Macrobenthic invertebrate survey of Waneta and Lamoka Lakes, Schuyler County, New York, September, 1996.

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1 133 Macrobenthic invertebrate survey of Waneta and Lamoka Lakes, Schuyler County, New York, September, Matthew Albright Willard N. Harman INTRODUCTION Schuyler County encompasses 331 square miles of the Appalachian Plateau in the Finger Lakes region (Curatolo, 1991). The county's economy is largely tourism-based, relying heavily upon its aquatic natural resources. The eutrophication of some of these water bodies, including Waneta and Lamoka Lakes (Figure 1), has had detrimental impacts on traditional lakes uses. Macrophyte growth and algal blooms have negatively impacted swimming. fishing, boating, and the perceived aesthetic qualities oftheses areas. These changes are believed to have lead to a decline in both tourism activities and in lakeside property values. In order to address these concerns, Schuyler County initiated an aquatic vegetation control program in 1986 (Curatolo. 1991). This program was to serve as an integrated approach to manage the county's aquatic resources through monitoring, research, macrophyte harvesting, upland treatment. and public education. In a comprehensive study was undertaken in order to provide baseline iniormation upon which lake management plans would be based (Curato ). This work included a characterization ofthe physical qualities of local lakes. surveys oftheir macroinvertebrate benthic populations. a description of macrophyte harvesting activities, a determination of the feasibility of conducting dredging operations in selected areas. and addressed means of reducing nutrient inputs to these lakes. This document describes the tirst of a two part benthic survey of Waneta and Lamoka Lakes. conducted 16 September A second sampling is planned for May, 1997 in order to ensure that various life stages of organisms are represented. This database. when compared to previous work (i. e. Curatolo. 1991) and future works, will indicate the impacts of macroph}te management on the benthic faunal communities. The background material for this report is derived almost exclusively from Curatolo BACKGROUND Waneta Lake (Figure 2) covers 781 acres in surface area. of which over 90% is located in Schuyler County. The lake occupies a volume of approximately acre-feet and has about 6.8 miles ofshoreline. This eutrophic lake has a maximum depth of<30 feet. Approximately 450 year-round and seasonal dwellings and a Boy Scout camp line the shore. The substrate varies

2 134 LAMOKA LAKE Figure I. The location of Waneta, Lamoka, and Seneca Lakes in Schuyler County, New York (modified from Curatolo, 1991).

3 .. '6' :: ,', ,,:1 5 "~ i, \I.~ I ;I~!/~ I I~ i ~ i~1 ~!~, rt\ \c.,) Q.7!~I! all J~.\.,.\. '~,.'1\",.... '.). '/ I i I ',3 I,, I.\ 1" "\ I " 4 ~._.-... EB o Scout Point BoAT LAUNCH SITES UNLOADING EXTENT OF SAMPLING PUBLIC Camp N SITES VEGETATION STATION ACCESS 1000 fl 135 Figure 2. Waneta Lake, Schuyler County, New York, showing sampling stations (modified from Curatolo, 1991).

