FAMILY Engraulidae Gill, anchovies

Transcription

1 FAMILY Engraulidae Gill, anchovies [=Engraulinae, Stolephoriformes, Coilianini, Anchoviinae, Setipinninae, Cetengraulidi] Notes: Spelling in prevailing recent practice Engraulinae Gill, 1861a:55 [ref. 1766] (subfamily) Engraulis [stem corrected to Engraulid- by Gill 1872:17 [ref ], confirmed by Goode 1876:70 [ref. 1832], by Steyskal 1980:170 [ref ], by Whitehead, Nelson & Wongratana 1988:306 [ref. 5725] and by Kottelat 2013b:52 [ref ]; Eschmeyer 1998:2455 [ref ], Zhang 2001:119 [ref ] and Nelson 2006:130 [ref ] used Engraulidae] Stolephoriformes Bleeker, :89, 125 [ref. 428] (subfamily) Stolephorus [stem Stolephor- confirmed by Fowler 1931b:111 [ref ]] Coilianini Bleeker, :137 [ref. 428] (phalanx = tribe) Coilia [stem corrected to Coili- by Jordan & Seale 1925:28 [ref. 2899], confirmed by Fowler 1931c:198 [ref ], by Whitehead, Nelson & Wongratana 1988:306 [ref. 5725] and by Nelson 2006:131 [ref ]] Anchoviinae Jordan & Seale, 1925:28 [ref. 2499] (subfamily) Anchovia [on p. 30 as Anchoviina] Setipinninae Fowler, 1931c:198 [ref ] (subfamily) Setipinna Cetengraulidi Fowler, 1958b:6 [ref. 1470] (tribe) Cetengraulis GENUS Amazonsprattus Roberts, pygmy anchovies [=Amazonsprattus Roberts [T. R.], 1984:317] Notes: [ref. 5829]. Masc. Amazonsprattus scintilla Roberts, Type by original designation (also monotypic). Valid as Amazonsprattus Roberts, 1984 in Engraulidae -- (Whitehead et al. 1988:393 [ref. 5725], Kullander & Ferraris in Reis et al. 2003:39 [ref ]). Current status: Valid as Amazonsprattus Roberts, Engraulidae. Species Amazonsprattus scintilla Roberts, Rio Negro pygmy anchovy [=Amazonsprattus scintilla Roberts [T. R.], 1984:318, Figs. 1-3] Notes: [Proceedings of the California Academy of Sciences (Series 4) v. 43 (no. 20); ref. 5829] Rio Jufari between Castanheiro Grande and Santa Fe, Brazil. Current status: Valid as Amazonsprattus scintilla Roberts, Engraulidae.

2 Distribution: Amazon River basin in lower Negro River, Brazil. Habitat: freshwater. GENUS Anchoa Jordan & Evermann, anchovies [=Anchoa (subgenus of Anchoviella) Jordan [D. S.] & Evermann [B. W.], 1927:501, Anchovietta (subgenus of Anchoa) Nelson [G. J.], 1986:895] Notes: [ref. 2453]. Fem. Engraulis compressus Girard, Type by original designation (also monotypic). Valid as Anchoa Jordan & Evermann, (Grande 1985:245 [ref. 6466], Nelson 1986 [ref. 5735], Whitehead et al. 1988:339 [ref. 5725], Castro-Aguirre et al. 1999:126 [ref ], Nizinski & Munroe 2003:766 [ref ], Kullander & Ferraris in Reis et al. 2003:39 [ref ], Angulo et al. 2013:990 [ref ]). Current status: Valid as Anchoa Jordan & Evermann, Engraulidae. (Anchovietta) [ref. 5735]. Fem. Stolephorus naso Gilbert & Pierson, Type by original designation. Synonym of Anchoa Jordan & Evermann, (Kullander & Ferraris in Reis et al. 2003:39 [ref ]). Current status: Synonym of Anchoa Jordan & Evermann, Engraulidae. Species Anchoa analis (Miller, 1945) - longfin Pacific anchovy [=Anchoviella analis Miller [R. R.], 1945:266, Fig. 1] Notes: [Journal of the Washington Academy of Sciences v. 35 (no. 8); ref ] Laguna Mexcaltitán, Nayarit, near Texpan, west coast of Mexico, near 23 N, 'W. Current status: Valid as Anchoa analis (Miller, 1945). Engraulidae. Distribution: Eastern Pacific: western Mexico, mostly in lagoons and estuaries. Species Anchoa argentivittata (Regan, 1904) - silverstripe anchovy, Regan's anchovy [=Engraulis (Stolephorus) argentivittatus Regan [C. T.], 1904:255, Anchovia arenicola Meek [S. E.] & Hildebrand [S. F.], 1923:201, Pl. 13 (fig. 1)] Notes: [Annals and Magazine of Natural History (Series 7) v. 13 (no. 76) (art. 30); ref ] Las Peñas, Jalisco [Puerto Vallarta], Mexico.

3 Current status: Valid as Anchoa argentivittata (Regan, 1904). Engraulidae. Distribution: Eastern Pacific. (arenicola) [Field Museum of Natural History, Publications, Zoölogical Series v. 15 (publ. 215); ref. 2963] Taboga Island, (Pacific) Panama. Current status: Synonym of Anchoa argentivittata (Regan, 1904). Engraulidae. Species Anchoa belizensis (Thomerson & Greenfield, 1975) - Belize anchovy [=Anchoviella belizensis Thomerson [J. E.] & Greenfield [D. W.], 1975:50, Fig. 1] Notes: [Copeia 1975 (no. 1); ref. 7478] Sibun River, Belize. Current status: Valid as Anchoa belizensis (Thomerson & Greenfield, 1975). Engraulidae. Distribution: Pacific slope, Belize and Guatemala. Habitat: freshwater. Species Anchoa cayorum (Fowler, 1906) - Key anchovy [=Anchovia choerostoma cayorum Fowler [H. W.], 1906:85, Fig. 4] Notes: [Proceedings of the Academy of Natural Sciences of Philadelphia v. 58; ref. 1371] Hailer's Rock, Florida Keys, Florida, U.S.A. Current status: Valid as Anchoa cayorum (Fowler, 1906). Engraulidae. Distribution: Western Atlantic. Habitat: marine Species Anchoa chamensis Hildebrand, Chame Point anchovy [=Anchoa chamensis Hildebrand [S. F.], 1943:54, Fig. 19] Notes: [Bulletin of the Bingham Oceanographic Collection Yale University v. 8 (art. 2); ref ] Chame Point, (Pacific) Panama. Current status: Valid as Anchoa chamensis Hildebrand, Engraulidae. Distribution: Eastern Pacific. Species Anchoa choerostoma (Goode, 1874) - Bermuda anchovy

4 [=Engraulis choerostomus Goode [G. B.], 1874:125] Notes: [American Journal of Science and Arts (Series 3) v. 8 (no. 43, art. 12); ref ] Hamilton Harbor, Bermuda. Current status: Valid as Anchoa choerostoma (Goode, 1874). Engraulidae. Distribution: Western Atlantic: Bermuda. Species Anchoa colonensis Hildebrand, narrow-striped anchovy [=Anchoa hepsetus colonensis Hildebrand [S. F.], 1943:60, Fig. 22] Notes: [Bulletin of the Bingham Oceanographic Collection Yale University v. 8 (art. 2); ref ] Vicinity of Colón, (Atlantic) Panama. Current status: Valid as Anchoa colonensis Hildebrand, Engraulidae. Distribution: Western Atlantic. Species Anchoa compressa (Girard, 1858) - deepbody anchovy [=Engraulis compressus Girard [C. F.], 1858:336] Notes: [General report upon zoology of the several Pacific railroad routes, 1857; ref. 4911] San Diego, California, U.S.A. Current status: Valid as Anchoa compressa (Girard, 1858). Engraulidae. Distribution: Eastern Pacific. Species Anchoa cubana (Poey, 1868) - Cuban anchovy [=Engraulis cubanus Poey [F.], 1868:420, Stolephorus astilbe Jordan [D. S.] & Rutter [C. M.], 1897:95] Notes: [Repertorio Fisico-Natural de la Isla de Cuba v. 2; ref. 3505] Cuba. Current status: Valid as Anchoa cubana (Poey, 1868). Engraulidae. Distribution: Western Atlantic. (astilbe) [Proceedings of the Academy of Natural Sciences of Philadelphia v. 49; ref ] Kingston, Jamaica. Current status: Synonym of Anchoa cubana (Poey, 1868). Engraulidae.

