Taxonomical updates in The Scorpion Files for Scorpionidae (2008 )

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1 Taxonomical updates in The Scorpion Files for Scorpionidae (2008 ) Taxa Status Distribution Comments Reference Aops New genus Australia The family Urodacidae was abolished by Soleglad, Fet & Kovarik in 2005 and the subfamily Urodacinae was transfered to Scorpionidae. Volschenk & Prendini have not accepted this decision and retained the family Urodacidae (and placed the new genus and species in this family because of its close relationship to Urodacus). The taxonomy of The Scorpion Files follows Soleglad & Fet (2005), and I have here listed Aops in Scorpionidae. This is formally not correct, but is the best choice in the current situation until a new revision on higher scorpion systematics is published. Cazierius granulosus Teruel, 2013 Cazierius neibae New sp. Dominican Republic Volschenk ES, Prendini L. Aops oncodactylus, gen. et sp. nov, the frist troglobitic urodacid (Urodacidae: Scorpiones), with a re-assessment of cavernicolus, troglobitic and troglomorphic scorpions. Invertebrate Systematics. 2008;22:235. New sp. Cuba Teruel R. La subfamilia Diplocentrinae (Scorpiones: Scorpionidae) en Cuba. Sexta parte: Nueva especie de Cazierius Francke Revista Iberica de Arachnologia (23):43-8. Kovarik F, Teruel R. Three New Scorpion Species from the

2 Cazierius scaber (Pocock, 1893) Cazierius tatae Santiago- Blay, 2009 Jamaica New name: Heteronebo scaber (Pocock, 1893) Synonymization Regarded as junior synonym of C. garridoi Armas, 2005 and synonymised. Dominican Republic, Greater Antilles (Scorpiones: Buthidae, Scorpionidae). Euscorpius (187):1-27. Teruel R. Los escorpiones Diplocentrinos de Jamaica (Scorpiones: Scorpionidae: Diplocentronae). Boletin Sociedad Entomológica Aragonesa. 2009; (44): de Armas LF. Acerca de algunos alacranes (Scorpiones: Buthidae, Scorpionidae) recientemente descritos o registrados de Puerto Rico e Isla Mona, Antillas Mayores. Boletin Sociedad Entomológica Aragonesa. 2009(45):298. Cryptoiclus Teruel & Kovarik, 2012 Cryptoiclus rodriguezi Teruel & Kovarik, 2012 New sp. Puerto Rico Santiago-Blay JA. Systematics and some aspects of the biology of the scorpions (Arachnida) of the greater Puerto Rico region: A biosystematic synopsis. Entomological News Jan- Feb;120 (1): New genus Cuba Endemic to eastern Cuba. Teruel R, Kovarik F. Scorpions of Cuba: Jakub Rolcik - Clarion Production; 2012 (page ) New sp. Cuba Endemic to eastern Cuba. Teruel R, Kovarik F. Scorpions of Cuba: Jakub Rolcik - Clarion

3 Didymocentus armasi Teruel & Rodriguez, 2008 Didymocentus jaumei Armas, 1976 Didymocentus sanfelipensis Armas, 1976 Diplocentrus bicolor Contreras-Felix & Santibanez- Lopez, 2011 Diplocentrus duende Santibanez-Lopez & Gonzalez-Santillan, 2017 Production; 2012 (page ) New sp. Cuba Teruel R, Rodriguez TM. La subfamilia Diplocentrinae (Scorpiones: Scorpionidae) en Cuba. Quinta parte: el genero Didymocentrus Kraepelin Boletin Sociedad Entomológica Aragonesa. 2008(42):53-78 Restored species Restored species Species is valid and is redescribed and updated as to their respective diagnosis and geographical distribution. Species is valid and is redescribed and updated as to their respective diagnosis and geographical distribution. Teruel R, Rodriguez TM. La subfamilia Diplocentrinae (Scorpiones: Scorpionidae) en Cuba. Quinta parte: el genero Didymocentrus Kraepelin Boletin Sociedad Entomológica Aragonesa. 2008(42):53-78 Teruel R, Rodriguez TM. La subfamilia Diplocentrinae (Scorpiones: Scorpionidae) en Cuba. Quinta parte: el genero Didymocentrus Kraepelin Boletin Sociedad Entomológica Aragonesa. 2008(42):53-78 New sp. Mexico Contreras-Felix G, Santibanez- Lopez CE. Diplocentrus bicolor sp. n. (Scorpiones: Diplocentridae) from Jalisco, Mexico. Zootaxa. 2011(2992):61-8. New sp. Mexico Santibanez-Lopez CE, Gonzalez-Santillan E. A new species of Diplocentrus (Scorpiones: Diplocentridae)

4 Diplocentrus formosus Armas & Martin-Frias, 2003 Diplocentrus franckei Santobanez-Lopez, 2014 New status Reinstated as valid species. Previously synonymized with Diplocentrus tehuano Francke, with punctate pedipalp surfaces, a diagnostic character within the "mexicanus" group. C R Biol. 2017; In Press. Santibanez-Lopez CE, Francke OF, Prendini L. Systematics of the keyserlingii group of Diplocentrus Peters, 1861 (Scorpiones: Diplocentridae), with descriptions of three new species from Oaxaca, Mexico. American Museum Novitates (3777):1-47. New sp. Mexico Santibanez-Lopez CE. A new species of the genus Diplocentrus Peters, 1861 (Scorpiones, Diplocentridae) from Oaxaca, Mexico. Zookeys (412): Diplocentrus gladiator New sp. Mexico Species described in 1999, but I didn t learn about this species before October Thanks to Oscar Francke for informing me! Diplocentrus insularis Sagastume-Espinoza, Longhorn & Santiban ez- Lopez, 2015 Beutelspacher CR, Trujillo Olivera M. Una especie nueva de Diplocentrus Peters (Scorpionida: Diplocentridae) de Chiapas, Mexico. Rev Nica Ent. 1999;50:1-11. New sp. Honduras Sagastume-Espinoza KO, Longhorn SJ, Santibanez-Lopez CE. A new scorpion species of genus Diplocentrus Peters, 1861 (Scorpiones: Diplocentridae) endemic to Islas de la Bahia, Honduras. C R Biol May