4 136 from silty to organic clay. Eurasian milfoil (Myriophyllum <spicatum) dominated most ofthe shallows, with plant diversity being considerably higher at the southernmost shore. At the time of sampling (16 September), all substrates were oxygenated. Lamoka Lake (Figure 3) is interconnected with Mill Pond, located in western Schuyler County at 1099 feet in elevation. These shallow «47 ft) eutrophic lakes together occupy 826 acres in surface area, acre-feet in volume, and have approximately 11.3 miles ofshoreline. In 1990, about 325 seasonal and year-round houses occupied the shoreline. Recreational activities include swimming, fishing, and boating (Curatolo, 1991). Substrates were dominated by anaerobic organic muds; a notable exception was observed at a drop-off at the northeastern shore, where compact sand was encountered. Here, evidence ofunionid clams (i.e. shells) were seen. The predominating macrophytes ~ncountered were the nonindigenous Eurasian milfoil (Myriophyllum <spicatum) and curly leafed pondweed (Potamoge/on crispus). Toward the southern, shallower end of the lake diversity was considerably higher, with Nuphar. Ceratophyllum demersum, He/eran/hera. and Vallisineria being well represented. Interestingly, at the time of sampling milfoil was absent from the Mill Pond area. Similar flora were encountered along the channel connecting Waneta and Lamoka Lakes. Access is provided to both lakes by boat launch sites maintained by the New York State Department ofconservation (NYSDEC). Neither lake is used as a potable water supply. Both lakes have been rated as moderately impaired on the NYSDEC 1989 Priority Problem List (NYSDEC, 1989). Despite inclement weather encountered throughout the day of sampling (45-50 F, steady rain), bird activity was considerable. Large numbers oftree swallows (lridoprocne hieolor) were active on Waneta Lake, presumably feeding on emerging aquatic insects. Two ospreys (Pandion haliae/us) and parasitic jaeger (S/ercorarius parasi/iclls) were seen over Lamoka Lake; the latter seabird is extremely rare in this region and was likely displaced by recent tropical storms (Butts, 1996). MONITORING AND WATER QUALITY Temperature, dissolved oxygen, ph. and conductivity were measured using a Hydrolab Scout II multiparameter water quality monitoring instrument which had been calibrated the day of data collection following manufacturer's operating manual (Hydro lab Corp., 1993). Readings were taken at I meter (m) intervals from the surface to the bottom. The maximum depth encountered at Waneta Lake was 8.1 meters (26.6 feet); that for Lamoka Lake was 12.2 meters (40.0 feet). Waneta was undergoing fall overturn, although slight. presumably temporary stratification was observed below 8 m. Here, temperature was approxin1ately 0.3 C less than the rest of the water column and dissolved oxygen was over 2 mg/lless than overlaying waters. Conductivity was between mmho/cm throughout. Temperature, dissolved oxygen. and ph profiles are graphically presented in Figure 4a.

5 137 FLEET!A-...-COVE /000 ft t N CREEK o BOAT LAUNCH SITES PUBLIC ACCESS UNLOADING SITES ~ -._.- EXTENT OF VEGETATION ffi SAMPLING STATION POND Figure 3. Lamoka Lake, Schuyler County, New York, showing sampling stations (modified from Curatolo, 1991).

6 138 A B l _.J L-..--L_J._ I _L~...L_l_..L-l.l.----.l.-_.l _I~L~ I, I I l 3 3 I l I a -., 4 5 '-' 6 os 0. II) Cl a -., 4 5 '-' 6 os 0. II) Cl i I li - I 1 j I 1'\ 12 I I 11 I.J 12 _ " Temperature -ph _ Dissolved Oxygen Figure 4. Profiles of temperature, ph, and dissolved oxygen for Waneta (A) and Lamoka (B) Lakes, 17 September, 1986.

7 139 Lamoka Lake was stratified at the time of sampling, with the thermocline located between 6 and 7 m. Hypolimnetic waters were essentially anoxic «0.3 mg/l), a situation similar to that reported by Curatolo (1991) for corresponding dates in 1988, 1989 and Conductivity was approximately 190 mmho/cm through the epi1imnion and increased below the thermocline to 213 mmho/cm at the bottom. Temperature, dissolved oxygen, and ph profiles are graphically presented in Figure 4b. Water transparency was measured using a standard 20 cm Secchi disk. Transparency was 1.9 m (6.2 ft) in Lamoka and 2.0 m (6.6 ft) in Waneta. Values reported by Curatolo (1991) indicate that Lamoka was somewhat more transparent and Waneta twice as transparent in 1996 compared to similar dates in recent years. BENTHIC INVERTEBRATE ANALYSIS METHODS Macroinvertebrates were collected from nine sites on Waneta Lake and nine sites on Lamoka Lake. These sites included eulittoral, littoral benthic, littoral vegetative, and profundal areas. The objective was to obtain taxa diversity and density data for comparison with previous (e.g. Curatolo, 1991) and future surveys in order to ascertain any effects of macrophyte and algae control programs, as well as other changes in water quality, on these organisms. Benthic samples were collected in triplicate using either a 23X23 cm 2 or a 15X15 cm 2 Ekman dredge. Larger dredges, using a conventional cable and messenger, were used in deeper locations; in shallower areas, smaller dredges were employed using extension handles. Eulittoral samples, where the substrate tended to be stony, were acquired by manually gathering all material within a 23X23 cm 2 quadrangle. Vegetative samples were collected by sweeping the macrophytes with a triangle net five times and rinsing the contents from the net. Effort was made to include the entire depth distribution of the macrophyte bed. Each sample was collected in quadruplicate. While it is recognized that these vegetative samples cannot be interpreted as truly quantitative. the attempt was made to standardize the method as much as possible so that future comparisons may be made. Upon retrieval, multiple samples were composited and passed through a #30 mesh brass screen. thus retaining all particles over 583 microns. This material was transferred to one-gallon plastic jars, to which 95% ethanol was added until a final concentration of approximately 70% ethanol was reached. Several milliliters ofrose bengal were added to each sample to later aid in the recognition ofbenthic organisms. Upon return to the laboratory. organisms were separated from the substrate by transferring each sample. spoonful at a time, into a white enamel pan and rinsing with adequate water to distribute the material. All benthic organisms were plucked and transferred to 6 dram vials containing 70% ethanol. Taxonomic identifications were made according to Pennak (1989), Peckarsky et al. (1990), and Merritt and Cummins (1996).