5 Species Anchoa curta (Jordan & Gilbert, 1882) - short anchovy [=Stolephorus curtus Jordan [D. S.] & Gilbert [C. H.], 1882:343] Notes: [Proceedings of the United States National Museum v. 4 (no. 237); ref. 2470] Astillero at Mazatlán, Sinaloa, western Mexico. Current status: Valid as Anchoa curta (Jordan & Gilbert, 1882). Engraulidae. Distribution: Eastern Pacific: Baja California to northern Peru. Habitat: freshwater, brackish, marine. Species Anchoa delicatissima (Girard, 1854) - slough anchovy [=Engraulis delicatissimus Girard [C. F.], 1854:154] Notes: [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 5769] San Diego, California, U.S.A. Current status: Valid as Anchoa delicatissima (Girard, 1854). Engraulidae. Distribution: Eastern Pacific. Species Anchoa eigenmannia (Meek & Hildebrand, 1923) - Eigenmann's anchovy [=Anchovia eigenmannia Meek [S. E.] & Hildebrand [S. F.], 1923:205, Pl. 14 (fig. 2)] Notes: [Field Museum of Natural History, Publications, Zoölogical Series v. 15 (publ. 215); ref. 2963] Taboga Island, (Pacific) Panama. Current status: Valid as Anchoa eigenmannia (Meek & Hildebrand, 1923). Engraulidae. Distribution: Eastern Pacific: southwestern Mexico to Ecuador. Species Anchoa exigua (Jordan & Gilbert, 1882) - slender anchovy [=Stolephorus exiguus Jordan [D. S.] & Gilbert [C. H.], 1882:342, Anchoa tropica Hildebrand [S. F.], 1943:80, Fig. 33] Notes: [Proceedings of the United States National Museum v. 4 (no. 237); ref. 2470] Astillero at Mazatlán, Sinaloa, western Mexico. Current status: Valid as Anchoa exigua (Jordan & Gilbert, 1882). Engraulidae. Distribution: Eastern Pacific: Baja California to Ecuador.

6 (tropica) [Bulletin of the Bingham Oceanographic Collection Yale University v. 8 (art. 2); ref ] Cupica Island, Colombia. Current status: Synonym of Anchoa exigua (Jordan & Gilbert, 1882). Engraulidae. Species Anchoa filifera (Fowler, 1915) - longfinger anchovy [=Anchoa filifera Fowler [H. W.], 1915:524, Fig. 2, Anchoa howelli Hildebrand [S. F.], 1943:106, Fig. 46, Anchoviella longipinna Beebe [W.] & Tee-Van [J.], 1928:48, Fig.] Notes: [Proceedings of the Academy of Natural Sciences of Philadelphia v. 67; ref ] Port of Spain, Trinidad Island, West Indies. Current status: Valid as Anchoa filifera (Fowler, 1915). Engraulidae. Distribution: Western Atlantic. (howelli) [Bulletin of the Bingham Oceanographic Collection Yale University v. 8 (art. 2); ref ] Market at Santos, Brazil. Current status: Synonym of Anchoa filifera (Fowler, 1915). Engraulidae. (longipinna) [Zoologica, Scientific Contributions of the New York Zoological Society v. 10 (no. 1); ref ] Bizoton, Haiti, West Indies. Current status: Synonym of Anchoa filifera (Fowler, 1915). Engraulidae. Species Anchoa helleri (Hubbs, 1921) - Heller's anchovy [=Anchoviella helleri Hubbs [C. L.], 1921:47] Notes: [Proceedings of the Biological Society of Washington v. 34; ref ] San Felipe Bay, Gulf of California, Mexico. Current status: Valid as Anchoa helleri (Hubbs, 1921). Engraulidae. Distribution: Eastern Pacific: Gulf of California. Species Anchoa hepsetus (Linnaeus, 1758) - broad-striped anchovy [=Esox hepsetus Linnaeus [C.], 1758:314, Atherina brownii Gmelin [J. F.], 1789:1397, Esox epsetus Bonnaterre [J. P.], 1788:175, Anchoa ginsburgi Hildebrand [S. F.], 1943:55, Fig. 20, Stolephorus perthecatus Goode [G. B.] & Bean [T. H.], 1882:434] Notes: [Systema Naturae, Ed. X v. 1; ref. 2787] America. Current status: Valid as Anchoa hepsetus (Linnaeus, 1758). Engraulidae.

7 Distribution: Western Atlantic. (brownii) [Caroli a Linné... Systema Naturae per regna tria naturae v. 1 (pt 3); ref ] Atlantic and Pacifics. Current status: Synonym of Anchoa hepsetus (Linnaeus, 1758). Engraulidae. (epsetus) [Tableau encyclopédique et méthodique des trois règnes de la nature... Ichthyologie; ref. 4940] American seas. Current status: Synonym of Anchoa hepsetus (Linnaeus, 1758). Engraulidae. (ginsburgi) [Bulletin of the Bingham Oceanographic Collection Yale University v. 8 (art. 2); ref ] Eslaques Bay, Gulf of Venezuela. Current status: Synonym of Anchoa hepsetus (Linnaeus, 1758). Engraulidae. (perthecatus) [Proceedings of the United States National Museum v. 5 (no. 297); ref. 1840] Pensacola, Florida, U.S.A. Current status: Synonym of Anchoa hepsetus (Linnaeus, 1758). Engraulidae. Species Anchoa ischana (Jordan & Gilbert, 1882) - slender anchovy [=Stolephorus ischanus Jordan [D. S.] & Gilbert [C. H.], 1882:340] Notes: [Proceedings of the United States National Museum v. 4 (no. 237); ref. 2470] (Pacific) Mexico. Current status: Valid as Anchoa ischana (Jordan & Gilbert, 1882). Engraulidae. Distribution: Eastern Pacific: Baja and Gulf of California to northern Peru, including Cocos and the Galápagos Islands. Species Anchoa januaria (Steindachner, 1879) - Rio anchovy [=Engraulis januarius Steindachner [F.], 1879:176 [58]] Notes: [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 80 (nos 1-2); ref. 4229] Harbor of Rio de Janeiro, Brazil. Current status: Valid as Anchoa januaria (Steindachner, 1879). Engraulidae. Distribution: Southwestern Atlantic: Brazil and Venezuela. Species Anchoa lamprotaenia Hildebrand, bigeye anchovy [=Anchoa lamprotaenia Hildebrand [S. F.], 1943:62, Fig. 23, Engraulis hiulcus Goode [G. B.] & Bean [T. H.], 1880:343]