5 Diplocentrus Izabal Armas & Trujillo, 2016 Diplocentrus kraepelini Santibanez-Lopez, Francke & Prendini, 2013 Diplocentrus lachua Armas. Trujillo & Agreda, 2011 Diplocentrus landelinoi Trujillo & Armas, 2012 Diplocentrus longimanus Santibanez-Lopez, Francke & Athanasiadis, New sp. Guatemala De Armas LF, Trujillo RE. A new species of Diplocentrus (Scorpionidae: Diplocentrinae) from western Izabal, Guatemala. Euscorpius. 2016(225):1-8. New sp. Mexico Santibanez-Lopez CE, Francke OF, Prendini L. Systematics of the keyserlingii group of Diplocentrus Peters, 1861 (Scorpiones: Diplocentridae), with descriptions of three new species from Oaxaca, Mexico. American Museum Novitates (3777):1-47. New sp. Guatemala de Armas LF, Trujillo RE, Agreda EO. Nueva especie de Diplocentrus Peters, 1851 (Scorpionidae: Diplocentrinae) del noroeste de alta Verapaz, Guatemala. Boletin de la SEA. 2011(49): New sp. Guatemala Trujillo RE, de Armas LF. Dos especie nuevas de Diplocentrus Peters, 1861 (Scorpionidae: Diplocentrinae) de Guatemala. Revista Iberica de Arachnologia (21): New sp. Mexico Santibanez-Lopez CE, Francke OF, Athanasiadis MC. The genus Diplocentrus Peters (Scorpiones: Diplocentridae) in

6 Diplocentrus majahuensis New sp. Mexico Species described in 2003, but I didn t learn about this species before October Thanks to Oscar Francke for informing me! Diplocentrus motagua Armas & Trujillo, 2009 Diplocentrus oxlajujbaktun Trujillo & Armas, 2012 Diplocentrus poncei Francke & Quijano-Ravell, 2009 Diplocentrus popti Trujillo & Armas, 2016 Morelos, Mexico. Revista Iberica de Arachnologia. 2011;19:3-13. Baldazo Monsivaiz JG. Diplocentrus majahuensis, nueva especie de alcran del estado de Guerrero, Mexico (Scorpiones, Diplocentridae). Entomologia Mexicana. 2003;2:73-7. New sp. Guatemala de Armas LF, Trujillo RE. Nueva especie de Diplocentrus Peters, 1861 (Scorpiones: Scorpionidae) de Guatemala. Boletin Sociedad Entomológica Aragonesa. 2009(45): New sp. Guatemala Trujillo RE, de Armas LF. Dos especie nuevas de Diplocentrus Peters, 1861 (Scorpionidae: Diplocentrinae) de Guatemala. Revista Iberica de Arachnologia (21): See Kolotl below. New sp. Mexico Authors using family Diplocentridae Francke OF, Quijano-Ravell AF. Una especie nueva Diplocentrus (Scorpiones: Diplocentridae) del estado de Michoacan, Mexico. Revista Mexicana de Biodiversidad 2009;80: New sp. Guatemala Trujillo RE, de Armas LF. Nueva especie de Diplocentrus

7 Diplocentrus sagittipalpus Santibanez-Lopez, Francke & Prendini, 2013 Diplocentrus sissomi Santibanez-Lopez, Francke & Prendini, 2013 peters, 1861 (Scorpiones: Scorpionidae: Diplocentrinae) del occidente de Guatemala. Revista Iberica de Arachnologia. 2016(28): New sp. Mexico Santibanez-Lopez CE, Francke OF, Prendini L. Systematics of the keyserlingii group of Diplocentrus Peters, 1861 (Scorpiones: Diplocentridae), with descriptions of three new species from Oaxaca, Mexico. American Museum Novitates (3777):1-47. New sp. Mexico Santibanez-Lopez CE, Francke OF, Prendini L. Systematics of the keyserlingii group of Diplocentrus Peters, 1861 (Scorpiones: Diplocentridae), with descriptions of three new species from Oaxaca, Mexico. American Museum Novitates (3777):1-47. Diplocentrus tenango New sp. Mexico Santibanez-Lopez CE, Francke OF. A new species of Diplocentrus (Arachnida: Scorpiones) from Oaxaca, Mexico. Zootaxa. 2008(1742):53-60 Heterometrus atrascorpius Mirza, Joshi, Desouza & Sanap, 2012 New sp. India Mirza ZA, Joshi D, Desouza G, Sanap RV. Description of a new species of scorpion of the genus

8 Heterometrus liangi Zhu & Yang, 2007 Heterometrus minotaurus Pliskova, Kovarik, Kosulic & Stahlavsky, 2016 Heterometrus serratus (Pocock, 1900) New sp. Vietnam Haven t seen this paper (in Chinese). Info based on bibliographical data only. Heterometrus Ehrenberg, 1828 (Scorpiones: Scorpionidae) from the Western Ghats, India. Indian Journal of Arachnology. 2012;1(2):1-8. Zhu MS, Yang X-F. Two species of the genus Heterometrus Ehrenberg, 1828 (Scorpionidae) from South Vietnam sold in pet shops in China. Acta Arachnologica Sinica. 2007;16 (2): New sp. Thailand Pliskova J, Kovarik F, Kosulic O, Stahlavsky F. Description of a new species of Heterometrus Ehrenberg, 1828 (Scorpiones: Scorpionidae) from Thailand with remarks about the utilization of cytogenetic data in taxonomy of the genus. Annales Zoologici. 2016;66(3): New status Restored from synonymy with H. indus (Geer, 1778). Kovarik F, Lowe G, Ranawana KB, Hoferek D, Jayarathne VAS, Pliskova J, et al. Scorpions of Sri Lanka (Arachnida, Scorpiones: Buthidae, Chaerilidae, Scorpionidae) with of the Genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972, stat. n. Euscorpius (220):1-133.

9 Heterometrus telanganaensis Javed, Mirza, Tampal & Lourenco, 2010 Heteronebo barahonae Teruel, de Armas & Kovarik, 2015 Heteronebo franckei Stockwell, 1985 Heteronebo jamaicae jamaicae Francke, 1978 New sp. India Maqsood Javed SM, Mirza ZA, Tampal F, Lourenco WR. A new species of the genus Heterometrus Ehrenberg, 1828 (Scorpiones: Scorpionidae) from India with notes on its natural history. Boletin de la SEA. 2010(47): New sp. Synonymization New status Hispaniola, Dominican Republic Synonymized with Heteronebo scaber (Pocock, 1893) Revalidated as species. Teruel R, de Armas LF, Kovarik F. Two new species of scorpions (Scorpiones: Buthidae, Scorpionidae) from Dominican Republic, Greater Antilles. Revista Iberica de Arachnologia (27): Teruel R. Los escorpiones Diplocentrinos de Jamaica (Scorpiones: Scorpionidae: Diplocentronae). Boletin Sociedad Entomológica Aragonesa. 2009; (44): Santibanez-Lopez CE, Francke OF, Prendini L. Phylogeny of the North American scorpion genus Diplocentrus Peters, 1861 (Scorpiones: Diplocentridae) based on morphology, nuclear and mitochondrial DNA. Arthropod Systematics & Phylogeny. 2014;72(3): Synonymization Synonymized with Heteronebo Teruel R. Los escorpiones

10 Heteronebo jamaicae occidentalis Francke, 1978 New status scaber (Pocock, 1893) Revalidated as species. Diplocentrinos de Jamaica (Scorpiones: Scorpionidae: Diplocentronae). Boletin Sociedad Entomológica Aragonesa. 2009; (44): Santibanez-Lopez CE, Francke OF, Prendini L. Phylogeny of the North American scorpion genus Diplocentrus Peters, 1861 (Scorpiones: Diplocentridae) based on morphology, nuclear and mitochondrial DNA. Arthropod Systematics & Phylogeny. 2014;72(3): Heteronebo jamaicae portlandensis Francke, 1978 Synonymization New status Synonymized with Heteronebo scaber (Pocock, 1893) Revalidated as species. Teruel R. Los escorpiones Diplocentrinos de Jamaica (Scorpiones: Scorpionidae: Diplocentronae). Boletin Sociedad Entomológica Aragonesa. 2009; (44): Santibanez-Lopez CE, Francke OF, Prendini L. Phylogeny of the North American scorpion genus Diplocentrus Peters, 1861 (Scorpiones: Diplocentridae) based on morphology, nuclear and mitochondrial DNA. Arthropod Systematics & Phylogeny. 2014;72(3): Synonymization Synonymized with Heteronebo Teruel R. Los escorpiones

11 Heteronebo morenoi (Armas, 1973) Heteronebo scaber (Pocock, 1893) Kolotl Santibanez-Lopez, Francke & Prendini, 2014 Kolotl magnus (Beutelspacher & López-Forment, 1991) scaber (Pocock, 1893) New status Cuba Raised from subspecies status. Previously Heteronebo bermudezi morenoi (Armas, 1973). Jamaica Previous name: Cazierius scaber (Pocock, 1893) + senior synonym of Heteronebo franckei Stockwell, 1985, H. jamaicae jamaicae Francke, 1978, H. jamaicae occidentalis Francke, 1978 & H. jamaicae portlandensis Francke, 1978 Diplocentrinos de Jamaica (Scorpiones: Scorpionidae: Diplocentronae). Boletin Sociedad Entomológica Aragonesa. 2009; (44): Teruel R, Rodriguez-Cabrera TM. The Missing Piece of the Puzzle Solved: Heteronebo Pocock, 1899 (Scorpiones: Scorpionidae) Occurs at Isla de Pinos, Cuba. Euscorpius. 2017(240):1-4. Teruel R. Los escorpiones Diplocentrinos de Jamaica (Scorpiones: Scorpionidae: Diplocentronae). Boletin Sociedad Entomológica Aragonesa. 2009; (44): New genus Western Mexico Santibanez-Lopez CE, Francke OF, Prendini L. Kolotl, n. gen. (Scorpiones: Diplocentridae), a New Scorpion Genus from Mexico. American Museum Novitates (3815):1-28. Previous name: Diplocentrus magnus Beutelspacher & López- Forment, Santibanez-Lopez CE, Francke OF, Prendini L. Kolotl, n. gen. (Scorpiones: Diplocentridae), a New Scorpion Genus from Mexico. American Museum Novitates (3815):1-28.

12 Kolotl poncei (Francke & Quijano-Ravell, 2009) Previous name: Diplocentrus poncei Francke & Quijano-Ravell, Pandiborellius Rossi, 2015 New status Previously Pandinurus (Pandiborellius) Rossi, 2015a. Pandiborellius arabicus (Kraepelin, 1894) Previous Pandinurus arabicus (Kraepelin, 1894) Santibanez-Lopez CE, Francke OF, Prendini L. Kolotl, n. gen. (Scorpiones: Diplocentridae), a New Scorpion Genus from Mexico. American Museum Novitates (3815):1-28. Kovarik F, Lowe G, Soleglad Kovarik F, Lowe G, Soleglad Pandinurus arabicus (Kraepelin, 1894) Previous Pandinus arabicus (Kraepelin, 1894).

13 Pandiborellius awashensis (Kovarik, 2012) Previous Pandinurus awashensis (Kovarik, 2012) Kovarik F, Lowe G, Soleglad Pandinurus awashensis (Kovarik, 2012) Previous Pandinus awashensis (Kovarik, 2012). New sp. Ethiopia Kovarik F. Review of the subgenus Pandinurus Fet, 1997 with descriptions of three new species (Scorpiones, Scorpionidae, Pandinus). Euscorpius. 2012(141):1-22.

14 Pandiborellius igdu Kovarik, Lowe, Soleglad & Pliskova, 2017 Pandiborellius insularis Kovarik, Lowe, Soleglad & Pliskova, 2017 Pandiborellius lanzai (Rossi, 2015) New sp. Ethiopia Kovarik F, Lowe G, Soleglad New sp. Eritrea Kovarik F, Lowe G, Soleglad Previously Pandinurus lanzai Rossi, Kovarik F, Lowe G, Soleglad

15 Pandinurus lanzai Rossi, 2015 Pandiborellius magrettii (Borelli, 1901) New sp. Somalia Previous Pandinurus magrettii (Borelli, 1901) Rossi A. Due ulteriori nuove specie del sottogenere Pandiborellius Rossi, 2015 dal Corno d'africa (Scorpiones, Scorpionidae). Arachnida - Rivista Aracnologica Italiana. 2015;1(2):2-36. Kovarik F, Lowe G, Soleglad Pandinurus magrettii (Borelli, 1901) Pandiborellius meidensis (Karsch, 1879) Previous Pandinus magrettii (Borelli, 1901). Kovarik F, Lowe G, Soleglad

16 Pandinurus meidensis (Karsch, 1879) Pandiborellius nistriae (Rossi, 2014) New comb Previous Pandinus meidensis (Karsch, 1879). Previous Pandinurus nistriae (Rossi, 2014) Kovarik F, Lowe G, Soleglad Pandinurus nistriae (Rossi, 2014) Previous Pandinus nistriae (Rossi, 2014).

17 Pandiborellius percivali (Pocock, 1902) New sp. Djibouti Previous Pandinurus percivali (Pocock, 1902) Rossi A. Notes on the distribution of Pandinus (Pandinus) Thorell, 1876 and Pandinus (Pandinurus) Fet, 1997 with the descriptions of two new species from Central African Republic. Onychium. 2014;10 (2013): Kovarik F, Lowe G, Soleglad Pandinurus percivali (Pocock, 1902) Previous Pandinus percivali (Pocock, 1902).

18 Pandiborellius somalilandus (Kovarik, 2013) Previous Kovarik F, Lowe G, Soleglad Pandinurus somalilandus (Kovarik, 2013) Previous Pandinus somalilandus (Kovarik, 2012). New sp. Somaliland Pandinoides Fet, 1997 New status Raised to genus from subgenus status within genus Pandinus Thorell, 1876 Kovarik F. Review of the subgenus Pandinurus Fet, 1997 with descriptions of three new species (Scorpiones, Scorpionidae, Pandinus). Euscorpius. 2012(141):1-22.

19 Pandinoides cavimanus (Pocock, 1888) Pandinoides duffmackayi Prendini, 2016 Pandinoides militaris Pocock, 1900 New sp. New status/ Kenya and Tanzania Previous Pandinus cavimanus (Pocock, 1888). Revalidated and redescribed. Prendini L. Redefinition and systematic revision of the East African scorpion genus Pandinoides (Scorpiones: Scorpionidae) with critique of the taxonomy of Pandinus, sensu lato. Bulletin of the American Museum of Natural History. 2016(407):1-67. Prendini L. Redefinition and systematic revision of the East African scorpion genus Pandinoides (Scorpiones: Scorpionidae) with critique of the taxonomy of Pandinus, sensu lato. Bulletin of the American Museum of Natural History. 2016(407):1-67.

20 Pandinus militaris Pocock, 1900 Synonymization This taxa was judged as a junior synonym of Pandinoides cavimanus Pandinops Birula, 1913 New status Raised to genus from subgenus status within genus Pandinus Thorell, 1876 Pandinops bellicosus (L. Koch, 1875) Pandinus boschisi (Caporiacco, 1937) Previous Pandinus bellicosus (L. Koch, 1875). Previously placed in Pandinus with a nomen dubium status. Kovarik F. A checklist of scorpions (Arachnida) in the collection of the Forschungsinstitut und Naturmuseum Senckenberg, Frankfurt am Main, Germany. Serket April;8 (1):1-23. Rossi A. Una revisione preliminare del genere Pandinus Thorell, 1876 con la descrizione di un nuovo sottogenere e due nuove specie dall'etiopia (Scorpiones, Scorpionisae). Arachnida - Rivista Aracnologica Italiana. 2015;1(3):37-52.

21 Pandinops colei (Pocock, 1896) Pandinops eritreaensis (Kovarik, 2003) Pandinops friedrichi Kovarik, 2016 Pandinops hawkeri (Pocock, 1900) Previous Pandinus colei (Pocock, 1896). Previous Pandinus eriatreaensis Kovarik, New sp. Somalia Kovarik F. Scorpions of the Scorpiones). Part VIII. Pandinops Birula, 1913 (Scorpionidae), With Description of Two New Species. Euscorpius. 2016(229):1-20. Synonymization Synonymized with Pandinops pugilator (Pocock, 1900). Kovarik F, Lowe G, Elmi HSA. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XIII. Review of Pandinops hawkeri, P. peeli, P.platycheles, and P. pugilator (Scorpionidae). Euscorpius. 2017(254):1-20.

22 Pandinops peeli (Pocock, 1900) Synonymization Previous Pandinus hawkeri (Pocock, 1900). Synonymized with Pandinops pugilator (Pocock, 1900). Kovarik F, Lowe G, Elmi HSA. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XIII. Review of Pandinops hawkeri, P. peeli, P.platycheles, and P. pugilator (Scorpionidae). Euscorpius. 2017(254):1-20. Pandinops platycheles (Werner, 1916) Previous Pandinus peeli (Pocock, 1900). Previous Pandinurus platycheles (Werner, 1916) Kovarik F, Lowe G, Elmi HSA. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XIII. Review of Pandinops hawkeri, P. peeli, P.platycheles, and P. pugilator (Scorpionidae). Euscorpius. 2017(254):1-20

23 Previous Pandinoides platycheles (Werner, 1916) Kovarik F, Lowe G, Pliskova J, Stahlavsky F. Scorpions of the Scorpiones). Part VII. Parabuthus Pocock, 1890 (Buthidae), with Description of P. hamar sp. n. and P. kajibu sp. n. from Ethiopia. Euscorpius. 2016(228):1-58. Pandinops pococki (Kovarik, 2000) Pandinops pugilator (Pocock, 1900) New status Previous Pandinus platycheles (Werner, 1916). Previous Pandinus pococki Kovarik, 2000 Previously synonymized with Pandinops hawkeri (Pocock, 1900). Kovarik F, Lowe G, Elmi HSA. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XIII. Review of Pandinops hawkeri, P. peeli, P.platycheles, and P. pugilator (Scorpionidae).

24 Pandinops turieli Kovarik, 2016 Pandinopsis Vachon, 1974 New status Raised to genus from subgenus status within genus Pandinus Thorell, 1876 Pandinurus Fet, 1997 New status Raised to genus from subgenus status within genus Pandinus Thorell, 1876 Pandinurus afar Kovarik, Lowe, Soleglad & Pliskova, 2017 Euscorpius. 2017(254):1-20. New sp. Ethiopia and Kovarik F. Scorpions of the Kenya Scorpiones). Part VIII. Pandinops Birula, 1913 (Scorpionidae), With Description of Two New Species. Euscorpius. 2016(229):1-20. New sp. Ethiopia Kovarik F, Lowe G, Soleglad

25 Pandinurus bartolozzii Rossi, 2015 Synonymization Synonymized with Pandinus viatoris (Pocock, 1890) Prendini L. Redefinition and systematic revision of the East African scorpion genus Pandinoides (Scorpiones: Scorpionidae) with critique of the taxonomy of Pandinus, sensu lato. Bulletin of the American Museum of Natural History. 2016(407):1-67. Pandinurus cianferonii Rossi, 2015 New sp. Synonymization Democratic Republic of the Congo Synonymized with P. pallidus (Kraepelin, 1894) Kovarik F, Lowe G, Soleglad

26 New sp. Somalia Rossi A. Ulteriori commenti sulla tassonomia dei generi Pandinus e Pandinurus, con la definizione di un nuovo sottogenere ed una nuova specie di Pandinurus dalla Somalia. Arachnida - Rivista Aracnologica Italiana. 2015;1(4): Pandinurus citernii (Borelli, New status Restored from synonymy with P. Kovarik F, Lowe G, Soleglad 1919) phillipsii (Pocock, 1896). Pandinurus exitialis (Pocock, Previous Pandinus exitialis (Pocock, 1888) 1888). Pandinurus flagellicauda Synonymization Synonymized with Pandinus Prendini L. Redefinition and

27 Rossi, 2015 viatoris (Pocock, 1890) systematic revision of the East African scorpion genus Pandinoides (Scorpiones: Scorpionidae) with critique of the taxonomy of Pandinus, sensu lato. Bulletin of the American Museum of Natural History. 2016(407):1-67. Pandinurus gregoryi (Pocock, 1896) New sp. Democratic Republic of the Congo Previous Pandinus gregoryi (Pocock, 1888). New status Considered a synonym of Pandinus exitialis (Pocock, 1888) by Kovarik, 2003, but restored species status in Kovarik s (2009) Illustrated catalog of scorpions (page 55). Kovarik F. Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with illustrated keys to Heterometrus and Pandinus

28 Pandinurus hangarale Kovarik, Lowe, Mazuch, Awale, Stundlowa & Stahlavsky, 2017 Pandinurus intermedius (Borelli, 1919) species. Prague: Clarion Production; New sp. Somaliland Kovarik F, Lowe G, Mazuch T, Awale AI, Stundlova J, Stahlavsky F. Scorpions of the Scorpiones). Part XII. Pandinurus hangarale sp. n. (Scorpionidae) from Somaliland and a Review of Type Locality and True Distribution of Pandinurus smithi (Pocock, 1897). Euscorpius. 2017(253):1-18. New status Restored from synonymy with P. Kovarik F, Lowe G, Soleglad phillipsii (Pocock, 1896). Pandinurus janae Rossi, 2015 Synonymization Synonymized with Pandinus percivali (Pocock, 1902). Prendini L. Redefinition and systematic revision of the East African scorpion genus Pandinoides (Scorpiones: Scorpionidae) with critique of the taxonomy of Pandinus, sensu lato. Bulletin of the American

29 Museum of Natural History. 2016(407):1-67. Pandinurus kmoniceki Kovarik, Lowe, Mazuxh, Pliskova & Stahlavsky, 2017 Pandinurus lorenzoi Rossi, 2015 New sp. Yemen New sp. Somaliland Kovarik F, Lowe G, Mazuch T, Pliskova J, Stahlavsky F. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XI. Pandinurus kmoniceki sp. n. (Scorpionidae) from Somaliland. Synonymization Synonymized with Pandinus viatoris (Pocock, 1890) Euscorpius. 2017(243):1-14. Prendini L. Redefinition and systematic revision of the East African scorpion genus Pandinoides (Scorpiones: Scorpionidae) with critique of the taxonomy of Pandinus, sensu lato. Bulletin of the American Museum of Natural History. 2016(407):1-67. New sp. Tanzania

30 Pandinurus mazuchi (Kovarik, 2011) Previously Pandinus mazuchi Kovarik, Kovarik F, Lowe G, Soleglad Pandinus mazuchi Kovarik, 2011 Pandinurus oromo Kovarik, Lowe, Soleglad & Pliskova, 2017 New sp. Ethiopia Kovarik F. A review of the subgenus Pandinus Thorell, 1876 with descriptions of two new species from Uganda and Ethiopia (Scorpiones: Scorpionidae). Euscorpius. 2011(129):1-18. New sp. Ethiopia Kovarik F, Lowe G, Soleglad

31 Pandinurus pallidus (Kraepelin, 1894) Pandinurus pantinii Rossi, 2015 Synonymization Previous Pandinus kraepelin (Pocock, 1894). Synonymized with Pandinus viatoris (Pocock, 1890) Prendini L. Redefinition and systematic revision of the East African scorpion genus Pandinoides (Scorpiones: Scorpionidae) with critique of the taxonomy of Pandinus, sensu lato. Bulletin of the American Museum of Natural History. 2016(407):1-67. Pandinurus phillipsii (Pocock, 1896) New sp. Malawi New status Restored from synonymy. Kovarik F, Lowe G, Soleglad

32 Pandinurus prendinii Rossi, 2015 Synonymization Synonymized with Pandinus sudanicus (Hirst, 1911). Prendini L. Redefinition and systematic revision of the East African scorpion genus Pandinoides (Scorpiones: Scorpionidae) with critique of the taxonomy of Pandinus, sensu lato. Bulletin of the American Museum of Natural History. 2016(407):1-67. Pandinurus pygmaeus Rossi, 2015 New sp. Synonymization South Africa Synonymized with Pandinus viatoris (Pocock, 1890) Prendini L. Redefinition and systematic revision of the East African scorpion genus Pandinoides (Scorpiones: Scorpionidae) with critique of the taxonomy of Pandinus, sensu lato. Bulletin of the American

33 Museum of Natural History. 2016(407):1-67. Pandinurus sabbadinii Rossi, 2015 New sp. Synonymization Democratic Republic of Congo Synonymized with Pandiborellius magrettii (Borelli, 1901) Kovarik F, Lowe G, Soleglad Pandinurus smithi (Pocock, 1897) New sp. Eritrea Previously Pandinus smithi (Pocock, 1987). Rossi A. Due ulteriori nuove specie del sottogenere Pandiborellius Rossi, 2015 dal Corno d'africa (Scorpiones, Scorpionidae). Arachnida - Rivista Aracnologica Italiana. 2015;1(2):2-36. Kovarik F, Lowe G, Soleglad

34 Pandinurus sudanicus (Hirst, 1911) Previous Pandinus sudanicus (Hirst, 1911). Pandinurus trailini (Kovarik, 2013) New status Sudan Previous status: Pandinus exitialis sudanicus Hirst, 1911 (in part) Pandinus magretti Birula, 1928 (in part). Previously Pandinus trailini Kovarik, Kovarik F. Review of the subgenus Pandinurus Fet, 1997 with descriptions of three new species (Scorpiones, Scorpionidae, Pandinus). Euscorpius. 2012(141):1-22. Kovarik F, Lowe G, Soleglad

35 Pandinus trailini Kovarik, 2013 Pandinurus vachoni (Rossi, 2014) New sp. Synonymization Ethiopia Synonymized with P. sudanicus (Hirst, 1911). Kovarik F. Pandinus (Pandinus) trailini sp. n. from Ethiopia (Scorpiones: Scorpionidae), with data on localities and life strategy. Euscorpius (163):1-14. Kovarik F, Lowe G, Soleglad Previous Pandinus vachoni (Rossi, 2014) New sp. Chad Rossi A. The fragmented peri-

36 Pandinus bottegoi Rossi, 2015 Synonymization Synonymized with P. platycheles (Werner, 1916). Saharan distribution of the subgenus Pandinurus Fet, 1997 with the description of a new species from Chad (Scorpiones, Scorpionidae, Pandinus). Serket. 2014;14(1):6-14. Kovarik F, Lowe G, Soleglad Pandinus camerounensis Lourenco, 2014 New sp. Synonymization Ethiopia Synonymized with Pandinus imperator (C. L. Koch, 1841) Rossi A. Una revisione preliminare del genere Pandinus Thorell, 1876 con la descrizione di un nuovo sottogenere e due nuove specie dall'etiopia (Scorpiones, Scorpionisae). Arachnida - Rivista Aracnologica Italiana. 2015;1(3): Prendini L. Redefinition and systematic revision of the East African scorpion genus Pandinoides (Scorpiones: Scorpionidae) with critique of

37 the taxonomy of Pandinus, sensu lato. Bulletin of the American Museum of Natural History. 2016(407):1-67. Pandinus dictator (Pocock, 1888) New sp. Cameroon Previous Pandinopsis dictator (Pocock, 1888). Pandinopsis is subgenus. Lourenco WR. Further considerations on the identity and distribution of Pandinus imperator (C. L. Koch, 1841) and description of a new species from Cameroon (Scorpiones: Scorpionidae). entomologische Mitteilungen aus dem Zoologischen Museum Hamburg. 2014;17(192): Kovarik F, Lowe G, Soleglad Pandinopsis dictator (Pocock, 1888) Previous Pandinus dictator (Pocock, 1888).

38 Pandinus lowei Kovarik, 2012 Previous Pandinus lowei (Kovarik, 2012). Kovarik F, Lowe G, Soleglad Pandinurus lowei (Kovarik, 2012) Previously Pandinurus lowei (Kovarik, 2012) New sp. Democratic Republic of Congo Kovarik F. Review of the subgenus Pandinurus Fet, 1997 with descriptions of three new species (Scorpiones, Scorpionidae, Pandinus). Euscorpius. 2012(141):1-22. Pandinus platycheles See below for previous status. Prendini L. Redefinition and

39 (Werner, 1916) systematic revision of the East African scorpion genus Pandinoides (Scorpiones: Scorpionidae) with critique of the taxonomy of Pandinus, sensu lato. Bulletin of the American Museum of Natural History. 2016(407):1-67. Pandinus platycheles (Werner, 1916) Previous Pandinoides platycheles (Werner, 1916) Kovarik F, Lowe G, Pliskova J, Stahlavsky F. Scorpions of the Scorpiones). Part VII. Parabuthus Pocock, 1890 (Buthidae), with Description of P. hamar sp. n. and P. kajibu sp. n. from Ethiopia. Euscorpius. 2016(228):1-58. Pandinus platycheles (Werner, 1916) Pandinus riccardoi Rossi, 2015 Synonymization Previous Pandinus platycheles (Werner, 1916). Synonymized with P. platycheles (Werner, 1916). Kovarik F, Lowe G, Soleglad

40 Pandinus roeseli (Simon, 1872) New sp. Synonymization Ethiopia Synonymized with Pandinus imperator (C. L. Koch, 1841) Rossi A. Una revisione preliminare del genere Pandinus Thorell, 1876 con la descrizione di un nuovo sottogenere e due nuove specie dall'etiopia (Scorpiones, Scorpionisae). Arachnida - Rivista Aracnologica Italiana. 2015;1(3): Prendini L. Redefinition and systematic revision of the East African scorpion genus Pandinoides (Scorpiones: Scorpionidae) with critique of the taxonomy of Pandinus, sensu lato. Bulletin of the American Museum of Natural History. 2016(407):1-67. New status Guinea Restored from synonymization with Pandinus imperator (Koch, 1841) Lourenco WR. Further considerations on the identity and distribution of Pandinus imperator (C. L. Koch, 1841) and description of a new species from Cameroon (Scorpiones:

41 Pandinus ugandaensis Kovarik, 2011 Pandinus ulderigoi Rossi, 2014 Pandinus viatoris (Pocock, 1890) Scorpionidae). entomologische Mitteilungen aus dem Zoologischen Museum Hamburg. 2014;17(192): New sp. Uganda Kovarik F. A review of the subgenus Pandinus Thorell, 1876 with descriptions of two new species from Uganda and Ethiopia (Scorpiones: Scorpionidae). Euscorpius. New sp. Central African Republic Previous Pandinus viatoris (Pocock, 1890). 2011(129):1-18. Rossi A. Notes on the distribution of Pandinus (Pandinus) Thorell, 1876 and Pandinus (Pandinurus) Fet, 1997 with the descriptions of two new species from Central African Republic. Onychium. 2014;10 (2013): Kovarik F, Lowe G, Soleglad Pandinurus viatoris (Pocock, Previous Pandinurus viatoris

42 1890) (Pocock, 1890). Oiclus nanus Teruel & Chazal, 2010 New Sp. Guadeloupe, Lesser Antilles Teruel R, Chazal L. A new species of the genus Oiclus Simon, 1880 (Scorpiones: Scorpionidae: Diplocentrinae) from Guadeloupe, Lesser Antilles. Euscorpius (92):1-9. Oiclus questeli Teruel, 2008 New Sp. Lesser Antilles Teruel R. A new species of Oiclus Simon 1880 (Scorpiones: Scorpionidae: Diplocentrinae) from Saint-Barthelemy, Lesser Antilles. Boletin Sociedad Entomológica Aragonesa. 2008(43):95-9. Scorpio birulai Fet, 1997 New status Morocco Previous subspecies status: Scorpio maurus subtypicus Birula, 1910 Scorpio ennedi Lourenco, Duhem & Cloudsley- Thompson, 2012 Lourenco WR. Reanalysis of the genus Scorpio Linnaeus 1758 in sub-saharan Africa and description of one new species from Cameroon (Scorpiones, Scorpionidae). Entomol Mitt Zool Mus Hamburg. 2009;15(181): New sp. Chad Lourenco WR, Duhem B, Cloudsley Thompson JL. Scorpions from Ennedi, Kapka

43 Scorpio fuliginosus Pallary, 1928 Scorpio fuscus (Ehrenberg, 1829) New status Morocco Previous status: Heterometrus fuliginosus Pallary, 1928 New status Israel, Palestina Previous subspecies status: Scorpio maurus fuscus Ehrenberg, 1829 Scorpio hesperus Birula, 1910 New status Morocco Previous subspecies status: Scorpio maurus hesperus Birula, 1910 and Tibesti, the mountains of Chad, with descriptions of nine new species (Scorpiones: Buthidae, Scorpionidae). Arthropoda Selecta. 2012;21(4): Lourenco WR. Reanalysis of the genus Scorpio Linnaeus 1758 in sub-saharan Africa and description of one new species from Cameroon (Scorpiones, Scorpionidae). Entomol Mitt Zool Mus Hamburg. 2009;15(181): Talal S, Tesler I, Sivan J, Ben- Shlomo R, Muhammad Tahir H, Prendini L, et al. Scorpion speciation in the Holy Land: Multilocus phylogeography corroborates diagnostic differences in morphology and burrowing behavior among Scorpio subspecies and justifies recognition as phylogenetic, ecological and biological species. Mol Phylogenet Evol May 15. Lourenco WR. Reanalysis of the genus Scorpio Linnaeus 1758 in sub-saharan Africa and description of one new species from Cameroon (Scorpiones,

44 Scorpio kruglovi (Ehrenberg, 1829) Scorpio mogadorensis Birula, 1910 Scorpio niger Lourenco & Cloudsley-Thompson, 2012 New status Israel, Jordan, Syria Previous subspecies status: Scorpio maurus kruglovi Ehrenberg, 1829 New status Morocco Previous subspecies status: Scorpio maurus mogadorensis Birula, 1910 Scorpionidae). Entomol Mitt Zool Mus Hamburg. 2009;15(181): Talal S, Tesler I, Sivan J, Ben- Shlomo R, Muhammad Tahir H, Prendini L, et al. Scorpion speciation in the Holy Land: Multilocus phylogeography corroborates diagnostic differences in morphology and burrowing behavior among Scorpio subspecies and justifies recognition as phylogenetic, ecological and biological species. Mol Phylogenet Evol May 15. Lourenco WR. Reanalysis of the genus Scorpio Linnaeus 1758 in sub-saharan Africa and description of one new species from Cameroon (Scorpiones, Scorpionidae). Entomol Mitt Zool Mus Hamburg. 2009;15(181): New sp. Niger Lourenco WR, Cloudsley Thompson JL. About the enigmatic presence of the genus Scorpio Linnaeus, 1758 in Congo with the description of a new species from Niger (Scorpiones, Scorpionidae). Serket. 2012;13(1/2):1-7.

45 Scorpio occidentalis Werner, 1936 Scorpio palmatus (Ehrenberg, 1829) Scorpio propinquus (Simon, 1872) New status New status Mali, Mauritania & Senegal Israel, Palestina, Egypt Previous subspecies status: Scorpio maurus occidentalis Werner, 1936 Previous subspecies status: Scorpio maurus palmatus Ehrenberg, 1829 New status Israel, Syria Previous subspecies status: Scorpio maurus propinquus (Simon, 1872) Lourenco WR. Reanalysis of the genus Scorpio Linnaeus 1758 in sub-saharan Africa and description of one new species from Cameroon (Scorpiones, Scorpionidae). Entomol Mitt Zool Mus Hamburg. 2009;15(181): Talal S, Tesler I, Sivan J, Ben- Shlomo R, Muhammad Tahir H, Prendini L, et al. Scorpion speciation in the Holy Land: Multilocus phylogeography corroborates diagnostic differences in morphology and burrowing behavior among Scorpio subspecies and justifies recognition as phylogenetic, ecological and biological species. Mol Phylogenet Evol May 15. Talal S, Tesler I, Sivan J, Ben- Shlomo R, Muhammad Tahir H, Prendini L, et al. Scorpion speciation in the Holy Land: Multilocus phylogeography corroborates diagnostic differences in morphology and burrowing behavior among Scorpio subspecies and justifies recognition as phylogenetic, ecological and biological

46 Scorpio punicus Fet, 2000 New status Algeria, Libya, Morocco & Tunisia Scorpio savanicola Lourenco, 2009 Scorpio sudanensis Lourenco & Cludsley-Thompson, 2009 Scorpio tassili Lourenco & Rossi, 2016 species. Mol Phylogenet Evol May 15. Previous subspecies status: Scorpio Lourenco WR. Reanalysis of the maurus tunetanus Birula, 1910 genus Scorpio Linnaeus 1758 in sub-saharan Africa and description of one new species from Cameroon (Scorpiones, Scorpionidae). Entomol Mitt Zool Mus Hamburg. 2009;15(181): New sp. Cameroon Lourenco WR. Reanalysis of the genus Scorpio Linnaeus 1758 in sub-saharan Africa and description of one new species from Cameroon (Scorpiones, Scorpionidae). Entomol Mitt Zool Mus Hamburg. 2009;15(181): New sp. Sudan Lourenco WR, Cloudsley Thompson JL. A new species of the genus Scorpio Linnaeus 1758 from Sudan (Scorpiones, Scorpionidae). Boletin Sociedad Entomológica Aragonesa (45): New sp. Algeria Lourenco WR, Rossi A. Confirmation of a new species of Scorpio Linnaeus, 1758 in the Tassili N Ajjer Mountains, South Algeria (Scorpiones: Scorpionidae). Onychium. 2016;12:11-8.

47 Scorpio weidholzi Werner, 1929 Urodacus butleri Volschenk, Harvey & Prendini, 2012 New status Morocco Previous subspecies status: Scorpio maurus weidholzi Werner, 1929 Lourenco WR. Reanalysis of the genus Scorpio Linnaeus 1758 in sub-saharan Africa and description of one new species from Cameroon (Scorpiones, Scorpionidae). Entomol Mitt Zool Mus Hamburg. 2009;15(181): New sp. Australia Volschenk ES, Harvey MS, Prendini L. A new species of Urodacus (Scorpiones: Urodacidae) from Western Australia. American Museum Novitates. 2012(3748):1-18.