8 140 Organisms were enumerated and weighed by taxa. Wet weight was taken by removing the organisms from their vials and setting on blotting paper for 15 minutes prior to weighing to the nearest 0.1 mg on an electronic balance (Wheat, 1993). No attempt was made to count Oligochaetes, as they tended to fragment during processing. For benthic sites, where sample size was more standardized, data were converted to units per mete~ RESULTS AND DISCUSSION A description ofeach sample site, including sample type, sampling method, substrate characterization, and dominant macrophytes, is summarized in Table 1 (refer to Figure 1 for site locations). This survey revealed a total of 58 taxa; 44 and 45 taxa were encountered in Waneta Lake and Lamoka Lakes, respectively (Table 2). Generally, diversity was greater in Lamoka, where the number of taxa per site averaged 15.0, compared to 12.2 taxa per site found in Waneta. The southernmost reaches of Lamoka Lake (sites L6-L9) exhibited the greatest diversity. Here, 32 taxa were discovered, with each site averaging This area is relatively shallow and macrophytes were likewise diverse. OveralL the Order Trichoptera (caddis flies) showed the greatest diversity, with 7 genera representing 3 families being encountered. Tables 3-20 describe the macrobenthic invertebrates for each site visited, including numbers and wet weights of each taxa found in Waneta and Lamoka Lakes. For benthic sites, abundance and biomass have been projected to units/mete~. A more intensive interpretation of these data will follow a second sampling to occur in May, This later survey will complement the present data in that larval aquatic stages not present in the fall will be included. REFERENCES Butts, W.L Personal communication. Biological Field Station, Cooperstown, NY. Curatolo, J Final Report. The aquatic vegetation control program in Schuyler County, p. Schuller County Soil and Water Conservation District, Montour Falls, NY. Hydrolab Corporation Scout II operating manual. Hydrolab Corp. Austin, TX. Merritt, R.W., and K.W Cummins (eds.) Aquatic insects of North America. Kendall/Hunt Publishing Company. Dubuque, IA. New York State Dept. Of Envir. Cons., New York State water quality, Bureau of Monitoring and Assessment. Division of Water. NYSDEC. Albany, NY. Peckarsky, B.L., P.R. Fraissinet, M.A. Penton, and OJ. Conklin, Jr Freshwater macroinvertebrates of Northeastern North America. Cornell University Press. Ithaca, NY. Pennak, R.W Freshwater invertebrates of the United States, 3rd Ed. John Wiley and Sons, Inc. New York.

9 141 Site Sample Type Dominant Vegetation Substrate Characterization Waneta WI Sub., 3 small dredges Myriophyllum spicatum organic clay W2 Sub., 3 small dredges M spicatum organic clay W3 Veg. 4X5 sweeps M spicatum organic clay W4 Sub., 3 large dredges none fine silty clay W5 Veg. 4X5 sweeps organic clay organic clay W6 Sub., 3 small dredges M spicatum organic detritus W7 Sub., 1 quadrangle none stones (eulittoral) W8 Veg. 4X5 sweeps Ceratophyllum demersum organic clay W9 Veg. 4X5 sweeps Nuphar variegatum, Valisanaria organic mud americana Lamoka L1 Sub., 3 large dredges L2 Sub., 3 small dredges L3 Veg. 4X5 sweeps L4 Veg. 4X5 sweeps L5 Sub., 3 small dredges L6 Veg. 4X5 sweeps L7 Veg. 4X5 sweeps L8 Veg. 4X5 sweeps L9 Veg. 4X5 sweeps none none M spicatum M spicatum M spicatum Ai spicatum N variegatm M spicatum Heteranthera dubia, C. demersum, V americana organic mud (anoxic) compact sand organic mud peaty organic mud organic mud organic mud organic mud organic mud organic mud Table 1. Description of macrobenthic invertebrate sampling sites, Waneta and Lamoka Lakes, 16 September Sub.= substrate sample, Veg.= vegetative sample, small dredge=.0232 m 2, large dredge=.0523 m 2, quadrangie=.0523 m 2

10 142 All Sites, 9/16/96 Waneta Lamoka Platyhelminthes Turbellaria Tricladida Planariidae Nemotoda Annelida Oligochaeta Haplotaxida Naididae Tubificidae Lumbriculida Lumbriculidae Hirudinea Pharyngobdellida Erpobdellidae Erpobdella Dina parva Rhynchobdellida Glossiphoniidae HeJobdeJJa He/obdella fusca HeJobdeJJa stagnalis Batracobdella PlacobdeJJa Piscicolidae A4yzobdeJJalugubris Mollusca Bivalvia Paleoheterodonta Unionidae Lampsilis radiata E/Jiptio complanatl/s Veneroida Sphaeriidae Sphaerium Pisidium Gastropoda Basommatrophora Lymnaeidae Lymnaea cojumejja Table 2. Summary of macrobenthic invertebrates collected from Waneta and Lamoka Lakes, 16 September, 1996.

11 143 Waneta Lamoka Planorbldae Gyraulus parvus * * Gyraulus hirsutus * * Promentus exaeuous * Physidae Physa integra * Physa sayii * * Mesogastropoda Valvatidae Valvata triearinata * * Hydrobiidae Amnieo/a limosa * * Arthropoda Arachnida Acariformes Hyd rodromidae Hydrodroma * * Limnesiidae * Pionidae * Arrenuridae Arrenurus * Crustacea Isopoda Asellidae Caeeidotea * * Amphipoda Gammaridae Gammarus * * Talitridae Hyalella azteea * * Insecta Ephemeroptera Baetidae Paraeloeodes * * Caenidae Caenis * * Odonata (Anisoptera) Libellulidae Erythemis * * Corduliidae Epitheea * Table 2 (cont.). Summary of macrobenthic invertebrates collected from Waneta and Lamoka Lakes, 16 September, 1996.

12 144 /Waneta Lamoka Odonata (Zygoptera) Coenagrionidae Coenagrion or Enallagma * * Lestidae Lestes * Hemiptera Pleidae Paraplea * Mesoveliidae Mesovelia * * Lepidoptera Nepticulidae * Pyralidae Acentria * * Trichoptera Leptoceridae Oecetis * Ylodes * Leptocerus * * Hydroptilidae Orthatrichia * * Oxythira * * Hydoptilla * Polycentropodidae Cemotina * Coleoptera Curculionidae * * Dytisicidae * Chrysomelidae Pyrrha/ta * Diptera Ceratopogonidae I Sphaeromias * * Probezzia * * Dasyhelea * Bezzia or Palpomyia * * Chaoboridae Chaoborus * * Chironomidae * * Table 2 (cont.). Summary of macrobenthic invertebrates collected from Waneta and Lamoka Lakes, 16 September, 1996.

13 145 Waneta Lake, Site 1 (benthic) 9/16/96 #/sample g/sample #/m"2 g/m"2 Platynelmlntnes Turbellaria Tricladida Planariidae Annelida Oligochaeta Haplotaxida Tubificidae Lumbriculida Lumbriculidae Hirudinea Rhynchobdellida Glossiphoniidae He/obdella stagnalis Batracobdella Placobdella Mollusca Gastropoda Mesogastropoda Hydrobiidae Amnico/a limosa Arthropoda Crustacea Isopoda Asellidae Caecidotea Amphipoda Gammaridae Gammarus Talitridae Hyalella azteca Insecta Diptera Ceratopogonidae Sphaeromias Chaoboridae Chaoborus Chironomidae NA NA NA NA Table 3. Summary ofmacrobenthic invertebrates collected from Waneta Lake, Site #1, 9/16/96 (see Figure 2 for site locations).

14 146 Waneta Lake, Site 2 (benthic) 9/16/96 #/sample g/sample #/m"2 g/m"2 IPlatyhelminthes Turbellaria Tricladida Planariidae Nematoda Annelida Oligochaeta Lumbriculida Lumbriculidae NA NA Mollusca Gastropoda Mesogastropoda Hydrobiidae Amnicola limosa Arthropoda Crustacea Amphipoda Talitridae Hyalel/a azteca Insecta Trichoptera Hydroptilidae Orthotrichia Diptera Ceratopogonidae Sphaeromias Chaoboridae Chaoborus Chironomidae Table 4. Summary of macrobenthic invertehrates collected from Waneta Lake, Site #2, 9/16/96 (see Figure 2 for site locations).

15 147 Waneta Lake, Site 3 (vegetative) 9/16/96 #/sample g/sample PlatyhelminThes Turbellaria Tricladida Planariidae Annelida Oligochaeta Haplotaxida Naididae NA Mollusca Gastropoda Basommatrophora Planorbidae Gyraulus parvus Mesogastropoda Valvatidae Valvata tn'carinata Hydrobiidae Amnicola limosa Arthropoda Arachnida Acariformes Hydrodromidae Hydrodroma Crustacea Amphipoda Talitridae Hyalella azteca Insecta Ephemeroptera Caenidae Caenis Diptera Chironomidae Table 5. Summary ofmacrobenthic invertebrates collected from Waneta Lake, Site #3,9/16/96 (see Figure 2 for site locations).

16 148 Waneta Lake, Site 4 (benthic) 9/16/96 #/sample g/sample #/m"2 g/m"2 Annelida Oligochaeta Lumbriculida Lumbriculidae Arthropoda Insecta Diptera Chaoboridae Chaoborus Chironomidae NA NA Table 6. Summary of macrobenthic invertebrates collected from Waneta Lake, Site #4, 9/16/96 (see Figure 2 for site locations).

17 149 Waneta Lake, Site 5 (vegetative) 9/16/96 #/sample g/sample Platynelmlnthes Turbellaria Tricladida Planariidae Annelida Oligochaeta Haplotaxida Naididae NA Mollusca Gastropoda Basommatrophora Planorbidae Gyraulus panus Physidae Physa Mesogastropoda Hydrobiidae Amnicola limosa Arthropoda Crustacea Amphipoda Talitridae Hya/e/la azteca Insecta Ephemeroptera Baetidae Paracloeodes Caenidae Caenis Odonata (Zygoptera) Coenagrionidae Coenagrion or Ena//agma Lepidoptera Nepticulidae Trichoptera Hydroptilidae Orthotrichia Oxythira Hydoptilla Coleoptera Curculionidae Table 7. Summary ofmacrobenthic invertebrates collected from Waneta Lake, Site #5, 9/16/96 (see Figure 2 for site locations).

18 150 #/sample g/sample Diptera Ceratopogonidae Bezzia or Palpomyia Chaoboridae Chaoborus Chironomidae Table 7 (cont.). Summary of macrobenthic invertebrates collected from Waneta Lake, Site #5, 9/16/96 (see Figure 2 for site locations).

19 151 Waneta Lake, Site 6 (benthic) 9/16/96 #/sample g/sample #/m"2 g/m"2 IPlatyhelminthes Turbellaria Tricladida Planariidae Annelida Oligochaeta Haplotaxida Naididae NA NA Tubificidae NA NA Lumbriculida Lumbriculidae NA NA Hirudinea Rhynchobdellida Glossiphoniidae Helobdella stagnalis Mollusca Gastropoda Basommatrophora Planorbidae Gyraulus parvus Mesogastropoda Hydrobiidae Amnicola limosa Arthropoda Arachnida Acariformes Hydrodromidae Hydrodroma 'I Limnesiidae Crustacea Amphipoda Talitridae Hyalella azteca Insecta Ephemeroptera Caenidae Caenis Odonata (Zygoptera) Coenagrionidae Coenagrion or Enallagma Table 8. Summary ofmacrobenthic invertebrates collected from Waneta Lake, Site #6, 9/16/96 (see Figure 2 for site locations).

20 152 #/sample g/sample #/mi\2 g/ml\2 Lepidoptera Nepticulidae Coleoptera Dytisicidae Diptera Ceratopogonidae Probezzia Bezzia or Palpomyia Chaoboridae ChaobonJs Chironomidae Table 8 (cont.). Summary of macrobenthic invertebrates collected from Waneta Lake, Site #6, 9/16/96 (see Figure 2 for site locations).

21 153 Waneta Lake, Site 7 (benthic) 9/16/96 #/sample g/sample #/m"2 g/m"2 IPlatyhelminthes Turbellaria Tricladida Planariidae Annelida Oligochaeta Haplotaxida Naididae NA NA Tubificidae NA NA Lumbriculida Lumbriculidae NA NA Hirudinea Pharyngobdellida Erpobdellidae Erpobdella Arthropoda Crustacea Amphipoda Gammaridae Gammarus Insecta Diptera Chironomidae Table 9. Summary of macrobenthic invertebrates collected from Waneta Lake, Site #7, 9/16/96 (see Figure 2 for site locations).

22 154 Waneta Lake, Site 8 (vegetative) 9/16/96 #/sample g/sample Platyh-elmlnthes Turbellaria Tricladida Planariidae Annelida Oligochaeta Haplotaxida Naididae NA Hirudinea Pharyngobdellida Erpobdellidae Dina pana Rhynchobdellida Piscicolidae A4yzobdeNalugubris Mollusca Gastropoda Basommatrophora Planorbidae Gyraulus panus Physidae Physa integra Mesogastropoda Valvatidae Valvata tricarinata Hydrobiidae Amnicola limosa Arthropoda Arachnida Acariformes Pionidae Crustacea Amphipoda Talitridae HyaleNa azteca Insecta Ephemeroptera Caenidae Caenis Table 10. Summary of macrobenthic invertebrates collected from Waneta Lake, Site #8, 9/16/96 (see Figure 2 for site locations).

23 155 #/sample g/sample Odonata (Zygoptera) Coenagrionidae Coenagrion or Enal/agma Lepidoptera Nepticulidae Diptera Ceratopogonidae Bezzia Chironomidae or Palpomyia Table 10 (cont.). Summary ofmacrobenthic invertebrates collected from Waneta Lake, Site #8, 9/16/96 (see Figure 2 for site locations).

24 156 Waneta Lake, Site 9 (vegetative) 9/16/96 #/sample g/sample rplatynelmrnthes Turbellaria Tricladida Planariidae Mollusca Bivalvia Veneroida Sphaeriidae Sphaerium Gastropoda Basommatrophora Lymnaeidae Lymnaea columella Planorbidae Gyraulus hirsutus Physidae Physa sayii Mesogastropoda Hydrobiidae Amnicola limosa Arthropoda Crustacea Isopoda Asellidae Caecidotea Amphipoda Talitridae Hya/ella azteca Insecta Ephemeroptera Caenidae Caenis Odonata (Anisoptera) LibelluJidae Erythemis Odonata (Zygoptera) Coenagrionidae Coenagrion or Enallagma Table 11. Summary of macrobenthic inv~rtebrates collected from Waneta Lake, Site #9, 9/16/96 (see Figure 2 for site locations).

25 157 #/sample g/sample Hemiptera Mesoveliidae Mesovelia Lepidoptera Nepticulidae Pyralidae Acentria Trichoptera Leptoceridae Leptocerus Coleoptera Curculionidae Chrysomelidae Pyrrhalta Diptera Ceratopogonidae Bezzia or Palpomyia Chironomidae Table 11 (cont.). Summary of macrobenthic invertebrates collected from Waneta Lake, Site #9, 9/16/96 (see Figure 2 for site locations).

26 158 Lamoka Lake, Site 1 (benthic) 9/16/96 #/sample g/sample #/m!l2 g/m!l2 Annelida Oligochaeta Haplotaxida Tubificidae Lumbriculida Lumbriculidae Arthropoda Insecta Diptera Chaobol idae Chaoborus Chironomidae NA NA NA NA Table 12. Summary ofmacrobenthic invertebrates collected from Lamoka Lake, Site # 1, 9/16/96 (see Figure 3 for site locations).

27 159 Lamoka Lake, Site 2 (benthic) 9/16/96 #/sample g/sample #/m A 2 g/ml\2 IAnnelida Oligochaeta Haplotaxida Naididae NA NA Tubificidae NA NA Lumbriculida Lumbriculidae NA NA Hirudinea Rhynchobdellida Glossiphoniidae He/obdella stagnalis Mollusca Bivalvia Paleoheterodonta Unionidae * Lampsilis radiata (shells only) * Elliptio comp/anatus (shells only) Veneroida Sphaeriidae Sphaerium Gastropoda Mesogastropoda Hydrobiidae Amnico/a /imosa Arthropoda Crustacea Amphipoda Talitridae Hya/ella azteca Insecta Ephemeroptera Caenidae Caenis Diptera Ceratopogonidae Probezzia Chironomidae Table 13. Summary ofmacrobenthic invertebrates collected from Lamoka Lake, Site #2, 9116/96 (see Figure 3 for site locations).

28 1 160 Lamoka Lake, Site 3 (vegetative) 9/16/96 #/sample g/sample IPlatyhelminthes Turbellaria Tricladida Planariidae Annelida Oligochaeta Haplotaxida Naididae NA Hirudinea Rhynchobdedida Glossiphoniidae Helobdella stagnalis Mollusca Bivalvia Veneroida Sphaeriidae Sphaerium Gastropoda Basommatrophora Planorbidae Gyraulus parvus Promentus exacuous Physidae Physa sayii Mesogastropoda Valvatidae Valvata tricarinata Hydrobiidae Amnicola limosa Arthropoda Crustacea Isopoda Asellidae Caecidotea Amphipoda Gammaridae Gammarus Talitridae Hyalella azteca Table 14. Summary of macrobenthic invertebrates collected from Lamoka Lake, Site #3, 9/16/96 (see Figure 3 for site locations).

29 161 Insecta Odonata (Anisoptera) Corduliidae Epitheca Odonata (Zygoptera) Coenagrionidae Coenagrion or Enal/agma Trichoptera Leptoceridae Leptocerus Diptera Ceratopogonidae Bezzia or Palpomyia Chironomidae #/sample g/sample '1 Table 14 (conl). Summary of macrobenthic invertebrates collected from Lamoka Lake, Site #3, 9/16/96 (see Figure 3 for site locations).

30 162 Lamoka Lake. Site 4 (vegetative) 9/16/96 #/sample g/sample IPlatyhelminthes Turbellaria Tricladida Planariidae Annelida Oligochaeta Haplotaxida Naididae NA Hirudinea RhynchobdelHda Glossiphoniidae Helobdel/a stagnalis Mollusca Gastropoda Basommatrophora Planorbidae Gyraulus parvus Physidae Physa sayii Mesogastropoda Valvatidae Valvata tricarinata Hydrobiidae Amnicola limosa Arthropoda Crustacea Amphipoda Talitridae Hyalel/a azteca Insecta Ephemeroptera Caenidae Caenis Odonata (Zygoptera) Coenagrionidae Coenagrion or Ena/lagma Diptera Chironomidae Table 15. Summary of macrobenthic invertebrates collected from Lamoka Lake, Site #4, 9/16/96 (see Figure 3 for site locations).

31 163 Lamoka Lake, Site 5 (benthic) 9/16/96 #/sample g/sample #/m/l2 g/m/l2 IAnnellda Oligochaeta Lumbriculida Lumbriculidae NA NA Arthropoda Crustacea Amphipoda Talitridae Hyalella azteca Insecta Diptera Ceratopogonidae Sphaeromias Chaoboridae Chaoborus Chironomidae Table 16. Summary of macrobenthic invertebrates collected from Lamoka Lake, Site #5, 9/16/96 (see Figure 3 for site locations).

32 164 Lamoka Lake, Site 6 (vegetative) 9/16/96 #/sample g/sample IPlatynelmlnthes Turbellaria Tricladida Planariidae Annelida Oligochaeta Haplotaxida Naididae NA Hirudinea Rhynchobdeilida Glossiphoniidae Helobdella fusca Mollusca Bivalvia Veneroida Sphaeriidae Sphaerium Pisidium Gastropoda Basommatrophora Planorbidae Gyraulus parvus Gyraulus hirsutus Promentus exacuous Physidae Physa sayii Mesogastropoda Hydrobiidae Amnicola limosa Arthropoda Arachnida Crustacea Isopoda Asellidae Caecidotea Amphipoda Talitridae Hyalella azteca Table 17. Summary of macrobenthic invertebrates collected from Lamoka Lake, Site #6, 9/16/96 (see Figure 3 for site locations).

33 165 Insecta Ephemeroptera Baetidae Paracloeodes Caenidae Caenis Odonata (Anisoptera) Corduliidae Epitheca Odonata (Zygoptera) Coenagrionidae Coenagrion or Enallagma Trichoptera Leptoceridae Oecetis Leptocerus Hydroptilidae Oxythira Diptera Ceratopogonidae Bezzia or Palpomyia Chironomidae #/sample g/sample Table 17 (cont.). Summary of macrobenthic invertebrates collected from Lamoka Lake, Site #6, 9/16/96 (see Figure 3 for site locations).

34 166 Lamoka Lake, Site 7 (vegetative) 9/16/96 Platyhelminthes Turbellaria Tricladida Planariidae Hirudinea Rhynchobdellida Glossiphoniidae He/obdeJla stagnalis BatraeobdeJla Mollusca Bivalvia Veneroida Sphaeriidae Sphaerium Gastropoda Basommatrophora Planorbidae Gyrau/us hirsutus Mesogastropoda Hydrobiidae Amnieo/a /imosa Arthropoda Arachnida Acariformes Hydrodromidae Hydrodroma Crustacea Isopoda Asellidae Caeeidotea Amphipoda Talitridae Hya/e//a azteea Insecta Ephemeroptera Baetidae Parae/oeodes Caenidae Caenis #/sample g/sample Table 18. Summary of macrobenthic invertebrates collected from Lamoka Lake, Site #7,9/16/96 (see Figure 3 for site locations).

35 167 Odonata (Anisoptera) Libellulidae Erythemis Corduliidae Epitheca Odonata (Zygoptera) Coenagrionidae Coenagrion or Ena//agma Hemiptera Pleidae Parap/ea Mesoveliidae Mesove/ia Trichoptera Leptoceridae Y/odes Leptocerus Polycentropodidae Cemotina Diptera Ceratopogonidae Dasyhe/ea Bezzia or Pa/pomyia Chironomidae #/sample g/sample Table 18 (cont.). Summary of macrobenthic invertebrates collected from Lamoka Lake, Site #7, 9/l6/96 (see Figure 3 for site locations).

36 168 Lamoka Lake, Site 8 (vegetative) 9/16/96 #/sample g/sample IPlatyhelminthes Turbellaria Tricladida Planariidae Annelida Oligochaeta Haplotaxida Naididae NA Mollusca Bivalvia Veneroida Sphaeriidae Pisidium Gastropoda Basommatrophora Physidae Physa sayii Hydrobiidae Amnico/a limosa Arthropoda Arachnida Acariformes Arrenuridae Arrenurus Crustacea Amphipoda Talitridae Hyalel/a azteea Insecta Ephemeroptera Baetidae Parae/oeodes Caenidae Caenis Odonata (Anisoptera) Corduliidae Epitheea Odonata (Zygoptera) Coenagrionidae Coenagrion or Enallagma Table 19. Summary of macrobenthic invertebrates collected from Lamoka Lake, Site #8, 9/16/96 (see Figure J for site locations).

37 169 #/sample g/sample Lepidoptera Nepticulidae Pyralidae Acentria Trichoptera Leptoceridae Oecetis Leptocerus Hydroptilidae Orthotrichia Oxythira Polycentropodidae Cernotina Diptera Ceratopogonidae Dasyhe/ea Bezzia or Palpomyia Chaoboridae Chaoborus Chironomidae Table 19 (cont.). Summary ofmacrobenthic invertebrates collected from Lamoka Lake, Site #8, 9/16/96 (see Figure 3 for site locations).

38 170 Lamoka Lake,Site 9 (veqetative) 9/16/96 PlatYhelminthes Turbellaria Tricladida Planariidae Annelida Oligochaeta Haplotaxida Naididae Hirudinea #/sample 18 NA g/sample Rhynchobdellida Glossiphoniidae Helobdella Batracobdella Mollusca Bivalvia Veneroida Sphaeriidae Sphaerium Gastropoda Basommatrophora Planorbidae Gyraulus parvus Physidae Physa sayii Mesogastropoda Hydrobiidae Amnicola limosa Arthropoda Crustacea Amphipoda Talitridae Hyalella azteca Insecta Ephemeroptera Caenidae Caenis Odonata (Anisoptera) Corduliidae Epitheca Table 20. Summary of macrobenthic invertebrates collected from Lamoka Lake, Site #9, 9/16/96 (see figure 3 for site locations).

39 171 #/sample g/sample Odonata (Zygoptera) Coenagrionidae Coenagrion or Enallagma Lestidae Lestes Lepidoptera Nepticulidae Trichoptera Leptoceridae Oeeetis Ylades Leptacerus Polycentropodidae Cematina Coleoptera Curculionidae Diptera Ceratopogonidae Dasyhelea Bezzia or Palpomyia Chaoboridae Chaoborus Chironomidae Table 20 (cont.). Summary of macrobenthic invertebrates collected from Lamoka Lake, Site #9, 9/16/96 (see Figure 3 for site locations).