8 Notes: [Bulletin of the Bingham Oceanographic Collection Yale University v. 8 (art. 2); ref ] Key West, Florida, U.S.A. Current status: Valid as Anchoa lamprotaenia Hildebrand, Engraulidae. Distribution: Western Atlantic. (hiulcus) [Proceedings of the United States National Museum v. 2 (no. 98); ref ] Clearwater Harbor, Florida, U.S.A. Current status: Synonym of Anchoa lamprotaenia Hildebrand, Engraulidae. Species Anchoa lucida (Jordan & Gilbert, 1882) - bright anchovy [=Stolephorus lucidus Jordan [D. S.] & Gilbert [C. H.], 1882:341] Notes: [Proceedings of the United States National Museum v. 4 (no. 237); ref. 2470] Mazatlán, Sinaloa, western Mexico. Current status: Valid as Anchoa lucida (Jordan & Gilbert, 1882). Engraulidae. Distribution: Eastern Pacific: Baja and Gulf of California to Ecuador. Habitat: brackish, marine Species Anchoa lyolepis (Evermann & Marsh, 1900) - dusky anchovy, shortfinger anchovy [=Stolephorus lyolepis Evermann [B. W.] & Marsh [M. C.], 1900:89, Fig. 13, Anchoa lyolepis continentalis Cervigón [F.], 1969:238, Anchoa nasuta Hildebrand [S. F.] & Carvalho [J. de P.], 1948:288, Fig. 1, Anchovia platyargyrea Fowler [H. W.], 1911:216, Fig. 4] Notes: [Bulletin of the U. S. Fish Commission v. 20 (pt 1) [for 1900]; ref ] Culebra Island, Puerto Rico. Current status: Valid as Anchoa lyolepis (Evermann & Marsh, 1900). Engraulidae. Distribution: Western Atlantic. (continentalis) [Memoria de la Sociedad de Ciencias Naturales La Salle v. 29; ref ] Current status: Synonym of Anchoa lyolepis (Evermann & Marsh, 1900). Engraulidae. (nasuta) [Copeia 1948 (no. 4); ref ] São Sebastien, São Sebastien Island, Brazil. Current status: Synonym of Anchoa lyolepis (Evermann & Marsh, 1900). Engraulidae.

9 (platyargyrea) [Proceedings of the Academy of Natural Sciences of Philadelphia v. 63; ref. 1382] St. Martin Island, West Indies. Current status: Synonym of Anchoa lyolepis (Evermann & Marsh, 1900). Engraulidae. Species Anchoa marinii Hildebrand, Marini's anchovy [=Anchoa marinii Hildebrand [S. F.], 1943:99, Fig. 42, Anchovia platana Marini [T. L.], 1935:446] Notes: [Bulletin of the Bingham Oceanographic Collection Yale University v. 8 (art. 2); ref ] Off Isla Flores, Montevideo, Uruguay. Current status: Valid as Anchoa marinii Hildebrand, Engraulidae. Distribution: Southwestern Atlantic: Brazil and Argentina. (platana) [Physis (Buenos Aires) v. 11; ref ] Not available, mentioned in passing with no distinguishing features; from Argentina. Current status: Synonym of Anchoa marinii Hildebrand, Engraulidae. Species Anchoa mitchilli (Valenciennes, in Cuvier & Valenciennes, 1848) - bay anchovy [=Engraulis mitchilli Valenciennes [A.], in Cuvier & Valenciennes, 1848:50, Anchoa mitchilli diaphana Hildebrand [S. F.], 1943:91, Fig. 38, Engraulis louisiana Lesueur [C. A.], in Cuvier & Valenciennes, 1848:51] Notes: [Histoire naturelle des poissons v. 21; ref. 1013] New York, U.S.A. Current status: Valid as Anchoa mitchilli (Valenciennes, 1848). Engraulidae. Distribution: Western Atlantic: Atlantic and Gulf coasts. (diaphana) [Bulletin of the Bingham Oceanographic Collection Yale University v. 8 (art. 2); ref ] Grand Isle, Louisiana, U.S.A. Current status: Synonym of Anchoa mitchilli (Valenciennes, 1848). Engraulidae. (louisiana) [Histoire naturelle des poissons v. 21; ref. 1013] Current status: Synonym of Anchoa mitchilli (Valenciennes, 1848). Engraulidae. Species Anchoa mundeola (Gilbert & Pierson, 1898) - false Panama anchovy [=Stolephorus mundeolus Gilbert [C. H.] & Pierson [C. J.], in Jordan & Evermann, 1898:2812] Notes: [Bulletin of the United States National Museum No. 47; ref. 2445] Panama Bay, (Pacific) Panama.

10 Current status: Valid as Anchoa mundeola (Gilbert & Pierson, 1898). Engraulidae. Distribution: Eastern Pacific: southern Gulf of California to Colombia. Species Anchoa mundeoloides (Breder, 1928) - Northern Gulf anchovy [=Anchovia mundeoloides Breder [C. M., Jr.], 1928:9, Fig. 5, Anchoa schultzi Hildebrand [S. F.], 1943:41, Fig. 12] Notes: [Bulletin of the Bingham Oceanographic Collection Yale University v. 2 (art. 2); ref. 636] San Felipe Bay, Gulf of California, Mexico. Current status: Valid as Anchoa mundeoloides (Breder, 1928). Engraulidae. Distribution: Eastern Pacific: southern Baja and Gulf of California to southern Mexico and possibly to Costa Rica. (schultzi) [Bulletin of the Bingham Oceanographic Collection Yale University v. 8 (art. 2); ref ] Extreme northern end of the Gulf of California, near the mouth of the Colorado River. Current status: Synonym of Anchoa mundeoloides (Breder, 1928). Engraulidae. Species Anchoa nasus (Kner & Steindachner, 1867) - longnose anchovy [=Engraulis nasus Kner [R.] & Steindachner [F.], 1867:388 [33], Pl. 2 (fig. 17), Stolephorus cultratus Gilbert [C. H.], 1892:544, Stolephorus naso Gilbert [C. H.] & Pierson [C. J.], in Jordan & Evermann, 1898:2813, Engraulis peruanus Steindachner [F.], 1879:178 [60]] Notes: [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 54 (pt 3); ref. 2640] Chincha Islands, Peru. Current status: Valid as Anchoa nasus (Kner & Steindachner, 1867). Engraulidae. Distribution: Eastern Pacific: lower outer Baja and Gulf of California to Peru. (cultratus) [Proceedings of the United States National Museum v. 14 (no. 880); ref. 1626] Santa Margarita Island, off Baja California, Mexico. Current status: Synonym of Anchoa nasus (Kner & Steindachner, 1867). Engraulidae. (naso) [Bulletin of the United States National Museum No. 47; ref. 2445] Panama Bay, (Pacific) Panama.

11 Current status: Synonym of Anchoa nasus (Kner & Steindachner, 1867). Engraulidae. (peruanus) [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 80 (nos 1-2); ref. 4229] Callao, Peru. Current status: Synonym of Anchoa nasus (Kner & Steindachner, 1867). Engraulidae. Species Anchoa panamensis (Steindachner, 1876) - Panama anchovy [=Engraulis panamensis Steindachner [F.], 1876:589 [39]] Notes: [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 72 (1. Abth.); ref ] (Pacific) Panama. Current status: Valid as Anchoa panamensis (Steindachner, 1876). Engraulidae. Distribution: Eastern Pacific: El Salvador to northern Peru. Species Anchoa parva (Meek & Hildebrand, 1923) - little anchovy [=Anchovia parva Meek [S. E.] & Hildebrand [S. F.], 1923:202, Pl. 10 (fig. 2)] Notes: [Field Museum of Natural History, Publications, Zoölogical Series v. 15 (publ. 215); ref. 2963] Porto Bello, (Atlantic) Panama. Current status: Valid as Anchoa parva (Meek & Hildebrand, 1923). Engraulidae. Distribution: Western Atlantic. Habitat: freshwater, brackish, marine. Species Anchoa pectoralis Hildebrand, bigfin anchovy [=Anchoa pectoralis Hildebrand [S. F.], 1943:52, Fig. 18, Anchoa ubatubae Hildebrand [S. F.] & Carvalho [J. de P.], 1948:290, Fig. 2] Notes: [Bulletin of the Bingham Oceanographic Collection Yale University v. 8 (art. 2); ref ] Vigia, Pará State, Brazil. Current status: Valid as Anchoa pectoralis Hildebrand, Engraulidae. Distribution: Southwestern Atlantic. (ubatubae) [Copeia 1948 (no. 4); ref ] Ubatuba, northern part of São Paulo State, Brazil.

12 Current status: Synonym of Anchoa pectoralis Hildebrand, Engraulidae. Species Anchoa scofieldi (Jordan & Culver, 1895) - Scofield's anchovy [=Stolephorus scofieldi Jordan [D. S.] & Culver [G. B.], in Jordan, 1895:410] Notes: [Proceedings of the California Academy of Sciences (Series 2) v. 5; ref. 2394] Astillero at Mazatlán, Sinaloa, western Mexico. Current status: Valid as Anchoa scofieldi (Jordan & Culver, 1895). Engraulidae. Distribution: Eastern Pacific: central Mexico to Ecuador. Habitat: marine Species Anchoa spinifer (Valenciennes, in Cuvier & Valenciennes, 1848) - spicule anchovy [=Engraulis spinifer Valenciennes [A.], in Cuvier & Valenciennes, 1848:39, Anchoa argenteus Schultz [L. P.], 1949:45, Figs. 5, 6b, Engraulis thrissoides Müller [J.] & Troschel [F. H.], 1849:639] Notes: [Histoire naturelle des poissons v. 21; ref. 1013] Cayenne, French Guiana. Current status: Valid as Anchoa spinifer (Valenciennes, 1848). Engraulidae. Distribution: South and Central America: French Guiana, Guyana, Panama, Suriname and Venezuela and eastern Pacific from Guatemala to Ecuador and possibly Peru. Habitat: freshwater, brackish, marine. (argenteus) [Proceedings of the United States National Museum v. 99 (no. 3235); ref. 3967] Lake Maracaibo, 1 kilometer off Pueblo Viejo, Venezuela. Current status: Synonym of Anchoa spinifer (Valenciennes, 1848). Engraulidae. Distribution: Western Atlantic; eastern Pacific. Habitat: freshwater, brackish, marine. (thrissoides) [Reisen in Britisch-Guiana; ref. 3072] Cuyuni River, Guyana. Current status: Synonym of Anchoa spinifer (Valenciennes, 1849). Engraulidae. Habitat: freshwater, brackish, marine. Species Anchoa starksi (Gilbert & Pierson, 1898) - Starks's anchovy [=Stolephorus starksi Gilbert [C. H.] & Pierson [C. J.], in Jordan & Evermann, 1898:2813, Engraulis clarki Hildebrand [S. F.], 1943:19, Fig. 5] Notes: [Bulletin of the United States National Museum No. 47; ref. 2445] Panama.

13 Current status: Valid as Anchoa starksi (Gilbert & Pierson, 1898). Engraulidae. Distribution: Eastern Pacific: Guatemala to Ecuador and possibly Peru. (clarki) [Bulletin of the Bingham Oceanographic Collection Yale University v. 8 (art. 2); ref ] Balboa, Canal Zone, (Pacific) Panama. Current status: Synonym of Anchoa starksi (Gilbert & Pierson, 1898). Engraulidae. Species Anchoa tricolor (Spix & Agassiz, 1829) - piquitinga anchovy [=Engraulis tricolor Spix [J. B. von] & Agassiz [L.], 1829:51, Pl. 23 (fig. 1), Engraulis lemniscatus Cuvier [G.], 1829:323, Engraulis piquitinga Spix [J. B. von], in Spix & Agassiz, 1829:51, Pl. 23 (fig. 1), Anchoviella salvatoris Fowler [H. W.] & Bean [B. A.], 1923:6] Notes: [Reise nach Griechenland und der Türkei; ref. 13] Bahia and Pará, Brazil. Current status: Valid as Anchoa tricolor (Spix & Agassiz, 1829). Engraulidae. Distribution: Southwestern Atlantic. (lemniscatus) [Le Règne Animal (Edition 2) v. 2; ref. 995] Brazil. Current status: Synonym of Anchoa tricolor (Spix & Agassiz, 1829). Engraulidae. (piquitinga) [Reise nach Griechenland und der Türkei; ref. 13] Current status: Synonym of Anchoa tricolor (Spix & Agassiz, 1829). Engraulidae. (salvatoris) [Proceedings of the United States National Museum v. 63 (no. 2488); ref. 1474] Rio de Janeiro, Brazil. Current status: Synonym of Anchoa tricolor (Spix & Agassiz, 1829). Engraulidae. Species Anchoa trinitatis (Fowler, 1915) - Trinidad anchovy [=Anchovia trinitatis Fowler [H. W.], 1915:527, Fig. 3] Notes: [Proceedings of the Academy of Natural Sciences of Philadelphia v. 67; ref ] Port of Spain, Trinidad Island, West Indies. Current status: Valid as Anchoa trinitatis (Fowler, 1915). Engraulidae. Distribution: Western Atlantic. Habitat: brackish, marine

14 Species Anchoa walkeri Baldwin & Chang, Walker's anchovy [=Anchoa walkeri Baldwin [W. J.] & Chang [N. H. C.], 1970:140, Figs. 1, 2A] Notes: [Pacific Science v. 24 (no. 1); ref. 7716] San Blas, Nayarit, Mexico. Current status: Valid as Anchoa walkeri Baldwin & Chang, Engraulidae. Distribution: Eastern Pacific: Gulf of California to Ecuador. GENUS Anchovia Jordan & Evermann, in Jordan, anchovies [=Anchovia Jordan [D. S.] & Evermann [B. W.], in Jordan, 1895:411] Notes: [ref. 2394]. Fem. Engraulis macrolepidotus Kner, Type by monotypy. Genus described more fully in Jordan & Evermann 1896:449 [ref. 2443]. Valid as Anchovia Jordan & Evermann, (Grande 1985:245 [ref. 6466], Whitehead et al. 1988:377 [ref. 5725], Castro-Aguirre et al. 1999:123 [ref ], Nizinski & Munroe 2003:768 [ref ], Kullander & Ferraris in Reis et al. 2003:39 [ref ], Angulo et al. 2013:990 [ref ]). Current status: Valid as Anchovia Jordan & Evermann, Engraulidae Species Anchovia clupeoides (Swainson, 1839) - Zabaleta anchovy [=Engraulis clupeoides Swainson [W.], 1839:388, Anchovia nigra Schultz [L. P.], 1949:39, Fig. 4, Engraulis productus Poey [F.], 1866:380] Notes: [The natural history and classification v. 2; ref. 4303] Pernambuco coast, Brazil. Current status: Valid as Anchovia clupeoides (Swainson, 1839). Engraulidae. Distribution: Western Atlantic. (nigra) [Proceedings of the United States National Museum v. 99 (no. 3235); ref. 3967] Lake Maracaibo at Yacht Club, Maracaibo, Venezuela. Current status: Synonym of Anchovia clupeoides (Swainson, 1839). Engraulidae. (productus) [Repertorio Fisico-Natural de la Isla de Cuba v. 1; ref ] Matanzas, Cuba. Current status: Synonym of Anchovia clupeoides (Swainson, 1839). Engraulidae.

15 Species Anchovia landivarensis Greenfield & Greenfield, Landivar anchovy (author) [=Anchovia landivarensis Greenfield [D. W.] & Greenfield [T.], 1975:104, Fig. 1] Notes: [Fieldiana Zoology V. 65 (no. 9); ref ] Belize City. Belize, Central America. Current status: Valid as Anchovia landivarensis Greenfield & Greenfield, Engraulidae. Distribution: Belize, Central America. Habitat: marine Species Anchovia macrolepidota (Kner, 1863) - bigscale anchovy [=Engraulis macrolepidotus Kner [R.], 1863:224, "Fig. 7" (unpublished), Stolephorus branchiomelas Eigenmann [C. H.], 1918:682, Anchovia magdalenae Hildebrand [S. F.], 1943:23, Fig. 7, Stolephorus rastralis Gilbert [C. H.] & Pierson [C. J.], in Jordan & Evermann, 1898:2811] Notes: [Sitzungsberichte der Königlich Bayerischen Akademie der Wissenschaften zu München 1863, v. 2; ref. 5002] Río Bayano, (Pacific) Panama. Current status: Valid as Anchovia macrolepidota (Kner, 1863). Engraulidae. Distribution: Eastern Pacific: Gulf of California and outer Baja to northern Peru. (branchiomelas) [Proceedings of the American Philosophical Society v. 56 (no. 7); ref ] Mouth of River Dagua, Pacific coast of Colombia. Current status: Synonym of Anchovia macrolepidota (Kner, 1863). Engraulidae. (magdalenae) [Bulletin of the Bingham Oceanographic Collection Yale University v. 8 (art. 2); ref ] Magdalena Bay, Baja California, Mexico. Current status: Synonym of Anchovia macrolepidota (Kner, 1863). Engraulidae. (rastralis) [Bulletin of the United States National Museum No. 47; ref. 2445] Panama Bay, (Pacific) Panama. Current status: Synonym of Anchovia macrolepidota (Kner, 1863). Engraulidae. Habitat: brackish, marine Species Anchovia surinamensis (Bleeker, 1865) - Surinam anchovy [=Stolephorus surinamensis Bleeker [P.], 1865:178, Anchovia pallida Starks [E. C.], 1913:9, Pl. 1, Anchoviella potiana Schultz [L. P.] & Simões de Menezes [R.], 1951:235, Fig. 1,

16 Anchoviella venezuelae Fowler [H. W.], 1931:406, Fig. 6] Notes: [Nederlandsch Tijdschrift voor de Dierkunde v. 3; ref ] Suriname. Current status: Valid as Anchovia surinamensis (Bleeker, 1865). Engraulidae. Distribution: Estuaries and lower parts of rivers; Trinidad to Amazon River, northeastern South America: Brazil, French Guiana, Guyan, Suriname, Trinidad and Tobado and Venezuela. Habitat: freshwater, brackish. (pallida) [Leland Stanford Jr. University Publications, University Series; ref. 4197] Market at Pará, Brazil. Current status: Synonym of Anchovia surinamensis (Bleeker, 1865). Engraulidae. Habitat: freshwater, brackish. (potiana) [Journal of the Washington Academy of Sciences v. 41 (no. 7); ref ] Poti and Parnaiba rivers, Teresina, Piaui State, Brazil. Current status: Synonym of Anchovia surinamensis (Bleeker, 1865). Engraulidae. Habitat: freshwater, brackish. (venezuelae) [Proceedings of the Academy of Natural Sciences of Philadelphia v. 83; ref. 1410] Caño Guanoco, Venezuela. Current status: Synonym of Anchovia surinamensis (Bleeker, 1865). Engraulidae. Habitat: freshwater, brackish. GENUS Anchoviella Fowler, anchovies [=Anchoviella (subgenus of Anchovia) Fowler [H. W.], 1911:211, Amplova Jordan [D. S.] & Seale [A.], 1925:31] Notes: [ref. 1382]. Fem. Engraulis perfasciatus Poey, Type by original designation. Valid as Anchoviella Fowler, (Grande 1985:245 [ref. 6466], Cervigón 1987 [ref ], Whitehead et al [ref. 5725], Castro-Aguirre et al. 1999:125 [ref ], Nizinski & Munroe 2003:766 [ref ], Kullander & Ferraris in Reis et al. 2003:39 [ref ], Loeb 2012:13 [ref ], Angulo et al. 2013:990 [ref ]). Current status: Valid as Anchoviella Fowler, Engraulidae. (Amplova) [ref. 2499]. Fem. Anchovia brevirostra Meek & Hildebrand, Type by original designation (also monotypic). Synonym of Anchoviella Fowler, (Whitehead et al. 1988:323 [ref. 5725], Castro-Aguirre et al. 1999:125 [ref ]). Current status: Synonym of Anchoviella Fowler, Engraulidae. Species Anchoviella alleni (Myers, 1940) - Allen's anchovy [=Amplova alleni Myers [G. S.], 1940:441] Notes: [Proceedings of the California Academy of Sciences (Series 4) v. 23 (no. 29) (art. 78); ref ] Lake Cashiboya, Ucayali and Morona

17 rivers, Peruvian Amazon system. Current status: Valid as Anchoviella alleni (Myers, 1940). Engraulidae. Distribution: Upper Amazon River basin, Peru. Habitat: freshwater. Species Anchoviella balboae (Jordan & Seale, 1926) - Balboa anchovy [=Amplova balboae Jordan [D. S.] & Seale [A.], 1926:411, Anchovia brevirostra Meek [S. E.] & Hildebrand [S. F.], 1923:198, Pl. 12 (fig. 1)] Notes: [Bulletin of the Museum of Comparative Zoology v. 67 (no. 11); ref. 2500] Balboa, Canal Zone, (Pacific) Panama. Current status: Valid as Anchoviella balboae (Jordan & Seale, 1926). Engraulidae. Distribution: Eastern Pacific: Honduras and Nicaragua to Panama. (brevirostra) [Field Museum of Natural History, Publications, Zoölogical Series v. 15 (publ. 215); ref. 2963] Balboa, Canal Zone, (Pacific) Panama. Current status: Synonym of Anchoviella balboae (Jordan & Seale, 1926). Engraulidae. Species Anchoviella blackburni Hildebrand, Blackburn's anchovy [=Anchoviella blackburni Hildebrand [S. F.], 1943:129, Fig. 57] Notes: [Bulletin of the Bingham Oceanographic Collection Yale University v. 8 (art. 2); ref ] Jacque Point, Gulf of Venezuela. Current status: Valid as Anchoviella blackburni Hildebrand, Engraulidae. Distribution: Caribbean Sea. Species Anchoviella brevirostris (Günther, 1868) - snubnose anchovy [=Engraulis brevirostris Günther [A.], 1868:392, Anchoviella brasiliensis Hildebrand [S. F.], 1943:138, Fig. 62, Anchoviella hildebrandi Carvalho [J. de P.], 1950:78, Fig. 1] Notes: [Catalogue of the fishes in the British Museum v. 7; ref. 1990] Caxoeira, Río Paraguaçú, Bahia, Brazil. Current status: Valid as Anchoviella brevirostris (Günther, 1868). Engraulidae. Distribution: Atlantic estuaries and river mouths: Brazil, French Guiana, Guyana, Suriname and Venezuela.

18 (brasiliensis) [Bulletin of the Bingham Oceanographic Collection Yale University v. 8 (art. 2); ref ] Rio Ribeira de Iguapé, São Paulo, Brazil. Current status: Synonym of Anchoviella brevirostris (Günther, 1868). Engraulidae. (hildebrandi) [Boletim do Instituto Paulista Oceanografia v. 1 (no. 2); ref ] Río Paraguaçú at Caxoeira, Bahia. Current status: Synonym of Anchoviella brevirostris (Günther, 1868). Engraulidae. Species Anchoviella carrikeri Fowler, Carriker's anchovy [=Anchoviella carrikeri Fowler [H. W.], 1940:73, Fig. 28] Notes: [Proceedings of the Academy of Natural Sciences of Philadelphia v. 92; ref. 1436] Mouth of Río Chapare, Cochabamba, Bolivia. Current status: Valid as Anchoviella carrikeri Fowler, Engraulidae. Distribution: Amazon River basin: Bolivia and Brazil. Habitat: freshwater. Species Anchoviella cayennensis (Puyo, 1946) - Cayenne anchovy [=Stolephorus cayennensis Puyo [J.], 1946:101 [2], Fig. 1, Anchoviella victoriae Hildebrand [S. F.] & Carvalho [J. de P.] 1948:292, Fig. 3] Notes: [Bulletin de la Société d'histoire Naturelle de Toulouse v. 80 (1945); ref ] Cayenne River at Macouria, about 4 kilometers from its mouth, French Guiana. Current status: Valid as Anchoviella cayennensis (Puyo, 1946). Engraulidae. Distribution: Atlantic coastal river mouths: Brazil, French Guiana, Suriname and Trinidad and Tobago. Habitat: brackish. (victoriae) [Copeia 1948 (no. 4); ref ] Victoria, Espirito Santo State, Brazil. Current status: Synonym of Anchoviella cayennensis (Puyo, 1946). Engraulidae. Habitat: brackish. Species Anchoviella elongata (Meek & Hildebrand, 1923) - elongate anchovy [=Anchovia elongata Meek [S. E.] & Hildebrand [S. F.], 1923:198, Pl. 12 (fig. 2)] Notes: [Field Museum of Natural History, Publications, Zoölogical Series v. 15 (publ. 215); ref. 2963] Mindi Cut, Canal Zone, Panama. Current status: Valid as Anchoviella elongata (Meek & Hildebrand, 1923). Engraulidae.

19 Distribution: Western Atlantic. Species Anchoviella guianensis (Eigenmann, 1912) - Guyana anchovy [=Stolephorus guianensis Eigenmann [C. H.], 1912:447, Pl. 62 (fig. 5)] Notes: [Memoirs of the Carnegie Museum v. 5 (no. 1); ref. 1227] Bartica rocks, Guyana. Current status: Valid as Anchoviella guianensis (Eigenmann, 1912). Engraulidae. Distribution: Caribbean to Amazon River basin: Brazil, Colombia, French Guiana, Guyana, Suriname and Venezuela. Habitat: freshwater, brackish. Species Anchoviella jamesi (Jordan & Seale, 1926) - James's anchovy [=Amplova jamesi Jordan [D. S.] & Seale [A.], 1926:410] Notes: [Bulletin of the Museum of Comparative Zoology v. 67 (no. 11); ref. 2500] Rio Jutahy, tributary of rio Solimões, Amazonas State, Brazil. Current status: Valid as Anchoviella jamesi (Jordan & Seale, 1926). Engraulidae. Distribution: Orinoco River basin and Amazon River basin: Brazil, Colombia, Peru and Venezuela. Habitat: freshwater. Species Anchoviella juruasanga Loeb, Loeb's anchovy (author) [=Anchoviella juruasanga Loeb [M. V.] 2012] Notes: [Neotropical Ichthyology v. 10 (no. 1); ref ] Rio Trombetas, upstream from mouth of Lago do Jacaré at Reserva Biológica de Trombetas 1 20'S, 56 51'W, Pará, Brazil. Current status: Valid as Anchoviella juruasanga Loeb, Clupeidae. Distribution: Middle and lower Amazon River basin, Brazil. Habitat: freshwater. Species Anchoviella lepidentostole (Fowler, 1911) - broadband anchovy [=Anchovia lepidentostole Fowler [H. W.], 1911:214, Fig. 3, Anchoviella hubbsi Hildebrand [S. F.], 1943:128, Fig. 56,

20 Anchoviella iheringi Fowler [H. W.], 1941:124, Fig. 2, Anchoviella nitida Hildebrand [S. F.] & Carvalho [J. de P.], 1948:294, Fig. 4] Notes: [Proceedings of the Academy of Natural Sciences of Philadelphia v. 63; ref. 1382] Suriname. Current status: Valid as Anchoviella lepidentostole (Fowler, 1911). Engraulidae. Distribution: Atlantic and South America, estuaries and entering rivers: Brazil, French Guiana, Guyana, Suriname and Venezuela. Habitat: freshwater, brackish, marine. (hubbsi) [Bulletin of the Bingham Oceanographic Collection Yale University v. 8 (art. 2); ref ] Rio Ribeira de Iguapé, São Paulo, Brazil. Current status: Synonym of Anchoviella lepidentostole (Fowler, 1911). Engraulidae. Habitat: freshwater, brackish, marine. (iheringi) [Proceedings of the Academy of Natural Sciences of Philadelphia v. 93; ref. 1437] Rio Jaguaribe, Brazil. Current status: Synonym of Anchoviella lepidentostole (Fowler, 1911). Engraulidae. Habitat: freshwater, brackish, marine. (nitida) [Copeia 1948 (no. 4); ref ] Victoria, Espirito Santo State, Brazil. Current status: Synonym of Anchoviella lepidentostole (Fowler, 1911). Engraulidae. Habitat: freshwater, brackish, marine. Species Anchoviella manamensis Cervigón, Manamo anchovy [=Anchoviella manamensis Cervigón [F.] 1982:220, Figs. 5-6] Notes: [Los recursos pesqueros del Rio Orinoco y su explotacion; ref ] Caño Mánamo, near Tucupita, Orinoco Delta, Venezuela. Current status: Valid as Anchoviella manamensis Cervigón, Engraulidae. Distribution: Orinoco River Delta, Venezuela. Habitat: freshwater. Species Anchoviella miarcha (Jordan & Gilbert, 1882) - Mazatlan anchovy (author) [=Stolephorus miarchus Jordan [D. S.] & Gilbert [C. H.], 1882:344] Notes: [Proceedings of the United States National Museum v. 4 (no. 237); ref. 2470] Mazatlán harbor, Sinaloa, western Mexico. Current status: Valid as Anchoviella miarcha (Jordan & Gilbert, 1882). Engraulidae. Distribution: Eastern Pacific. Species Anchoviella perezi Cervigon, Guayana anchovy (author)

21 [=Anchoviella perezi Cervigón [F.], 1987:27, Fig. 2 et seq.] Notes: [Contribuciones Cientificas, Escula de Ciencas ADD No. 14; ref ] Laguna La Ceiba, Los Castillos de Guayana, Venezuela. Current status: Valid as Anchoviella perezi Cervigón, Engraulidae. Distribution: Oronico River basin delta, Venezuela. Habitat: brackish. Species Anchoviella perfasciata (Poey, 1860) - flat anchovy, Poey's anchovy [=Engraulis perfasciatus Poey [F.], 1860:312] Notes: [Memorias sobre la historia natural de la Isla de Cuba v. 2; ref. 3499] Cuba. Current status: Valid as Anchoviella perfasciata (Poey, 1860). Engraulidae. Distribution: Western Atlantic. Species Anchoviella vaillanti (Steindachner, 1908) - Vaillant's anchovy [=Engraulis vaillanti Steindachner [F.], 1908:193] Notes: [Anzeiger der Akademie der Wissenschaften in Wien v. 45 (no. 13); ref ] Rio São Francisco at Joazeiro and Ciudade da Barra; Rio Preto and Rio Grande do Norte, Brazil. Current status: Valid as Anchoviella vaillanti (Steindachner, 1908). Engraulidae. Distribution: Middle São Francisco River, Brazil. Habitat: freshwater. GENUS Cetengraulis Günther, anchovetas [=Cetengraulis Günther [A.], 1868:383] Notes: [ref. 1990]. Fem. Engraulis edentulus Cuvier, Type by subsequent designation. Type designated by Jordan & Evermann 1896:450 [ref. 2443]. Valid as Cetengraulis Günther, (Grande 1985:245 [ref. 6466], Whitehead 1967:132 [ref. 6464], Whitehead et al. 1988:381 [ref. 5725], Castro-Aguirre et al. 1999:122 [ref ], Nizinski & Munroe 2003:768 [ref ]). Current status: Valid as Cetengraulis Günther, Engraulidae.

22 Species Cetengraulis edentulus (Cuvier, 1829) - Atlantic anchoveta [=Engraulis edentulus Cuvier [G.], 1829:323, Engraulis brevis Poey [F.], 1866:379, Stolephorus garmani Evermann [B. W.] & Marsh [M. C.], 1899:352, Stolephorus gilberti Evermann [B. W.] & Marsh [M. C.], 1899:352, Stolephorus robertsi Jordan [D. S.] & Rutter [C. M.], 1897:95, Hildebrandichthys setiger Schultz [L. P.], 1949:49, Fig. 7] Notes: [Le Règne Animal (Edition 2) v. 2; ref. 995] Rio de Janeiro, Brazil [original locality Jamaica]. Current status: Valid as Cetengraulis edentulus (Cuvier, 1829). Engraulidae. Distribution: Western Atlantic. (brevis) [Repertorio Fisico-Natural de la Isla de Cuba v. 1; ref ] Cuba. Current status: Synonym of Cetengraulis edentulus (Cuvier, 1829). Engraulidae. (garmani) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 25 (for 1899) Appendix 7; ref. 1283] Puerto Real, Puerto Rico. Current status: Synonym of Cetengraulis edentulus (Cuvier, 1829). Engraulidae. (gilberti) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 25 (for 1899) Appendix 7; ref. 1283] Palo Seco, near San Juan, Puerto Rico. Current status: Synonym of Cetengraulis edentulus (Cuvier, 1829). Engraulidae. (robertsi) [Proceedings of the Academy of Natural Sciences of Philadelphia v. 49; ref ] Kingston, Jamaica. Current status: Synonym of Cetengraulis edentulus (Cuvier, 1829). Engraulidae. (setiger) [Proceedings of the United States National Museum v. 99 (no. 3235); ref. 3967] Near mouth of Caño de Sagua, about 25 kilometers north of Sinamaica, Venezuela. Current status: Synonym of Cetengraulis edentulus (Cuvier, 1829). Engraulidae. Species Cetengraulis mysticetus (Günther, 1867) - anchoveta, Pacific anchoveta [=Engraulis mysticetus Günther [A.], 1867:604, Cetengraulis engymen Gilbert [C. H.] & Pierson [C. J.], in Jordan & Evermann, 1898:2815, Stolephorus opercularis Jordan [D. S.] & Gilbert [C. H.], 1882:275]

23 Notes: [Proceedings of the Zoological Society of London 1866 (pt 3); ref. 1984] Panama (Pacific). Current status: Valid as Cetengraulis mysticetus (Günther, 1867). Engraulidae. Distribution: Eastern Pacific. (engymen) [Bulletin of the United States National Museum No. 47; ref. 2445] Panama Bay, (Pacific) Panama. Current status: Synonym of Cetengraulis mysticetus (Günther, 1867). Engraulidae. (opercularis) [Proceedings of the United States National Museum v. 4 (no. 227); ref ] Punta San Felipe, Gulf of California, Mexico (stomach content). Current status: Synonym of Cetengraulis mysticetus (Günther, 1867). Engraulidae. GENUS Coilia Gray, grenadier anchovies [=Coilia (subgenus of Engraulis) Gray [J. E.], 1830:Pl. 85 (v. 1), Chaetomus McClelland [J.], 1844:393, 405, Demicoilia Jordan [D. S.] & Seale [A.], 1925:28, Leptonurus Bleeker [P.], 1849:14, Mystus Lacepède [B. G. E.], 1803:466, Thrissa Cuvier [G.], 1816:176, Thrissocles Jordan [D. S.] & Evermann [B. W.], in Jordan, 1917:98] Notes: [ref. 1878]. Fem. Engraulis (Coilia) hamiltonii Gray, 1830 (= Mystus ramcarati Hamilton, 1822). Type by monotypy. Also appeared in Gray 1831:9 [ref. 1879] as a new genus. Valid as Coilia Gray, (see Whitehead 1967: [ref. 6464], Uyeno in Masuda et al. 1984:20 [ref. 6441], Grande 1985:245 [ref. 6466], Whitehead et al. 1988:460 [ref. 5725], Kottelat 1989:4 [ref ], Ataur Rahman 1989:240 [ref ], Kuang in Pan et al. 1991:42 [ref ], Liu 1995:558 [ref ], Wongratana et al. 1999:1699 [ref ], Ataur Rahman 2003:264 [ref ], Kottelat 2013:52 [ref ]). Current status: Valid as Coilia Gray, Engraulidae. (Chaetomus) [ref. 2927]. Masc. Choetomus playfairii McClelland, Type by subsequent designation. Type not by monotypy, second species included on p. 406 [earliest type designation not researched]. Usually spelled Choetomus (originally with a diphthong), but McClellan's derivation is clearly based on chaeto- a lock of hair. Synonym of Coilia Gray, (Whitehead et al. 1988:460 [ref. 5725] as Choetomus, Kottelat 2013:52 [ref ]). Current status: Synonym of Coilia Gray, Engraulidae. (Demicoilia) [ref. 2499]. Fem. Coilia quadragesimalis Valenciennes, Type by original designation (also monotypic). Synonym of Coilia Gray, (Whitehead 1967:149 [ref. 6464], Whitehead et al. 1988:460 [ref. 5725], Kottelat 2013:52 [ref ]). Current status: Synonym of Coilia Gray, Engraulidae.

24 (Leptonurus) [ref. 320]. Masc. Leptonurus chrysostigma Bleeker, Type by monotypy. Synonym of Coilia Gray, (Whitehead et al. 1988:460 [ref. 5725], Kottelat 2013:52 [ref ]). Current status: Synonym of Coilia Gray, Engraulidae. (Mystus) [ref. 4930]. Masc. Mystus clupeoides Lacepède, 1803 (= Clupea mystus Linnaeus, 1758). Type by monotypy. Objectively invalid; preoccupied by Mystus Scopoli, 1777 in fishes. In the synonymy of Coilia Gray, (Whitehead 1967:148 [ref. 6464], Whitehead et al. 1988:460 [ref. 5725], Kottelat 2013:51 [ref ]). Current status: Synonym of Coilia Gray, Engraulidae. (Thrissa) [ref. 993]. Fem. Clupea mystus Linnaeus, type by monotypy. Replacement name for Mystus Lacepède; appeared as 'Thrissa Cuv. Mystus Lac.'; therefore the type is the same as for Mystus Lacepède. Objectively invalid; preoccupied by Thrissa Rafinesque, Spelled Thryssa by Swainson 1838:279 [ref. 4302]. Treated as valid as Thrissa Cuvier, 1816 (Grande 1985:246 [ref. 6466], Whitehead et al. 1988:421 [ref. 5725], Kuang in Pan et al. 1991:40 [ref ]). Synonym of Coilia Gray, (Kottelat 2013:52 [ref ]). Current status: Synonym of Coilia Gray, Engraulidae. (Thrissocles) [ref. 2407]. Masc. Clupea mustus Linnaeus, Type by being a replacement name. Replacement for Thrissa Cuvier, Synonym of Thryssa Cuvier, (see Whitehead 1967:140 [ref. 6464], Whitehead & Wongratana 1986:206 [ref. 6284], Whitehead et al. 1988:421 [ref. 5725]). Synonym of Coilia Cuvier, (Kottelat 2013:52 [ref ]). Current status: Synonym of Coilia Cuvier, Engraulidae. Species Coilia borneensis Bleeker, Bornean grenadier anchovy [=Coilia borneensis Bleeker [P.], 1852:45, Coilia polyfilis Volz [W.], 1903:559] Notes: [Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen. v. 24 (8); ref ] Bandjarmasin, Borneo, Indonesia. Current status: Valid as Coilia borneensis Bleeker, Engraulidae. Distribution: Western Pacific. Habitat: freshwater, brackish, marine. (polyfilis) [Zoologischer Anzeiger v. 26 (no. 703); ref. 4531] Banju asin, Palembang, Sumatra, Indonesia. Current status: Synonym of Coilia borneensis Bleeker, Engraulidae. Habitat: freshwater, brackish, marine Species Coilia coomansi Hardenberg, Cooman's grenadier anchovy [=Coilia coomansi Hardenberg [J. D. F.], 1934:294]

25 Notes: [Treubia Buitenzorg v. 14; ref ] Lower course of Kapuas River, western Kalimantan [Borneo]. Current status: Valid as Coilia coomansi Hardenberg, Engraulidae. Distribution: Western Pacific. Species Coilia dussumieri Valenciennes, in Cuvier & Valenciennes, goldspotted grenadier anchovy [=Coilia dussumieri Valenciennes [A.], in Cuvier & Valenciennes, 1848:81, Pl. 610, Leptonurus chrysostigma Bleeker [P.], 1849:14, Demicoilia margaritifera Jordan [D. S.] & Seale [A.], 1926:363, Coilia quadrifilis Günther [A.], 1868:403] Notes: [Histoire naturelle des poissons v. 21; ref. 1013] Mumbai, India. Current status: Valid as Coilia dussumieri Valenciennes, Engraulidae. Distribution: Indo-West Pacific. Habitat: freshwater, brackish, marine. (chrysostigma) [Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen. v. 22 [8]; ref. 320] Madura Straits near Kammal and Surabaya, Java, Indonesia. Current status: Synonym of Coilia dussumieri Valenciennes, Engraulidae. Habitat: freshwater, brackish, marine. (margartifera) [Bulletin of the Museum of Comparative Zoology v. 67 (no. 11); ref. 2500] Colombo, Sri Lanka. Current status: Synonym of Coilia dussumieri Valenciennes, Engraulidae. Habitat: freshwater, brackish, marine. (quadrifilis) [Catalogue of the fishes in the British Museum v. 7; ref. 1990] Malay Peninsula. Current status: Synonym of Coilia dussumieri Valenciennes, Engraulidae. Habitat: freshwater, brackish, marine. Species Coilia grayii Richardson, Gray's grenadier anchovy [=Coilia grayii Richardson [J.], 1845:no p., Pl. 54 (figs. 1-2)] Notes: [Ichthyology.--Part 2. The zoology of the voyage of H. M. S. Sulphur; ref ] China Seas. Current status: Valid as Coilia grayii Richardson, Engraulidae. Distribution: Indo-West Pacific. Habitat: brackish, marine

26 Species Coilia lindmani Bleeker, Lindman's grenadier anchovy [=Coilia lindmani Bleeker [P.], 1857:48, Coilia macrognathus aequidentata Chabanaud [P.], 1924:59] Notes: [Acta Societatis Regiae Scientiarum Indo-Neêrlandicae v. 3; ref. 359] Mussi River, Palembang, Indonesia. Current status: Valid as Coilia lindmani Bleeker, Engraulidae. Distribution: Asia. Habitat: freshwater, brackish. (aequidentata) [Bulletin du Muséum National d'histoire Naturelle (Série 1) v. 30 (no. 1); ref ] Mouth of Saigon River, southern Vietnam. Current status: Synonym of Coilia lindmani Bleeker, Engraulidae. Habitat: freshwater, brackish. Species Coilia macrognathos Bleeker, longjaw grenadier anchovy [=Coilia macrognathos Bleeker [P.], 1852:436] Notes: [Natuurkundig Tijdschrift voor Nederlandsch Indië v. 3; ref ] Pamangkat, Indonesia. Current status: Valid as Coilia macrognathos Bleeker, Engraulidae. Distribution: Western Pacific. Species Coilia mystus (Linnaeus, 1758) - Osbeck's grenadier anchovy [=Clupea mystus Linnaeus [C.], 1758:319, Mystus clupeoides Lacepède [B. G. E.], 1803:466, 467, Coilia mystus jiulongjiangensis Liu [W.-B.], 1995:562 [English p. 565], Choetomus playfairii McClelland [J.], 1844:405, Pl. 24 (fig. 3), Osteoglossum prionostoma Basilewsky [S.], 1855:244] Notes: [Systema Naturae, Ed. X v. 1; ref. 2787] Indian Ocean. Current status: Valid as Coilia mystus (Linnaeus, 1758). Engraulidae. Distribution: Western Pacific. Habitat: freshwater, brackish, marine. (clupeoides) [Histoire naturelle des poissons (Lacepéde) v. 5; ref. 4930] Indian Ocean. Current status: Synonym of Coilia mystus (Linnaeus, 1758). Engraulidae. Habitat: freshwater, brackish, marine.

27 (jiulongjiangensis) [Oceanologica et Limnologica Sinica v. 26 (no. 5); ref ] Jiulongjiang, China. Current status: Synonym of Coilia mystus (Linnaeus, 1758). Engraulidae. Habitat: freshwater, brackish, marine. (playfairii) [Calcutta Journal of Natural History v. 4 (for 1843); ref. 2927] China. Current status: Synonym of Coilia mystus (Linnaeus, 1758). Engraulidae. Habitat: freshwater, brackish, marine. (prionostoma) [Nouveaux mémoires de la Société impériale des naturalistes de Moscou v. 10; ref. 200] Gulf of Tschiliensis, China. Current status: Synonym of Coilia mystus (Linnaeus, 1758). Engraulidae. Habitat: freshwater, brackish, marine. Species Coilia nasus Temminck & Schlegel, Japanese grenadier anchovy [=Coilia nasus Temminck [C. J.] & Schlegel [H.], 1846:243, Pl. 109 (fig. 4), Coilia brachygnathus Kreyenberg [W.] & Pappenheim [P.], 1908:96, Coilia ectenes Jordan [D. S.] & Seale [A.], 1905:517, Fig. 1, Coilia rendahli Jordan [D. S.] & Seale [A.], 1926:362, Coilia ectenes taihuensis Yuan [C.-M.], Lin [J.-B.], Liu [J.-H.] & Qin [A.-L.], 1977:135] Notes: [Fauna Japonica Parts 10-14; ref. 4374] Japan. Current status: Valid as Coilia nasus Temminck & Schlegel, Engraulidae. Distribution: Western North Pacific. Habitat: freshwater, brackish, marine. (brachygnathus) [Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1908; ref ] Tungting Lake; Hankau, east central China. Current status: Synonym of Coilia nasus Temminck & Schlegel, Engraulidae. Distribution: Asia. Habitat: freshwater. (ectenes) [Proceedings of the United States National Museum v. 29 (no. 1433); ref ] Shanghai, China. Current status: Synonym of Coilia nasus Temminck & Schlegel, Engraulidae. Habitat: freshwater, brackish, marine. (rendahli) [Bulletin of the Museum of Comparative Zoology v. 67 (no. 11); ref. 2500] Shanghai, China. Current status: Synonym of Coilia nasus Temminck & Schlegel, Engraulidae. Habitat: freshwater, brackish, marine. (taihuensis) [On a new subspecies of Coilia from China -- Coilia ectenes taihuensis ssp. nov. Pp in: Yangtse River region Coilia resources investigational report.; ref ] Taihu, Kiangsu Province, China. Current status: Synonym of Coilia nasus Temminck & Schlegel, Engraulidae. Habitat: freshwater, brackish, marine.