO. SMITH, D.H. AND D. M. LEHMKC'HL Analysis of two problematic North Ameriran caddisfly specles: Oecetts auara (Banks) and Oeretrs dzs~uncta (Bank5i (Trichoptera: Leptoceridar). Quaest. Fnto~nol. 16: SMITH, S. D. 1968a Thr Arctopsychrnae of Idaho. Pan-Pac. Entomol. 44:

37 November 1986 Herrmann et a].-distribution and Records of Trichoptera b. The Rhyarophila of the Salmon River drainage of Idaho with special reference to larvae. Ann. Entomol. Soc. Amer SMITH, S. D. AND K. L. MANUEL Reconsideration of the nearctic species of the Rhyacophtla acropedes subgroup based on adults (Trichoptera: Rhyacophilidae). In Proc. Fourth Int. Symp. Trichoptera, J. C. Morse, ed. Dr. W. Junk, The Hague. STANFORD, J. A. AND J.V. WARD Preliminary interpretations of the distribution of Hydropsychidae in a regulated river. In Proc. Third Int. Symp. on Trichoptera, G. P. Moretti, ed. Dr. W. Junk, The Hague. SWEGMAN, B. C. AND L. C. FERRINGTON, JR New records of western Trichoptera with notes on their biology. Great Basin Nat. 40: UNZICKER, J. D., L. AGGUS AND L. 0. WARREN A preliminary list of the Arkansas Trichoptera. J. Ga. Entomol. Soc. 5: WALLACE, J. B The larval retreat and food of Arctopsyche; with phylogenetic notes on feeding adaptations in Hydropsychidae larvae (Trichoptera). Ann. Entomol. Soc. Amer. 68: WALTZ, R. D. AND W. P. MCCAFFERTY New caddisfly records for New Mexico (Insecta: Trichoptera). Southwest. Nat. 28: WARD,J. V Bottom fauna-substrate relationships in a northern Colorado trout stream: 1945 and Ecology 56: Comparative limnology of differentially regulated sections of a Colorado mountain river. Arch. Hydrobiol Altitudinal distribution and abundance of Trichoptera in a Rocky Mountain stream. In Proc. Third Int. Symp. on Trichoptera, G. P. Moretti, ed. Dr. W Junk, The Hague. WARD, J. V. AND R. G. DUFFORD Longitudinal and seasonal distribution of macroinvertebrates and epilithic algae in a Colorado springbrook-pond system. Arch. Hydrobiol. 86: WIGGINS,G. B Larvae and pupae of two North American limnephilid caddisfly genera (Trichoptera: Limnephilidae). Bull. Brooklyn Entomol. Soc. 63: Contributions to the systematics of the caddisfly family Limnephilidae (Trichoptera). 11. Can. Entomol. 107: Larvae of the North American caddisfly genera (Trichoptera). University of Toronto Press. Toronto Trichoptera. In An introduction to the aquatic insects of North America, R. W. Merrit and K. W. Cummins, eds. Kendall/Hunt, Dubuque. WIGGINS, G. B. AND J. S. RICHARDSON Revision and synopsis of the caddisfly genus Drcosmoecus (Trichoptera: Limnephilidae; Dicosmoecinae). Aquatic Insects 4: WIGGINS,G. B., J. S. WEAVER 111 AND J. D. UUNZICKER. In Press. Revision of the caddisfly family Uenoidae (Trichoptera). Can. Entomol. YAMAMOTO, T. AND G. B. WIGGINS A comparative study of the North American species in the caddisfly genus Mystacides. Can. J. Zool. 42: YAMAMOTO, T. AND H. H. ROSS A phylogenetic outline of the caddisfly genus Mystacrdes (Trichoptera: Leptoceridae). Can. Entomol. 98: Address of authors: SJH, Dept. of Life Sciences, L'niv. of Southern Colorado, Pueblo, CO 81001; DER, 1588 S. Clermont St., Denver, CO 80222; and JDU, llltnors Natural Hzstory Sumey, 607 E. Peabody, Champargn, IL

38 LINKED CITATIONS - Page 1 of 3 - You have printed the following article: Distribution and Records of Colorado Trichoptera Scott J. Herrmann; David E. Ruiter; John D. Unzicker The Southwestern Naturalist, Vol. 31, No. 4. (Nov. 10, 1986), pp Stable URL: This article references the following linked citations. If you are trying to access articles from an off-campus location, you may be required to first logon via your library web site to access JSTOR. Please visit your library's website or contact a librarian to learn about options for remote access to JSTOR. Literature Cited Comparative Limnology and Biota of Mine Spoils Ponds in Colorado Steven P. Canton The Southwestern Naturalist, Vol. 27, No. 1. (Feb. 19, 1982), pp Stable URL: The Aquatic Insects, with Emphasis on Trichoptera, of a Colorado Stream Affected by Coal Strip-Mine Drainage Steven P. Canton; James V. Ward The Southwestern Naturalist, Vol. 25, No. 4. (Feb. 16, 1981), pp Stable URL: New and Little Known Species of Caddis Flies D. G. Denning American Midland Naturalist, Vol. 42, No. 1. (Jul., 1949), pp Stable URL:

39 LINKED CITATIONS - Page 2 of 3 - The Biosystematics of the Caddis fly Genus Nectopsyche in North America with Emphasis on the Aquatic Stages James D. Haddock American Midland Naturalist, Vol. 98, No. 2. (Oct., 1977), pp Stable URL: Life History and Ecology of Hesperophylax occidentalis (Banks) (Trichoptera: Limnephilidae) from Three Springs in the Piceance Basin, Colorado Robert J. Martinson; James V. Ward Freshwater Invertebrate Biology, Vol. 1, No. 3. (Aug., 1982), pp Stable URL: Feeding Habits of Trichoptera in a Mountain Stream John O. Mecom Oikos, Vol. 23, No. 3. (1972), pp Stable URL: Colorado Semidrainage Mountain Lakes Robert W. Pennak Limnology and Oceanography, Vol. 14, No. 5. (Sep., 1969), pp Stable URL: Synoptic Notes on Some Nearctic Limnephilid Caddisflies. (Trichoptera, Limnephilidae) Herbert H. Ross American Midland Naturalist, Vol. 43, No. 2. (Mar., 1950), pp Stable URL: An Annotated Key to the Nearctic Males of Limnephilus (Trichopetra, Limnephilidae) Herbert H. Ross; Don R. Merkley American Midland Naturalist, Vol. 47, No. 2. (Mar., 1952), pp Stable URL:

40 LINKED CITATIONS - Page 3 of 3 - Macroinvertebrates of a Colorado High Mountain Stream Robert A. Short; James V. Ward The Southwestern Naturalist, Vol. 25, No. 1. (Jan. 30, 1980), pp Stable URL: New Caddisfly Records for New Mexico (Insecta: Trichoptera) R. D. Waltz; W. P. McCafferty The Southwestern Naturalist, Vol. 28, No. 4. (Dec. 9, 1983), pp Stable URL: