Master triathletes have not reached limits in their Ironman triathlon performance

Size: px
Start display at page:

Download "Master triathletes have not reached limits in their Ironman triathlon performance"

Transcription

1 Scnd J Med Sci Sports 2012: : doi: /j x 2012 John Wiley & Sons A/S Mster trithletes hve not reched limits in their Ironmn trithlon performnce M. Stiefel 1, B. Knechtle 1,2, R. Lepers 3 1 Institute of Generl Prctice nd for Helth Services Reserch, University of Zurich, Zurich, Switzerlnd, 2 Gesundheitszentrum St. Gllen, St. Gllen, Switzerlnd, 3 INSERM U1093, Fculty of Sport Sciences, University of Burgundy, Dijon, Frnce Corresponding uthor: Bet Knechtle, MD, Fchrzt FMH für Allgemeinmedizin, Gesundheitszentrum St. Gllen, Vdinstrsse 26, 9001 St. Gllen, Switzerlnd. Tel: , Fx: , E-mil: et.knechtle@hispeed.ch Accepted for puliction 2 April 2012 The purpose of this study ws to nlyze the prticiption nd performnce trends of mle trithletes in the Ironmn Switzerlnd from 1995 to Prticiption trends of ll finishers ged etween 18 nd 64 yers were nlyzed over the 16-yer period y considering four 4-yer periods , , , nd , respectively. The 3.8-km swimming, 180-km cycling, 42-km running times, nd totl rce times were nlyzed for the top 10 trithletes in ech ge group from 18 to 64 yers. The prticiption of mster trithletes ( 40 yers old) incresed over the yers, representing on verge 23%, 28%, 37%, nd 48% of totl mle finishers during the four 4-yer periods, respectively. Over the period, trithletes older thn 40 yers significntly improved their performnce in swimming, cycling, running, nd in the totl time tken to complete the rce. The question whether mster Ironmn trithletes hve yet reched limits in their performnce during Ironmn trithlon should e rised. Further studies investigting trining regimes, competition experience, or sociodemogrphic fctors re needed to gin etter insights into the phenomenon of the reltive improvement in ultr-endurnce performnce with dvncing ge. Nowdys, in society with n incresing prt of older dults, successful geing is of highest importnce for the whole popultion (Lee & Tnk, 1997; Wright & Perricelli, 2008). Regulr physicl ctivity is eneficil for overll helth (Lee & Tnk, 1997; Trppe, 2007; Rnsdell et l., 2009), protects from diseses like dietes or hypertension (Willims, 2009) nd even very intense exercise increses longevity (Snchis-Gomr et l., 2011). Athletes who mintin n ctive lifestyle ge grcefully with few helth prolems, while those who lpse into inctivity regress towrd generl popultion norms for fitness, weight control, nd helth prolems (Fulkner et l., 2008). Some studies even hypothesized tht lifestyle fctors hve stronger influences on physiologicl cpcity thn ge (Leyk et l., 2007, 2009). Therefore, mster thletes, defined s thletes who exceeded the ge requisite for success in elite competition, i.e., typiclly 35 to 40 yers (Rnsdell et l., 2009), represent n intriguing model for successful geing (Tnk & Sels, 2008; Wright & Perricelli, 2008; Armptzis et l., 2011). Helth enefits nd fitness re long with socil fctors, enjoyment, nd competition resons for msters prticiption in endurnce events (Reurn & Dscome, 2008). Recent studies hve ccounted for the enormous increse in prticiption of mster thletes (Rnsdell et l., 2009). Mster thletes prticiption rte incresed in different running events; from mrthon (Jokl et l., 2004; Lepers & Cttgni, 2011) to -km (Knechtle et l., 2011d) nd 161-km ultr-mrthons (Hoffmn et l., 2010), while the prticiption rte of younger thletes decresed. It hs een shown tht mster runners represented ~55% of the totl field in mrthon running (Leyk et l., 2009; Lepers & Cttgni, 2011), while their proportion reched ~70% in ultrmrthons (Hoffmn et l., 2010; Knechtle et l., 2011d). The ge-relted decline in performnce in mster thletes relted to chnges in physiologicl (e.g., oxygen crrying cpcity, muscle strength) nd morphologicl (e.g., muscle mss, percentge of ody ft) functionl chrcteristics, hs een evidenced y numerous studies (Rogers et l., 1990; Pollock et l., 1997; Hwkins et l., 2001; Pimentel et l., 2003; Fulkner et l., 2008; Korhonen et l., 2009). In generl, pek endurnce performnce is mintined until 35 yers of ge, followed y modest decreses until yers of ge, nd progressively steeper decline therefter (Tnk & Sels, 2003, 2008; Trppe, 2007; Reurn & Dscome, 2008; Wright & Perricelli, 2008; Rnsdell et l., 2009). The 1

2 Stiefel et l. rtes of the decline in performnce with ge differ etween the sports depending on the specific demnds of the event (Tnk & Sels, 2003; Wright & Perricelli, 2008). In mrthon running (Jokl et l., 2004; Leyk et l., 2007; Lepers & Cttgni, 2011), s well s in long-distnce trithlon (Lepers & Mffiuletti, 2011; Lepers et l., 2012), pek endurnce performnce level is usully ttined t n ge comprised etween 20 nd 40 yers while for ultr-running the ge of pek performnce moves to older ges, i.e., yers (Hoffmn, 2010; Knechtle et l., 2011d). Despite the numerous studies on the ge-relted decline in endurnce performnce, there re few investigtions on the performnce trends over time of mster thletes. Msters performnce trends hve een previously investigted t the New York City Mrthon (Jokl et l., 2004; Lepers & Cttgni, 2011). These studies showed for exmple tht over the period, mle runners older thn 50 yers of ge improved their running performnce (Jokl et l., 2004), while regrding the period, only runners older thn 65 yers improved their performnce. In contrst, performnce times of younger mrthoners t the New York City Mrthon remined quite stle over the lst three decdes (Lepers & Cttgni, 2011). Ironmn trithlon involving successively 3.8-km swimming, 180-km cycling, nd 42.2-km running is often considered s one of the world s most chllenging endurnce events nd represents n intriguing model of ultr-endurnce performnce (Lepers, 2008; Lepers et l., 2010; Knechtle et l., 2011). Compred to mrthon running, Ironmn trithlon is reltively young in the field of endurnce events where the first event ws held in Hwii in 1978 (Lepers, 2008). It hs een previously shown tht for elite trithletes, fter n initil phse of rpid improvement of Ironmn trithlon performnce during the 1980s, there ws reltive plteu, ut t lest in running nd cycling, there were smll improvements (Lepers, 2008). However, the performnce nd prticiption trends of mster thletes in this multi-sport ultrendurnce event hve not een previously investigted. Therefore, the im of the present study ws to nlyze the prticiption nd performnce trends of mle Ironmn trithletes, with specil emphsis on mster trithletes (i.e., 40 yers of ge), t the Ironmn Switzerlnd from 1995 to Respecting existing literture on running, we hypothesized oth n increse in prticiption nd n improvement in mster trithletes performnce over the 16-yer studied period. Mteril nd methods This study ws pproved y the institutionl review ord of St. Gllen, Switzerlnd, with wiver of the requirement for informed consent given tht the study involved the nlysis of pulicly ville dt. The dtset from this study ws otined from the rce wesite ( nd from the rce director. The ge t the time of competition nd swimming, cycling, running, nd totl time performnces of the mle finishers t the Ironmn Switzerlnd were nlyzed from 1995 to Femles were not ccounted in this study ecuse their prticiption in tht ultr-endurnce event ws low (~10% of the totl field). We focused on the Ironmn Switzerlnd s qulifying event for the Ironmn Trithlon World Chmpionship held every yer in Kon, Hwii ( The Ironmn Switzerlnd is with its qulifying plces for the Ironmn Hwii trithlon comprle to the other world s gretest Ironmn events. Between its inception in 1995 nd 2010, totl of mle trithletes finished the Ironmn Switzerlnd. The rce is held ech yer in the city of Zurich during July. The rce consists of 3.8-km swim (two lps) in the lke of Zurich where wetsuits re llowed; 180-km cycling course (two lps) with totl ltitude difference of 1260 m; nd completely flt running course over the mrthon distnce (four lps). Dt nlysis Becuse there is very smll numer of finisher trithletes older thn 64 yers, we considered only mle trithletes ged from 18 to 64 yers. The ge groups distinguish the ctegories s follows: yers, yers, yers, yers, yers, yers, yers, yers, nd yers, respectively. Since the numer of finisher trithletes in the ge group yers ws frequently lower thn 10, the ge-relted chnges in performnce were nlyzed for ge groups comprised etween 18 nd 59 yers. Swimming, cycling, running, nd totl performnce times were converted to minutes nd verged cross the first 10 trithletes of ech ge group. In order to simplify the nlysis over the 16 yers studied period ( ), we pooled dt into four periods of 4 yers: , , , nd , respectively. Therefore, for ech ge group nd ech 4-yer period, 40 time performnces were considered. Sttisticl nlysis Dt re reported s mens stndrd devition (SD) in the text. One-wy nlysis of vrince (ANOVA) ws used to compre the men swimming, cycling, running, nd totl times etween the different ge groups during the period. A two-wy ANOVA with etween fctors of ge nd 4-yer group ws used to compre percent of finishers per ge groups, nd swimming, cycling, running, nd totl times etween the 4-yer periods cross the ge groups. Performnces of one suject could hve een nlyzed severl times, inducing dt is ut ecuse of the low proility tht it hppens, we still considered our groups s independent. Tukey s post hoc nlyses were used to test differences within the ANOVAs when pproprite. A significnce level of P < 0.05 ws used to identify sttisticl significnce. Results Prticiption trends The numer of totl mle finishers incresed cross the four 4-yer periods: : 1819 finishers; : 2992 finishers; : 4364 finishers; nd : 6764 finishers, respectively. The men ge distriution of the mle finishers over the 4-yer periods is displyed in Fig. 1. The men numer of finishers older thn 30 yers of ge incresed during the four 4-yer periods (Fig. 1). During the lst 4-yer period, i.e., , the 5-yer ge rckets with the lrgest prticiption hve een yers nd yers 2

3 Mster trithletes improved their performnce () Numer of finishers () 35 Finishers (% totl) c Age groups (yers) Fig. 1. () Men numer of mle finishers per ge group for the four periods , , , nd t the Ironmn Switzerlnd trithlon. () Men percentge of mle finishers per ge group for the four periods , , , nd t Ironmn Switzerlnd trithlon. Vlues re mens SD., significntly different from , P < 0.05., significntly different from nd , P < c, significntly different from , , nd , P < with on verge more thn 400 finishers per yer. Over the studied period, the percentge of finishers significntly decresed etween 25 nd 34 yers, remined stle for ge group yers since nd significntly incresed etween 40 nd 49 yers (Fig. 1). Mster trithletes ( 40 yers old) represented on verge 23%, 28%, 37%, nd 48% of totl mle finishers during the , , , nd periods, respectively. Age-relted chnges in performnces The reltive contriution of ech prt of the overll Ironmn trithlon performnce remined constnt cross the ges. On verge, the 3.8-km swimming, 180-km cycling, nd 42-km running represented %, %, nd % of the totl time, respectively. The men ge-relted chnges for swimming, cycling, running, nd totl performnces throughout the period re shown in Fig. 2. For the three disciplines nd totl rce time, the performnce times incresed in curviliner mnner with dvncing ge. There ws significnt ge effect for swimming (F = 51.7; P < 0.001), cycling (F = 33.6; P < 0.001), running (F = 82.4 P < 0.001), nd totl (F = 64.8; P < 0.001) times. Swimming, running, nd totl time performnces were significntly lower in the ge groups yer nd older compred to younger ge groups rnging from 18 to 39 yers for swimming nd from 25 to 39 yers for running nd totl time. Cycling performnces were significntly lower in the ge groups yer nd older compred to younger ge groups. Figure 3 shows the ge-relted chnges in swimming, cycling, running, nd totl performnces cross the four 4-yer periods. There ws significnt ge group 4- yer period interction for swimming (F = 6.2; P < 0.001), cycling (F = 11.2; P < 0.001), running (F = 9.5; P < 0.001), nd overll rce time (F = 22.3; 3

4 Stiefel et l. Swim time (min) NS Swim time (% est performnce) ,,c, Cycle time (min) NS Cycle time (% est performnce) ,,,c,,c Run time (min) NS Run time (% est performnce) ,,,,,c,,c Totl time (min) NS Totl time (% est performnce) ,,,,c,,,c 500 Age groups (yers) Age groups (yers) Fig. 2. Age-relted chnges in swimming, cycling, running, nd totl performnces t Ironmn Switzerlnd trithlon. Dt were pooled from 1995 to Vlues re mens SD. NS, nonsignificnt difference etween ge groups. Fig. 3. Age-relted chnges in swimming, cycling, running, nd totl performnces cross the four periods , , , nd t Ironmn Switzerlnd trithlon. For clrity resons, SDs were not shown., significntly different (P < 0.01) from period., significntly different (P < 0.01) from period. c, significntly different (P < 0.01) from period. 4

5 Mster trithletes improved their performnce Totl time (min) yers yers yers yers yers yers Periods yers yers Fig. 4. Chnge in totl time performnces of the different ge groups cross the four periods , , , nd t Ironmn Switzerlnd trithlon. For clrity resons, SDs were not shown. P < 0.001) performnces. Swimming performnce times for ge groups older thn yers significntly decresed over the four periods. Cycling, running, nd totl performnces times for ge groups older thn yers significntly decresed over the four periods. The chnges in totl time performnces of the different ge groups cross the four periods re shown in Fig. 4. The performnce improvement ppered much lrger etween the lst two periods nd thn etween ny other period for the oldest ge groups nd yers, respectively. Discussion The min findings of the present study were () n increse in prticiption of mle mster Ironmn trithletes; they represented during the period lmost 50% of the totl field; nd () n improvement of mster trithletes performnces during the 16-yer studied period, while performnces of trithletes younger thn 40 yers old remined quite stle. Chnges in prticiption of mster trithletes It should e mentioned tht the rce orgniztion of Ironmn Switzerlnd hs limited strt plces during the lst yers, nd this entry cp hs incresed over the yers reching 2200 prticipnts in However, this limittion would proly not chnge the reltive proportion etween the different ge ctegories. Sociologicl nd demogrphic chnges my explin the increse in mster thletes prticiption in Ironmn trithlon. The development nd encourgement of helth nd sport policy ws common gol in the developed world during the pst five decdes (Lee & Tnk, 1997). The helthier wy of life resulted itself in higher rte of ctive dults nd is min fctor for the incresed life expectncy s well (Svidn et l., 2010), wht increses the proility of older competing in endurnce events too (Lee & Tnk, 1997; Rnsdell et l., 2009). Figure 5 shows the reltionship etween the chnges in percentge of the mle Ironmn Switzerlnd finishers nd the chnges in percentge of the Swiss generl popultion per ge group etween the two periods nd , respectively. During the 16-yer studied period, the percentge of ge groups older thn 40 yers incresed for oth Ironmn Switzerlnd finishers nd Swiss popultion, while the percent of ge groups comprised etween 18 nd 34 yers remined stle or decresed. This finding suggests tht the increse in older trithletes prticiption in Ironmn Switzerlnd my e linked to the demogrphic chnges in generl popultion. Interestingly, the reltive proportion of dults older thn 40 yers represented 52.5% of the Swiss popultion in 2010 ( wht is close to their recent reltive Ironmn prticiption rte. However, the proportion of older mster thletes incresed more thn the proportion of those ge groups in the generl Swiss popultion, suggesting tht older people hve chnged their wreness nd lifestyle during these lst two decdes. Indeed, the etter considertion y older people of the positive effects of physicl ctivity upon helth with dvncing ge, i.e., etter 5

6 Stiefel et l. Swiss popultion (%) Ironmn Switzerlnd finishers (%) r ² = 0.53, P = Fig. 5. Reltion etween the chnges in percentge of mle Ironmn Switzerlnd finishers nd the chnges in percentge of the Swiss generl popultion per ge group (18 24 yers, yers, yers, yers, yers, yers, yers, yers, nd yers) etween the two periods nd Dt of the Swiss generl popultion were otined from geing, might e the primry reson for this high prticiption rte (Lee & Tnk, 1997). The high prticiption rte of mle mster Ironmn trithletes my lso e due to gret prt of thletes who hve strted lter in their life with trithlon. The Ironmn trithlon is young in the field of ultr-endurnce event with the first Ironmn Hwii trithlon in tking plce in 1978 (Lepers, 2008). A study on Ironmn Lnzrote in Spin found tht Ironmn competitors hd on verge 6 yers of experience in trithlon, nd they cme from vriety of sporting ckgrounds with most from running ckground, i.e., 28%, followed y swimming nd cycling with 14% nd 13%, respectively (Gulin & Gffney, 1999). The ide of short trining histories is confirmed y n investigtion on mrthoners showing tht the mjority of mster thletes hve trining histories of less thn 7 yers of running (Leyk et l., 2009). Mster thletes were responsile for the considerle increse in totl mle finishers. We could expect tht the prticiption rte of mster trithletes will still increse in the future nd will soon e greter thn the prticiption rte of younger trithletes. Age-relted chnges in performnces The highest Ironmn trithlon performnce level of trithletes ged etween 25 nd 39 yers nd the lesser ge-relted decline in cycling performnce compred with running nd swimming performnces in trithlon re in ccordnce with previous studies (Bernrd et l., 2010; Lepers et l., 2010, 2012; Lepers & Mffiuletti, 2011). Physiologicl (e.g., concentric muscle contrctions) nd mechnicl (e.g., reltion etween mechnicl power output nd velocity, non-weight-ering spect) specificities of cycling compred with the two other disciplines my explin the lower ge-relted decline in cycling performnce (Bernrd et l., 2010; Lepers et l., 2010). Reductions in mximl oxygen uptke (VO 2mx ) re long with the decrese in lctte threshold, the loss of muscle mss, nd the increse in percent ody ft primry resons for the decline in functionl endurnce with ge (Pollock et l., 1997; Hwkins et l., 2001; Reurn & Dscome, 2008; Tnk & Sels, 2008; Korhonen et l., 2009; Rnsdell et l., 2009). The potentil slowing of some of the geing processes through thletic trining is known since the 1960s (Jokl et l., 2004). Physiologicl chrcteristics such s eroic cpcity (e.g., VO 2mx ) nd muscle strength decline t similr rte with incresing ge in mster thletes nd helthy controls (Rogers et l., 1990; Pollock et l., 1997; Person et l., 2002; Michelis et l., 2008). However, the higher seline levels in trined thletes represent n pprent ge dvntge of severl yers compred with ge-mtched sedentry dults (Person et l., 2002; Pimentel et l., 2003; Tnk & Sels, 2003, 2008; Michelis et l., 2008). Endurnce-trined dults hve impressive higher levels of VO 2mx compred to their sedentry counterprts, i.e., ~50 ml/kg/min vs ~33 ml/kg/min t the ge of ~60 yers (Rogers et l., 1990; Pollock et l., 1997; Hwkins et l., 2001; Pimentel et l., 2003). The numer of muscle fiers nd motor units decreses fter the ge of 50 yers, with greter loss from the fst-contrcting type II muscle fiers (Fulkner et l., 2008; Korhonen et l., 2009). Lifelong 6

7 Mster trithletes improved their performnce physicl ctivity does not pper to hve n impct on the loss in fier numer, ut the loss of fiers cn e uffered to some degree y hypertrophy of fiers tht remin. Especilly, older thletes who do weight trining seem to etter mintin their muscle mss (Pollock et l., 1997). Body ft is ~10% higher in sedentry compred to endurnce trined dults nd it increses over lifespn, in trined from ~10% to ~20%, while in untrined from ~20% to ~30% (Pollock et l., 1997; Pimentel et l., 2003). Chnges in performnce of mster trithletes According to our hypothesis, the mster trithletes improved their performnce cross the 16-yer study period. The economic development, with mjor technologicl, nutritionl, nd medicl dvnces, offered constnt elevtion of life resources in developed countries (Berthelot et l., 2008). This implied for thletes n improvement of trining fcilities, coching, trining techniques, nutritionl strtegies, nd equipment. Becuse of sociologicl chnges, older thletes ttined more nd more ccess to these improved trining possiilities (Rnsdell et l., 2009). Recent mster thletes re likely to hve hd longer nd etter ccess to these improved trining possiilities thn mster thletes in erlier yers. Wright nd Perricelli (2008) found in demogrphic survey of ~2600 prticipnts in the Ntionl Senior Olympic Gmes, tht most of the prticipnts were middle clss nd well educted. Approximtely 80% hd grduted from college studies nd 32% held postgrdute degrees. They concluded tht the Senior Olympic Gmes prticipnts my reflect the helth enefits of their socioeconomic sttus. Performnce improvements lso hppened ecuse of the considerle increse of mster Ironmn trithletes prticiption. It is ovious tht the higher prticiption rte increses the possiility of hving etter thletes. More successful thletes retin higher intrinsic motivtion to trin nd re more likely to compete in endurnce events (Medic et l., 2007). Accordingly, the etter condition of older thletes is likely to increse the competitive spirit, the prticiption, nd the performnce (Medic et l., 2007; Rnsdell et l., 2009). The dvncement of trining qulity is in ll proility reson for these performnce trends. The performnce improvements of mster trithletes in such n ultrendurnce discipline re mzing. Erlier investigtions suggested tht the performnce decline with dvncing ge ws ecuse of decresed trining volumes nd intensity. The reduced trining ws result of sociologicl fctors like incresed work nd fmily commitments, ehviorl fctors such s less intrinsic drive to trin hrd, nd physiologicl fctors like longer time needed to recover nd higher risk for injuries (Reurn & Dscome, 2008; Tnk & Sels, 2008; Korhonen et l., 2009; Rnsdell et l., 2009). Trining for n Ironmn trithlon is very demnding (O Toole, 1989; Gulin & Gffney, 1999; Knechtle et l., 2011c). It hs een shown tht recretionl Ironmn Switzerlnd prticipnts spend every week lmost 14 h trining; e.g., 2.4 h for swimming, 6.7 h for cycling, nd 4.7 h for running (Knechtle et l., 2011c). For comprison, hlf mrthoners spend 4 h (Knechtle et l., 2010) nd -km ultr-runners spend 7 h per week running (Knechtle et l., 2011). It is ovious tht to plce in the top 10 of n ge group ctegory, oth lrge trining volume nd high trining intensity re required (Gulin & Gffney, 1999; Knechtle et l., 2011, 2011c). However, it hs een suggested tht mster thletes could optimize their qulity of trining, so they could reduce their trining volume to sve time for dequte recovery nd sty free of injuries (Rnsdell et l., 2009). It hs een shown tht sport disciplines presented their highest progression curvture during their erly phse (Desgorces et l., 2008). Although Ironmn trithlon is young ultr-endurnce event, elite Ironmn trithletes seem to hve reched their performnce limits (Desgorces et l., 2008; Lepers, 2008) s it is oserved in more trditionl sports like mrthon (Jokl et l., 2004; Lepers & Cttgni, 2011). This rpid evolution of performnce in Ironmn trithlon my e sed on the rpidly growing populrity of the event nd enefits from the dvnces previously mde in the three sports involved in trithlon (Desgorces et l., 2008). In ddition, the fct tht some elite swimmers, cyclists, nd runners my strt Ironmn trithlon competitions lter in their life, hs nd could help to increse the level of performnce in mster thletes. Unfortuntely, we hve no informtion out the sportive ckground of these successful mster trithletes. Even if recretionl mster thletes hd reltively short trining histories nd little experience in trithlon (Gulin & Gffney, 1999; Leyk et l., 2009), these successful thletes proly hve lifelong histories of physicl ctivity (Wright & Perricelli, 2008). Methodologicl considertions We re wre of some unvoidle limittions of our study. First, femles were not ccounted ecuse of their lower prticiption rte. Previous studies on the gerelted decline in performnce concluded greter decline in femles thn in mles, ut it ws confounded y the reltive smller numer of femle vs mles especilly in the older ge groups (Tnk & Sels, 2003, 2008; Rnsdell et l., 2009; Lepers & Mffiuletti, 2011). Second, there ws no informtion out the individul fctors, like physiologicl (e.g., eroic cpcity) nd nthropometric (e.g., ody weight, len ody mss) prmeters, trining regimes, nd competition experience. Third, the influence of environmentl conditions such s wether conditions nd chnges in sport equipment ws unknown. Fourth, estimting the chnge in 7

8 Stiefel et l. percent of dropout in the different ge groups cross the yers would hve een interesting ut unfortuntely in the present study, we did not hve the exct numer of strters. Despite these limittions, the gret numer of mle trithletes considered in the present study my give new insight on the ge-relted decline in ultr-endurnce performnce. Perspectives The prticiption of mle mster Ironmn trithletes in Ironmn Switzerlnd incresed over the period, nd it will proly still grow in future. The improvement of mster trithletes performnce suggests tht these thletes my not rech their limits in Ironmn trithlon performnce. It would e interesting to nlyze the chnges in ge-relted decline in performnce t the World Ironmn trithlon chmpionship tking plce ech yer in Hwii during the lst three decdes nd to compre with the present results oserved on qulifying event. Further studies investigting trining regimes, competition experience, or socio-demogrphic fctors re needed to gin etter insights into the phenomenon of the reltive improvement in ultr-endurnce performnce with dvncing ge. Key words: geing, ge-relted decline, sport, endurnce performnce. Acknowledgements We thnk Mrk Klenthous, London, Englnd, for his help in trnsltion. References Armptzis A, Degens H, Bltzopoulos V, Rittweger J. Why do older sprinters rech the finish line lter? Exerc Sport Sci Rev 2011: 39: Bernrd T, Sultn F, Lepers R, Husswirth C, Brisswlter J. Age-relted decline in Olympic trithlon performnce: effect of locomotion mode. Exp Aging Res 2010: 36: Berthelot G, Thiult V, Tfflet M, Escolno S, El Helou N, Jouven X, Hermine O, Toussint JF. The citius end: world records progression nnounces the completion of rief ultr-physiologicl quest. PLoS ONE 2008: 6: e1552. Desgorces FD, Berthelot G, El Helou N, Thiult V, Guillume M, Tfflet M, Hermine O, Toussint JF. From Oxford to Hwii ecophysiologicl rriers limit humn progression in ten sport monuments. PLoS ONE 2008: 3: e3653. Fulkner JA, Dvis CS, Mendis CL, Brooks SV. The ging of elite mle thletes: ge-relted chnges in performnce nd skeletl muscle structure nd function. Clin J Sport Med 2008: 18: Gulin JP, Gffney PT. Ultrendurnce trithlon prticiption: typicl rce preprtion of lower level trithletes. J Sports Med Phys Fitness 1999: 39: Hwkins SA, Mrcell TJ, Victori Jque S, Wiswell RA. A longitudinl ssessment of chnge in VO 2mx nd mximl hert rte in mster thletes. Med Sci Sports Exerc 2001: 33: Hoffmn MD. Performnce trends in 161-km ultrmrthons. Int J Sports Med 2010: 31: Hoffmn MD, Ong JC, Wng G. Historicl nlysis of prticiption in 161 km ultrmrthons in North Americ. Int J Hist Sport 2010: 27: Jokl P, Sethi PM, Cooper AJ. Mster s performnce in the New York City Mrthon Br J Sports Med 2004: 38: Knechtle B, Knechtle P, Lepers R. Prticiption nd performnce trends in ultr-trithlons from 1985 to Scnd J Med Sci Sports 2011: 21: e82 e90. Knechtle B, Knechtle P, Rosemnn T, Senn O. Sex differences in ssocition of rce performnce, skin-fold thicknesses, nd trining vriles for recretionl hlf-mrthon runners. Percept Mot Skills 2010: 111: Knechtle B, Knechtle P, Rosemnn T, Senn O. Wht is ssocited with rce performnce in mle -km ultr-mrthoners nthropometry, trining or mrthon est time? J Sports Sci 2011: 29: Knechtle B, Knechtle P, Rüst CA, Rosemnn T. A comprison of nthropometric nd trining chrcteristics of Ironmn trithletes nd Triple Iron ultr-trithletes. J Sports Sci 2011c: 29: Knechtle B, Rüst CA, Rosemnn T, Lepers R. Age-relted chnges in -km ultr-mrthon running performnce. Age (Dordr) 2011d: doi: 10.7/s Korhonen MT, Mero AA, Alén M, Sipilä S, Häkkinen K, Liikvinio T, Viitslo JT, Hverinen MT, Suominen H. Biomechnicl nd skeletl muscle determinnts of mximum running speed with ging. Med Sci Sports Exerc 2009: 41: Lee MS, Tnk K. Significnce of helth fitness pprisl in n ging society. Appl Humn Sci 1997: 16: Lepers R. Anlysis of Hwii Ironmn performnces in elite trithletes from 1981 to Med Sci Sports Exerc 2008: 40: Lepers R, Cttgni T. Do older thletes rech limits in their performnce during mrthon running? Age (Dordr) 2011: doi: 10.7/s z Lepers R, Mffiuletti N. Age nd gender interctions in ultr-endurnce performnce: insight from trithlon. Med Sci Sports Exerc 2011: 43: Lepers R, Rüst CA, Stpley P, Knechtle B. Reltive improvements in endurnce performnce with ge: evidence from 25 yers of Hwii Ironmn rcing. Age (Dordr), 2012: 34(3): Lepers R, Sultn F, Bernrd T, Husswirth C, Brisswlter J. Age-relted chnges in trithlon performnces. Int J Sports Med 2010: 31: Leyk D, Erley O, Gorges W, Ridder D, Rüther T, Wunderlich M, Sievert A, Essfeld D, Piekrski C, Erren T. Performnce, trining nd lifestyle prmeters of mrthon runners ged yers: results of the PACE-study. Int J Sports Med 2009: 30: Leyk D, Erley O, Ridder D, Leurs M, Rüther T, Wunderlich M, Sievert A, Bum K, Essfeld D. Age-relted chnges in mrthon nd hlf-mrthon performnces. Int J Sports Med 2007: 28:

9 Mster trithletes improved their performnce Medic N, Strkes JL, Young BW. Exmining reltive ge effects on performnce chievement nd prticiption rtes in msters thletes. J Sports Sci 2007: 25: Michelis I, Kwiet A, Gst U, Boshof A, Antvorskov T, Jung T, Rittweger J, Felsenerg D. Decline of specific pek jumping power with ge in mster runners. J Musculoskelet Neuronl Interct 2008: 8: O Toole ML. Trining for ultrendurnce trithlons. Med Sci Sports Exerc 1989: 21: Person SJ, Young A, Mcluso A, Devito G, Nimmo MA, Coold M, Hrridge SD. Muscle function in elite mster weightlifters. Med Sci Sports Exerc 2002: 34: Pimentel AE, Gentile CL, Tnk H, Sels DR, Gtes PE. Greter rte of decline in mximl eroic cpcity with ge in endurnce-trined thn in sedentry men. J Appl Physiol 2003: 94: Pollock ML, Mengelkoch LJ, Grves JE, Lowenthl DT, Limcher MC, Foster C, Wilmore JH. Twenty-yer follow-up of eroic power nd ody composition of older trck thletes. J Appl Physiol 1997: 82: Rnsdell LB, Vener J, Huerty J. Mster thletes: n nlysis of running, swimming nd cycling performnce y ge nd gender. J Exerc Sci Fit 2009: 7: Reurn P, Dscome B. Endurnce performnce in msters thletes. Eur Rev Aging Phys Act 2008: 5: Rogers MA, Hgerg JM, Mrtin WH 3rd, Ehsni AA, Holloszy JO. Decline in VO 2mx with ging in mster thletes nd sedentry men. J Appl Physiol 1990: 68: Snchis-Gomr F, Olso-Gonzlez G, Corell D, Gomez-Crer MC, Vin J. Incresed verge longevity mong the Tour de Frnce cyclists. Int J Sports Med 2011: 32: Svidn A, Junker C, Cerny T, Ess S. Premture deths in Switzerlnd from : cuses nd trends. Swiss Med Wkly 2010: 24 (140): w Tnk H, Sels DR. Invited review: dynmic exercise performnce in msters thletes: insight into the effects of primry humn ging on physiologicl functionl cpcity. J Appl Physiol 2003: 95: Tnk H, Sels DR. Endurnce exercise performnce in msters thletes: ge-ssocited chnges nd underlying physiologicl mechnisms. J Physiol 2008: 586: Trppe S. Mrthon runners: how do they ge? Sports Med 2007: 37: Willims PT. Lower prevlence of hypertension, hypercholesterolemi, nd dietes in mrthoners. Med Sci Sports Exerc 2009: 41: Wright VJ, Perricelli BC. Age-relted rtes of decline in performnce mong elite senior thletes. Am J Sports Med 2008: 36:

TECHNICAL BULLETINApril 2016

TECHNICAL BULLETINApril 2016 SYN-86 TECHNICAL BULLETINApril 216 Synovex One-Feedlot Implnts in Feedlot Steers: Phse IIIB Studies in Nersk nd Texs Zoetis Florhm Prk, NJ 7932 Summry Two Phse IIIB studies 1,2 were conducted in feedlot

More information

Department of Animal and Aquacultural Sciences, Norwegian University of life Science, PO-Box N-1432 Ås, Norway.

Department of Animal and Aquacultural Sciences, Norwegian University of life Science, PO-Box N-1432 Ås, Norway. Presenttion in session H11.5 * The ehviour of horses in different pddock sizes, with nd without exercise G.H.M. Jørgensen * & K.E. Bøe. Deprtment of Animl nd Aquculturl Sciences, Norwegin University of

More information

Minnesota s Wild Turkey Harvest Fall 2016, Spring 2017

Minnesota s Wild Turkey Harvest Fall 2016, Spring 2017 Minnesot s Wild Turkey Hrvest Fll 2016, Spring 2017 Lindsey Messinger Frmlnd Wildlife Popultions nd Reserch Group Minnesot Deprtment of Nturl Resources Mdeli, Minnesot 15 August 2017 Summry of Seson Structure

More information

EMBASSY SWIMMING POOL & FITNESS SUITE Fitness Programme June - September 2017

EMBASSY SWIMMING POOL & FITNESS SUITE Fitness Programme June - September 2017 EMBASSY POOL & FITNESS SUITE Fitness Progrmme June - September 2017 7m Sturdy dy dy sdy nesdy sdy dy 7m - using the benefits of wter resistnce for low impct crdio nd muscle toning workout tht cn suit wide

More information

Open Access Regression Analysis-based Chinese Olympic Games Competitive Sports Strength Evaluation Model Research

Open Access Regression Analysis-based Chinese Olympic Games Competitive Sports Strength Evaluation Model Research Send Orders for Reprints to reprints@benthmscience.e The Open Cybernetics & Systemics Journl, 05, 9, 79-735 79 Open Access Regression Anlysis-bsed Chinese Olympic Gmes Competitive Sports Strength Evlution

More information

Nick Willis Discipline: Middle distance running Specialist events: 800m and 1500m

Nick Willis Discipline: Middle distance running Specialist events: 800m and 1500m Olympic Eduction Discipline: Middle distnce running Specilist events: 800m nd 1500m ws born in Lower Hutt (ner Wellington) in 1983.Running tlent obviously runs in the Willis fmily, s Nick nd his brother

More information

Small Game Hunter Lead Shot Communication Study. Executive Summary. A cooperative study conducted by:

Small Game Hunter Lead Shot Communication Study. Executive Summary. A cooperative study conducted by: 8 Smll Gme Hunter Led Shot Communiction Study Executive Summry A coopertive study conducted y: Minnesot Coopertive Fish nd Wildlife Reserch Unit Minnesot Deprtment of Nturl Resources 9 Smll Gme Hunter

More information

A Measurement Framework for National Key Performance Measures

A Measurement Framework for National Key Performance Measures MINISTERIAL COUNCIL ON EDUCATION, EMPLOYMENT, TRAINING AND YOUTH AFFAIRS Performnce Mesurement nd Reporting Tskforce Working for qulity eduction outcomes A Mesurement Frmework for Ntionl Key Performnce

More information

First Aid in School Policy

First Aid in School Policy First Aid in School Policy Approved y the Governing Body of West Monkton Church of Englnd Primry School: Reviewed y Governors nd Agreed: Septemer 2017 At West Monkton we im to e n inspirtionl community,

More information

The infection of tench (Tinca tinca) with Ligula intestinalis plerocercoids in Lake Beysehir (Turkey)

The infection of tench (Tinca tinca) with Ligula intestinalis plerocercoids in Lake Beysehir (Turkey) Bull. Eur. Ass. Fish Pthol., 23(5) 2003, 223 The infection of tench (Tinc tinc) with Ligul intestinlis plerocercoids in Lke Beysehir (Turkey) H. Yvuzcn Yıldız*, A. S. Korkmz nd Ö. Zencir Ankr University,

More information

Contents TRIGONOMETRIC METHODS PROBABILITY DISTRIBUTIONS

Contents TRIGONOMETRIC METHODS PROBABILITY DISTRIBUTIONS ontents UNIT 7 TRIGONOMETRI METHODS Lesson 1 Trigonometric Functions................... 462 1 onnecting ngle Mesures nd Liner Mesures.............. 463 2 Mesuring Without Mesuring.........................

More information

Characteristics, Expenditures, and Economic Impact of Resident and Nonresident Hunters and Anglers in North Dakota, , Season and Trends

Characteristics, Expenditures, and Economic Impact of Resident and Nonresident Hunters and Anglers in North Dakota, , Season and Trends Agriculturl Economics Report No. 389 June 1998 Chrcteristics, Expenditures, nd Economic Impct of Resident nd Nonresident Hunters nd Anglers in North Dkot, 1996-97, Seson nd Trends Tin D. Lewis Jy A. Leitch

More information

Fitness, obesity and risk of heat illness among army trainees

Fitness, obesity and risk of heat illness among army trainees Occuptionl Medicine Advnce Access published July 14, 2014 Occuptionl Medicine doi:10.1093/occmed/kqu062 Fitness, obesity nd risk of het illness mong rmy trinees S. A. Bedno 1, N. Urbn 2,3, M. R. Boivin

More information

Intended Hydration Strategies and Knowledge of Exercise-Associated Hyponatraemia in Marathon Runners: A Questionnaire-Based Study

Intended Hydration Strategies and Knowledge of Exercise-Associated Hyponatraemia in Marathon Runners: A Questionnaire-Based Study Journl of Athletic Trining 2018;53(7):696 702 doi: 10.4085/1062-6050-125-17 Ó by the Ntionl Athletic Triners Assocition, Inc www.ntjournls.org Exertionl Het Illness Intended Hydrtion Strtegies nd Knowledge

More information

ERRATA for Guide for the Development of Bicycle Facilities, 4th Edition (GBF-4)

ERRATA for Guide for the Development of Bicycle Facilities, 4th Edition (GBF-4) Dvid Bernhrdt, P.E., President Commissioner, Mine Deprtment of Trnsporttion Bud Wright, Executive Director 444 North Cpitol Street NW, Suite 249, Wshington, DC 20001 (202) 624-5800 Fx: (202) 624-5806 www.trnsporttion.org

More information

INVESTIGATION 2. What s the Angle?

INVESTIGATION 2. What s the Angle? INVESTIGATION 2 Wht s the Angle? In the previous investigtion, you lerned tht when the rigidity property of tringles is comined with the ility to djust the length of side, the opportunities for useful

More information

Announcements. CS 188: Artificial Intelligence Spring Today. P4: Ghostbusters. Exact Inference in DBNs. Dynamic Bayes Nets (DBNs)

Announcements. CS 188: Artificial Intelligence Spring Today. P4: Ghostbusters. Exact Inference in DBNs. Dynamic Bayes Nets (DBNs) CS 188: Artificil Intelligence Spring 2010 Lecture 21: DBNs, Viteri, Speech Recognition 4/8/2010 Written 6 due tonight Project 4 up! Due 4/15 strt erly! Announcements Course contest updte Plnning to post

More information

BASKETBALL SPEED AND AGILITY

BASKETBALL SPEED AND AGILITY SKETLL SPEED ND GILITY Off court Speed and gility Work: ox gility Drills: cone set up 5 yards apart, read and follow description Drill 1 : (1-2-3-4) Sprint around cones, make hard cuts Drill 2: 1-2 Sprint,

More information

Analysis of performance and age of the fastest 100- mile ultra-marathoners worldwide

Analysis of performance and age of the fastest 100- mile ultra-marathoners worldwide CLINICAL SCIENCE Analysis of performance and age of the fastest 100- mile ultra-marathoners worldwide Christoph Alexander Rüst, I Beat Knechtle, I,II Thomas Rosemann, I Romuald Lepers III I University

More information

The development of a truck concept to allow improved direct vision of vulnerable road users by drivers

The development of a truck concept to allow improved direct vision of vulnerable road users by drivers Loughorough University Institutionl Repository The development of truck concept to llow improved direct vision of vulnerle rod users y drivers This item ws sumitted to Loughorough University's Institutionl

More information

Garden City Community Park - Bike Terrain Park Consultation Process

Garden City Community Park - Bike Terrain Park Consultation Process To: From: City of Richmond Prks, Recretion nd Culturl Services Committee Mike Redpth, Senior Mnger, Prks Report to Committee To V~-- \Wht A 20 x>r3. \ Dte: My 13, 2013 File: 06-2345-20- MNOR1NoI01 Re:

More information

and that excessive pronation is one major cause of foot and leg problems of runners. The treatment modahties used on these runners are shown

and that excessive pronation is one major cause of foot and leg problems of runners. The treatment modahties used on these runners are shown Foot orthotic devices to modify selected spects of lower extremity mechnics* B. T. BATES,&dgger; Ph.D., L. R. OSTERNIG, Ph.D., B. MASON, M.S., AND L. S. JAMES, M.D., Eugene, Oregon From the Biomechnics-Sports

More information

2014 WHEAT PROTEIN RESPONSE TO NITROGEN

2014 WHEAT PROTEIN RESPONSE TO NITROGEN 2014 WHEAT PROTEIN RESPONSE TO NITROGEN Aron Wlters, Coopertor Ryford Schulze, Coopertor Dniel Hthcot, Extension Progrm Specilist Dr. Clrk Neely, Extension Stte Smll Grins Specilist Ryn Collett, Extension

More information

Exploring Impacts of Countdown Timers on Queue Discharge Characteristics of Through Movement at Signalized Intersections

Exploring Impacts of Countdown Timers on Queue Discharge Characteristics of Through Movement at Signalized Intersections Avilble online t www.sciencedirect.com ScienceDirect Procedi - Socil nd Behviorl Scienc es 96 ( 2013 ) 255 264 13th COTA Interntionl Conference of Trnsporttion Professionls (CICTP 2013) Exploring Impcts

More information

Unintentional Drownings Among New York State Residents,

Unintentional Drownings Among New York State Residents, Reserch Articles Unintentionl Drownings Among New York Stte Residents, 1988 1994 Mrilyn L. Browne, MS Elizbeth L. Lewis-Michl, PhD Alice D. Strk, MPH, DrPH SYNOPSIS Objective. This study exmines situtions

More information

CS 188: Artificial Intelligence Spring Announcements

CS 188: Artificial Intelligence Spring Announcements CS 188: Artificil Intelligence Spring 2011 Lecture 19: Dynmic Byes Nets, Nïve Byes 4/6/2011 Pieter Aeel UC Berkeley Slides dpted from Dn Klein. Announcements W4 out, due next week Mondy P4 out, due next

More information

For citation purposes, the electronic version is the definitive version of this article:

For citation purposes, the electronic version is the definitive version of this article: Professionl I s s u e s Use of Mouthgurds Among 12- to 14-Yer-Old Ontrio Schoolchildren Kusr Sdi Fkhruddin, BDS, MSc; Hereni P. Lwrence, DDS, MSc, PhD; Dvid J. Kenny, DDS, PhD, FRCD(C); Dvid Locker, BDS,

More information

An Indian Journal FULL PAPER ABSTRACT KEYWORDS. Trade Science Inc. The tennis serve technology based on the AHP evaluation of consistency check

An Indian Journal FULL PAPER ABSTRACT KEYWORDS. Trade Science Inc. The tennis serve technology based on the AHP evaluation of consistency check [Type text] [Type text] [Type text] ISSN : 097-7 Volume 0 Issue 0 BioTechnology 0 An Indin Journl FULL PAPER BTAIJ, 0(0), 0 [-7] The tennis serve technology bsed on the AHP evlution of consistency check

More information

APPLICATION OF POLISH CALCIUM SORBENTS IN CARBONATE LOOPING

APPLICATION OF POLISH CALCIUM SORBENTS IN CARBONATE LOOPING Physicochem. Probl. Miner. Process. 49(), 203, 20 www.minproc.pwr.wroc.pl/journl/ Physicochemicl Problems of Minerl Processing ISSN 643-049 (print) ISSN 2084-4735 (online) Received April 3, 202; reviewed;

More information

CHAPTER 4. Surface Root-zone Water Content and Bentgrass Water Stress. During Drydown for Selected Putting Green Construction.

CHAPTER 4. Surface Root-zone Water Content and Bentgrass Water Stress. During Drydown for Selected Putting Green Construction. CHAPTER 4 Surfce Root-zone Wter Content nd Bentgrss Wter Stress During Drydown for Selected Putting Green Construction Systems ABSTRACT Creeping entgrss, cool-seson grss used extensively in golf courses

More information

Age and gender interactions in short distance triathlon performance

Age and gender interactions in short distance triathlon performance Zurich Open Repository and Archive University of Zurich Main Library Strickhofstrasse 39 CH-8057 Zurich www.zora.uzh.ch Year: 2013 Age and gender interactions in short distance triathlon performance Etter,

More information

An Analysis of the Visual Demands Associated with Aviation Maintenance Inspectors. The Ohio State University College of Optometry, Columbus, OH 2

An Analysis of the Visual Demands Associated with Aviation Maintenance Inspectors. The Ohio State University College of Optometry, Columbus, OH 2 An Anlysis of the Visul Demnds Associted with Avition Mintennce Inspectors Gregory W. Good 1, Json J. Nichols 1, Mnoj V. Subbrm 1, Vn B. Nkgwr 2, Ronld W. Montgomery 2 1 The Ohio Stte University College

More information

1 Measurement. What you will learn. World s largest cylindrical aquarium. Australian Curriculum Measurement and Geometry Using units of measurement

1 Measurement. What you will learn. World s largest cylindrical aquarium. Australian Curriculum Measurement and Geometry Using units of measurement Austrlin Curriulum Mesurement nd Geometry Using units of mesurement hpter 1 Mesurement Wht you will lern 1.1 1.2 1.3 1.4 1.5 1.6 1.7 1.8 Conversion of units Perimeter Cirumferene Are Are of irle Surfe

More information

Ferskvannsavlusing small-scale tests: Short-term exposure tests to elucidate handling effects.

Ferskvannsavlusing small-scale tests: Short-term exposure tests to elucidate handling effects. Ferskvnnsvlusing smll-scle tests: Short-term exposure tests to elucidte hndling effects. The use of freshwter to control infesttions of the se louse Lepeophtheirus slmonis K on Atlntic slmon Slmo slr L.

More information

1970 BRITISH COHORT STUDY: SURVEY

1970 BRITISH COHORT STUDY: SURVEY 1970 BRITISH COHORT STUDY: 2016-18 SURVEY Self-completion Questionnire HOW TO FILL IN THE QUESTIONNAIRE Plese complete the questionnire using blck or blue ink. The questionnire will be red by scnner, so

More information

Multiple Meaning Words: Grade 3 to 5 More Teaching Tools at

Multiple Meaning Words: Grade 3 to 5 More Teaching Tools at pool pool pool pool smll re of still wter, typiclly one formed nturlly Pools of wter formed on the street fter the rin. smll, shllow ptch of liquid lying on surfce There s pool of oil under our cr. deep

More information

ABSTRACT VARIATION AMONG FISH SPECIES IN THE STOICHIOMETRY OF NUTRIENT EXCRETION. by Lisette Esmeralda Torres

ABSTRACT VARIATION AMONG FISH SPECIES IN THE STOICHIOMETRY OF NUTRIENT EXCRETION. by Lisette Esmeralda Torres ABSTRACT VARIATION AMONG FISH SPECIES IN THE STOICHIOMETRY OF NUTRIENT EXCRETION by Lisette Esmerld Torres This study investigtes how nutrient cycling rtes nd rtios vry mong fish species nd with body size,

More information

Biomechanical Analysis of Pressure Redistribution

Biomechanical Analysis of Pressure Redistribution Biomechnicl Anlysis Redistribution Dul PegAssist Conur Bllet-Style Sports (neutrl shoe) Dul Chrcteristics Dul + Generl PegAssist + Conur All mpping results re bsed upon following prmeters: Test Person

More information

AHP-based tennis service technical evaluation consistency test

AHP-based tennis service technical evaluation consistency test Avilble online.jocpr.com Journl of Chemicl nd Phrmceuticl Reserch, 0, ():7-79 Reserch Article ISSN : 097-78 CODEN(USA) : JCPRC AHP-bsed tennis service technicl evlution consistency test Mio Zhng Deprtment

More information

The Discussion of this exercise covers the following points: The open-loop Ziegler-Nichols method. The open-loop Ziegler-Nichols method

The Discussion of this exercise covers the following points: The open-loop Ziegler-Nichols method. The open-loop Ziegler-Nichols method Exercise 6-3 Level Process Control EXERCISE OBJECTIVE In this exercise, you will perform PID control of level process. You will use the open-loop step response method to tune the controller. DISCUSSION

More information

S. FURDEAN, D. LALESCU, Sandra Antonia MIHAILOV, A. GROZEA*

S. FURDEAN, D. LALESCU, Sandra Antonia MIHAILOV, A. GROZEA* GROWTH DYNAMIC OF THE MAIN MORPHOLOGICAL TRAITS IN A STERLET (ACIPENSER RUTHENUS) POPULATION REARED INTO RECIRCULATING AQUACULTURE SYSTEM, FROM 2 TO 6 MONTHS OLD S. FURDEAN, D. LALESCU, Sndr Antoni MIHAILOV,

More information

Working Paper: Reversal Patterns

Working Paper: Reversal Patterns Remember to welcome ll ides in trding. AND remember to reserve your opinion until you hve independently vlidted the ide! Working Pper: Reversl Ptterns Working Pper In this pper I wnt to review nd (hopefully)

More information

PCT MINIMUM DOCUMENTATION

PCT MINIMUM DOCUMENTATION Ref.: PCT Minimum Documenttion pge: 4.1.1 PCT MINIMUM TION INVENTORY S CCORDING TO PCT RULE 34.1 (PERIOD FROM 1920 TO 2000) Explntory Notes 1. On the following pges is given the inventory of ptent documents,

More information

IGF Research Project N Safer High Heels

IGF Research Project N Safer High Heels IGF Reserch Project 17868 N Sfer High Heels Survivl of compnies in the fshion nd sesonlly dependent sectors such s the footwer industry depends upon their ility to implement the ltest trends. Shoe ottoms

More information

Renewable Energy xxx (2011) 1e10. Contents lists available at SciVerse ScienceDirect. Renewable Energy

Renewable Energy xxx (2011) 1e10. Contents lists available at SciVerse ScienceDirect. Renewable Energy Renewle Energy xxx () e Contents lists ville t SciVerse ScienceDirect Renewle Energy journl homepge: www.elsevier.com/locte/renene Wke effect in wind frm performnce: Stedy-stte nd dynmic ehvior F. González-Longtt

More information

Chp. 3_4 Trigonometry.notebook. October 01, Warm Up. Pythagorean Triples. Verifying a Pythagorean Triple... Pythagorean Theorem

Chp. 3_4 Trigonometry.notebook. October 01, Warm Up. Pythagorean Triples. Verifying a Pythagorean Triple... Pythagorean Theorem Chp. 3_4 Trigonometry.noteook Wrm Up Determine the mesure of the vrile in ech of the following digrms: x + 2 x x 5 x + 3 Pythgoren Theorem - is fundmentl reltionship mongst the sides on RIGHT tringle.

More information

1985 BFS CLINICS. BFS Clinic-Assembly in Kamloops, British Columbia. Westsyde High School. Bob Bridges is the Football Coach.

1985 BFS CLINICS. BFS Clinic-Assembly in Kamloops, British Columbia. Westsyde High School. Bob Bridges is the Football Coach. 1985 BFS CLINICS BFS Clinic-Assembly in Kmloops, British Columbi. Westsyde High School. Bob Bridges is the Footbll Coch. BFS Clinicin Rick Anderson detils BFS Nutrition Progrm t Lke Hvsu High School in

More information

Standard Weight (W s ) Equation and Length Categories for Shovelnose Sturgeon

Standard Weight (W s ) Equation and Length Categories for Shovelnose Sturgeon North Americn Journl of Fisheries Mngement 18:9 7, 18 Copyright by the Americn Fisheries Society 18 Stndrd Weight (W s ) Eqution nd Length Ctegories for Shovelnose Sturgeon MICHAEL C. QUIST,* CHRISTOPHER

More information

8/1/2012. Introduction. Oxidative Stress. Oxidative Stress. Antioxidants. Antioxidant Mechanisms H2 O. Oxidative Stress is a hot topic:

8/1/2012. Introduction. Oxidative Stress. Oxidative Stress. Antioxidants. Antioxidant Mechanisms H2 O. Oxidative Stress is a hot topic: Introduction Oxidtive Stress is hot topic: ANTIOXIDANT RESEARCH & ITS APPLICATION FOR USE IN EXERCISING HORSES Crey A. Willims, Ph.D. Associte Professor Google = just under 8 million results Add horse

More information

ORIGINAL RESEARCH Effects of Different Bedding Materials on the Behavior of Horses Housed in Single Stalls

ORIGINAL RESEARCH Effects of Different Bedding Materials on the Behavior of Horses Housed in Single Stalls ORIGINAL RESEARCH Effects of Different Bedding Mterils on the Behvior of Horses Housed in Single Stlls Hnn Werhhn, MSc, Engel F. Hessel, PD Dr, Irene Bchhusen, MSc, nd Hermn F.A. Vn den Weghe, Prof Dr

More information

2014 Victorian Shooting Championship

2014 Victorian Shooting Championship 2014 Victorin Shooting Chmpionship VPCI, in conjunction with the Stte Coches nd the Stte Umpires invite ll PFA licensed petnque plyers in the Stte of Victori to tke prt in the 2014 Victorin Shooting Chmpionship.

More information

Grade 6. Mathematics. Student Booklet SPRING 2011 RELEASED ASSESSMENT QUESTIONS. Record your answers on the Multiple-Choice Answer Sheet.

Grade 6. Mathematics. Student Booklet SPRING 2011 RELEASED ASSESSMENT QUESTIONS. Record your answers on the Multiple-Choice Answer Sheet. Grde 6 Assessment of Reding, Writing nd Mthemtics, Junior Division Student Booklet Mthemtics SPRING 211 RELEASED ASSESSMENT QUESTIONS Record your nswers on the Multiple-Choice Answer Sheet. Plese note:

More information

Nanobiophysical exploration of transthyretin amyloid fibrils Final report

Nanobiophysical exploration of transthyretin amyloid fibrils Final report Nnoiophysicl explortion of trnsthyretin myloid firils Finl report Summry In this work we used AFM to follow the myloidogenetic pthwy of trnsthyretin (TTR) y imging the events leding to the formtion of

More information

Research Article Antioxidant Status in Elite Three-Day Event Horses during Competition

Research Article Antioxidant Status in Elite Three-Day Event Horses during Competition Oidtive Medicine nd Cellulr Longevity Volume 2012, Article ID 572090, 8 pges doi:10.1155/2012/572090 Reserch Article Antioidnt Sttus in Elite Three-Dy Event Horses during Competition Crey A. Willims 1

More information

Wonderland Road South from Southdale Road to Highway 402 DRAFT Assessment and Evaluation of Road Cross-Section Concepts

Wonderland Road South from Southdale Road to Highway 402 DRAFT Assessment and Evaluation of Road Cross-Section Concepts APPENDIX F 1 Socio-Economic 1.1 Property nd Access Impcts to properties Potentil impcts to ccess to existing residences/usinesses Totl numer of properties impcted Impcts to residences (e.g., displcement)

More information

Collettsville Road Race

Collettsville Road Race Merckx Time Tril April 7th 2017 4690 Collettsville School Dr. Collettsville, NC 28611 Get redy to strt your omnium weekend off with n evening TT. This course is the strt nd finish stretch of the Rod Rce

More information

Barbara Padalino, 1 Paola Zaccagnino, 2 and Pietro Celi 3,4. 1. Introduction

Barbara Padalino, 1 Paola Zaccagnino, 2 and Pietro Celi 3,4. 1. Introduction Veterinry Medicine Interntionl, rticle ID 87551, 8 pges http://dx.doi.org/1.1155/214/87551 Reserch rticle The Effect of Different Types of Physicl Exercise on the ehviourl nd Physiologicl Prmeters of Stndrdred

More information

Integration of modelling and monitoring to optimize network control: two case studies from Lisbon

Integration of modelling and monitoring to optimize network control: two case studies from Lisbon Aville online t www.sciencedirect.com ScienceDirect Procedi Engineering 70 ( 2014 ) 555 562 12th Interntionl Conference on Computing nd Control for the Wter Industry, CCWI2013 Integrtion of modelling nd

More information

The Pythagorean Theorem and Its Converse Is That Right?

The Pythagorean Theorem and Its Converse Is That Right? The Pythgoren Theorem nd Its Converse Is Tht Right? SUGGESTED LEARNING STRATEGIES: Activting Prior Knowledge, Mrking the Text, Shred Reding, Summrize/Prphrse/Retell ACTIVITY 3.6 How did Pythgors get theorem

More information

Regards, David C. Grossman, M.D., M.P.H. Chair, U.S. Preventive Services Task Force

Regards, David C. Grossman, M.D., M.P.H. Chair, U.S. Preventive Services Task Force Thnk you for your interest in the U.S. Preventive Services Tsk Force (USPSTF or Tsk Force), n independent, volunteer group of ntionl experts in prevention nd evidence-sed medicine. Our mission is to improve

More information

FREEWAY SYSTEM PROBLEMS AND DEFICIENCIES: PHYSICAL DESIGN, TRAFFIC SAFETY, AND TRAFFIC CONGESTION

FREEWAY SYSTEM PROBLEMS AND DEFICIENCIES: PHYSICAL DESIGN, TRAFFIC SAFETY, AND TRAFFIC CONGESTION Finl Drft s Approved by Technicl Subcommittee SEWRPC Plnning Report No. 7 A REGIONAL FREEWAY RECONSTRUCTION SYSTEM PLAN FOR SOUTHEASTERN WISCONSIN Chpter V FREEWAY SYSTEM PROBLEMS AND DEFICIENCIES: PHYSICAL

More information

No Gender Difference in Peak Performance in Ultra-Endurance Swimming Performance Analysis of the Zurich 12-h Swim from 1996 to 2010

No Gender Difference in Peak Performance in Ultra-Endurance Swimming Performance Analysis of the Zurich 12-h Swim from 1996 to 2010 Chinese Journal of Physiology 55(5): 346-351, 2012 DOI: 10.4077/CJP.2012.BAA053 No Gender Difference in Peak Performance in Ultra-Endurance Swimming Performance Analysis of the Zurich 12-h Swim from 1996

More information

Rules of Hockey including explanations. Effective from 1 January 2015

Rules of Hockey including explanations. Effective from 1 January 2015 Rules of Hockey including explntions Effective from 1 Jnury 2015 Copyright FIH 2014 The Interntionl Hockey Federtion Rue du Vlentin 61 CH 1004 Lusnne Switzerlnd Tel. : + 41 21 641 0606 Fx : + 41 21 641

More information

Daytime Habitat Selection by Resident Golden Eagles in Southern Idaho, USA

Daytime Habitat Selection by Resident Golden Eagles in Southern Idaho, USA Dytime Hbitt Selection by Resident Golden Egles in Southern Idho, USA Environmentl & Sttisticl Consultnts Chd LeBeu 1-04-14 Ryn Nielson, Eric Hllingstd, nd Dve Young cwlebeu@west-inc.com Introduction Risk

More information

Top-down regression of the avian oviduct during late oviposition in a small passerine bird

Top-down regression of the avian oviduct during late oviposition in a small passerine bird The Journl of Experimentl Biology 27, 263-268 Pulished y The Compny of Biologists 24 doi:1.1242/je.74 263 Top-down regression of the vin oviduct during lte oviposition in smll psserine ird T. D. Willims*

More information

VB-7253 Series. Application. Features. Applicable Literature

VB-7253 Series. Application. Features. Applicable Literature VB-7253 Series 1/2 to 2 Screwed NPT Stinless Steel Trim with Teflon Disc Stem Up Open, Two-Wy Vlves Generl Instructions Appliction VB-7253 series single set, stem down to close, two wy vlves control wter

More information

Responses of fish communities to sustained removals of perch (Perca fluviatilis)

Responses of fish communities to sustained removals of perch (Perca fluviatilis) Responses of fish communities to sustined removls of perch (Perc fluvitilis) SCIENCE FOR CONSERVATION Benjmin G. Ludgte nd Gerrd P. Closs Pulished y Deprtment of Conservtion PO Box -4 Wellington, New Zelnd

More information

A Step, Stride and Heading Determination for the Pedestrian Navigation System

A Step, Stride and Heading Determination for the Pedestrian Navigation System Journl of Globl Positioning Systems (2004) Vol. 3,. 1-2: 273-279 A Step, Stride nd Heding Determintion for the Pedestrin Nvigtion System Jeong Won Kim GNSS Technology Reserch Center, Chungnm Ntionl University,

More information

Report. Social Facilitation of Long-Lasting Memory Retrieval in Drosophila

Report. Social Facilitation of Long-Lasting Memory Retrieval in Drosophila Current iology 19, 1654 1659, Octoer 13, 2009 ª2009 Elsevier Ltd ll rights reserved DOI 116/j.cu.2009.08.017 Socil Fcilittion of Long-Lsting Memory Retrievl in Drosophil Report Mrie-nge Chud, 1,2,5 Guillume

More information

Response by anglers to a differential harvest regulation on three black bass species at Skiatook Lake, Oklahoma

Response by anglers to a differential harvest regulation on three black bass species at Skiatook Lake, Oklahoma 9 Response y nglers to differentil hrvest regultion on three lck ss species t Skitook Lke, Oklhom Jmes M. Long U.S. Geologicl Survey, Oklhom Coopertive Fish nd Wildlife Reserch Unit, Nturl Resource Ecology

More information

1. God 2. Family 3. Career

1. God 2. Family 3. Career Attributes for Success 1. Congenil, Likes People 2. Positive Outlook, Visionry 3. Cpble, Willing 4. Motivted, Committed 5. Sincere, Professionl 6. High Integrity 7. Fmily Oriented Benefits of n MK Creer...

More information

Workfit -SR, Dual Monitor Short Surface

Workfit -SR, Dual Monitor Short Surface User's Guide Workfit -SR, Dul Monitor Short Surfce Do not throw wy! Crdord locks needed for instlltion. Visit http://www.ergotron.com/workfi t-sr-instll for instlltion instructionl video. NOTE: 10 feet

More information

How To Demonstrate Guide E24 & E26

How To Demonstrate Guide E24 & E26 Pre-Demonstrtion Checklist Clen the entire mchine Chrge tteries Check squeegee ldes Bring proper rushes nd/or pds Clening cost nlysis Prepre quottion with lese pyments E24 & E26 literture nd fct sheet

More information

USA Field Hockey s Modifications to the 2017 FIH Rules of Hockey

USA Field Hockey s Modifications to the 2017 FIH Rules of Hockey USA Field Hockey hs estlished tht the Interntionl Hockey Federtion (FIH) will e the ultimte uthority for rules governing the sport of hockey in the United Sttes. This rule ook is reprinted under the uthority

More information

OFFSHORE USE OF FLOATING SHEERLEGS

OFFSHORE USE OF FLOATING SHEERLEGS Proceedings of the ASME 211 3th Interntionl Conference on Ocen, Offshore nd Arctic Engineering OMAE211 June 19-24, 211, Rotterdm, The Netherlnds OMAE211-49394 OFFSHORE USE OF FLOATING SHEERLEGS Vincent

More information

FOR OWNERS OF HORSE BUSINESSES

FOR OWNERS OF HORSE BUSINESSES W 371-B MANAGEMENT AND COST CONSIDERATIONS FOR OWNERS OF HORSE BUSINESSES Andrew P. Griffith, Assistnt Professor nd UT Extension Specilist Deprtment of Agriculturl nd Resource Economics BACKYARD VEGETABLES

More information

Rules of Hockey including explanations. Effective from 1 January 2017

Rules of Hockey including explanations. Effective from 1 January 2017 From 1 Jnury 2017 Rules of Hockey including explntions Effective from 1 Jnury 2017 Copyright FIH 2016 The Interntionl Hockey Federtion Rue du Vlentin 61 CH 1004 Lusnne Switzerlnd Tel. : + 41 21 641 0606

More information

Changes in transition times in Ironman Hawaii between 1998 and 2013

Changes in transition times in Ironman Hawaii between 1998 and 2013 Rüst et al. BMC Sports Science, Medicine, and Rehabilitation 2014, 6:37 RESEARCH ARTICLE Open Access Changes in transition times in Ironman Hawaii between 1998 and 2013 Christoph A Rüst 1, Thomas Rosemann

More information

Syngenta Soybean Insecticide Screening Study Beaumont, TX 2017

Syngenta Soybean Insecticide Screening Study Beaumont, TX 2017 I II III IV N Syngent Soyben Insecticide Screening Study Beumont, TX 2017 PLOT PLAN 20 1 LG 32 4 BX 19 8 GX 31 5 O 18 3 B 30 2 LGX 17 6 OX 29 7 G 16 1 LG 28 2 LGX 15 7 G 27 8 GX 14 3 B 26 6 OX 13 5 O 25

More information

Chapter 5. Triangles and Vectors

Chapter 5. Triangles and Vectors www.ck12.org Chpter 5. Tringles nd Vectors 5.3 The Lw of Sines Lerning Objectives Understnd how both forms of the Lw of Sines re obtined. Apply the Lw of Sines when you know two ngles nd non-included side

More information

Design and Calibration of Submerged Open Channel Flow Measurement Structures: Part 3 - Cutthroat Flumes

Design and Calibration of Submerged Open Channel Flow Measurement Structures: Part 3 - Cutthroat Flumes Uth Stte University DigitlCommons@USU Reports Uth Wter Reserch Lbortory Jnury 1967 Design nd Clibrtion of Submerged Open Chnnel Flow Mesurement Structures: Prt 3 - Cutthrot Flumes Gylord V. Skogerboe M.

More information

Available online at ScienceDirect. Procedia Materials Science 8 (2015 )

Available online at   ScienceDirect. Procedia Materials Science 8 (2015 ) Aville online t www.sciencedirect.com ScienceDirect Procedi Mterils Science 8 (2015 ) 914 923 Interntionl Congress of Science nd Technology of Metllurgy nd Mterils, SAM - CONAMET 2013 Effect of Het Input

More information

Study on Fish Migration through a Stone-Embedded Fish Passage Based on Preference

Study on Fish Migration through a Stone-Embedded Fish Passage Based on Preference Journl of Wter nd Environment Technology, Vol.13, No.1, 2015 Study on Fish Migrtion through Stone-Embedded Fish ssge Bsed on reference Rin FEBRINA 1) 2), Mshiko SEKINE 2), Hiroshi KANEMOTO 2), Koichi YAMAMOTO

More information

SWD Update. Elizabeth H. Beers Tree Fruit Research & Extension Center 1100 N. Western Ave. Wenatchee, Washington

SWD Update. Elizabeth H. Beers Tree Fruit Research & Extension Center 1100 N. Western Ave. Wenatchee, Washington SWD Updte Northwest Wholesle Orgnic Grower Meeting Cmpbell s Resort, Cheln, WA 5 Februry 213 9:3-1: m Elizbeth H. Beers Tree Fruit Reserch & Extension Center 11 N. Western Ave. Wentchee, Wshington US SWD

More information

The Effect of Stocking Density on Growth Parameters and Survival Rate of Rainbow Trout Alevins (Oncorhynchus mykiss)

The Effect of Stocking Density on Growth Parameters and Survival Rate of Rainbow Trout Alevins (Oncorhynchus mykiss) World Journl of Fish nd Mrine Sciences 4 (5): 480-485, 2012 ISSN 2078-4589 IDOSI Pulictions, 2012 DOI: 10.5829/idosi.wjfms.2012.04.05.6414 The Effect of Stocking Density on Growth Prmeters nd Survivl Rte

More information

bark bark bat bat Multiple Meaning Words: Kindergarten to Grade 2 More Teaching Tools at harsh sound made by a dog

bark bark bat bat Multiple Meaning Words: Kindergarten to Grade 2 More Teaching Tools at  harsh sound made by a dog the brk the brk bt bt hrsh sound mde by dog Mx, stop brking! outside cover of the trunks, brnches, nd roots of woody plnts The brk of this tree is very rough. club of wood or metl used to hit the bll in

More information

Report. Honolulu City Council. The Proposed Sale of City Housing Properties

Report. Honolulu City Council. The Proposed Sale of City Housing Properties Report t o m ~ t 4 rfl +r~ +kn L~J LI IC V 0 Honolulu City Council ~2c - C rn on The Proposed Sle of City Housing Properties Prepred By The Council Tsk Force to Review Utiliztion of All City Property (Tsk

More information

MAXIMUM CARDIAC PERFORMANCE OF RAINBOW TROUT (ONCORHYNCHUS MYKISS) AT TEMPERATURES APPROACHING THEIR UPPER LETHAL LIMIT

MAXIMUM CARDIAC PERFORMANCE OF RAINBOW TROUT (ONCORHYNCHUS MYKISS) AT TEMPERATURES APPROACHING THEIR UPPER LETHAL LIMIT The Journl of Experimentl Biology 199, 663 672 (1996) Printed in Gret Britin The Compny of Biologists Limited 1996 JEB0137 663 MAXIMUM CARDIAC PERFORMANCE OF RAINBOW TROUT (ONCORHYNCHUS MYKISS) AT TEMPERATURES

More information

PROTECTION FROM HAND-ARM TRANSMITTED VIBRATION USING ANTIVIBRATON GLOVES

PROTECTION FROM HAND-ARM TRANSMITTED VIBRATION USING ANTIVIBRATON GLOVES PROTECTION FROM HAND-ARM TRANSMITTED VIBRATION USING ANTIVIBRATON GLOVES VLADO GOGLIA IGOR ĐUKIĆ JOSIP ŢGELA Abstrct There re vrious wys of reducing the helth-hzrds of hnd-trnsmitted vibrtion. One of the

More information

Improved Race Times in Marathoners Older than 75 Years in the Last 25 Years in the World s Largest Marathons

Improved Race Times in Marathoners Older than 75 Years in the Last 25 Years in the World s Largest Marathons Chinese Journal of Physiology 9(3): 139-147, 216 139 DOI: 1.477/CJP.216.BAE382 Improved Race Times in Marathoners Older than 7 s in the Last s in the World s Largest Marathons Baschir Ahmadyar, Thomas

More information

Differences in Participation and Performance Trends in Age Group Half and Full Marathoners

Differences in Participation and Performance Trends in Age Group Half and Full Marathoners Chinese Journal of Physiology 57(4): 29-219, 214 29 DOI: 1.477/CJP.214.BAC219 Differences in Participation and Performance Trends in Age Group Half and Full Marathoners Danny Anthony 1, Christoph A. Rüst

More information

Performance in Olympic triathlon: changes in performance of elite female and male triathletes in the ITU World Triathlon Series from 2009 to 2012

Performance in Olympic triathlon: changes in performance of elite female and male triathletes in the ITU World Triathlon Series from 2009 to 2012 Rüst et al. SpringerPlus 213, 2:68 a SpringerOpen Journal RESEARCH Open Access Performance in Olympic triathlon: changes in performance of elite female and male triathletes in the ITU World Triathlon Series

More information

Name Class Date SAMPLE. Complete the missing numbers in the sequences below. 753, ,982. The area of the shape is approximately cm 2

Name Class Date SAMPLE. Complete the missing numbers in the sequences below. 753, ,982. The area of the shape is approximately cm 2 End of term: TEST A You will need penil. Yer 5 Nme Clss Dte 1 2 Complete the missing numers in the sequenes elow. 200 3926 4926 400 500 700 7926 753,982 553,982 Estimte the re of the shpe elow. The re

More information

United States Patent (19)

United States Patent (19) United Sttes Ptent (19) Mobley (11) 45) Ptent Number: Dte of Ptent: Nov. 19, 1991 (54) BICYCLE RACK FORMOUNTING ON A VAN 76 Inventor: Mrk H. Mobley, P.O. Box 636, Norris, Tenn. 37828 21 Appl. No.: 436,492

More information

Quantifying Efficacy and Avoidance Behavior by Tawny Mole Crickets (Orthoptera: Gryllotalpidae: Scapteriscus vicinus) to Three Synthetic Insecticides

Quantifying Efficacy and Avoidance Behavior by Tawny Mole Crickets (Orthoptera: Gryllotalpidae: Scapteriscus vicinus) to Three Synthetic Insecticides Quntifying Efficcy nd Avoidnce Behvior y Twny Mole Crickets (Orthopter: Gryllotlpide: Scpteriscus vicinus) to Three Synthetic Insecticides Authors: D. E. Silcox, C. E. Sorenson, nd R. L. Brndenurg Source:

More information

High rate of prey consumption in a small predatory fish on coral reefs

High rate of prey consumption in a small predatory fish on coral reefs Corl Reefs (1) 31:99 918 DOI 1.17/s338-1-894-z High rte of prey consumption in smll predtory fish on corl reefs W. E. Feeney O. M. Lönnstedt Y. Bosiger J. Mrtin G. P. Jones R. J. Rowe M. I. McCormick Received:

More information

Chapter 4 Group of Volunteers

Chapter 4 Group of Volunteers CHAPTER 4 SAFETY CLEARANCE, FREEBOARD AND DRAUGHT MARKS 4-1 GENERAL 4-1.1 This chpter specifies the minimum freebord for inlnd wterwy vessels. It lso contins requirements concerning the indiction of the

More information

Sparse, decorrelated odor coding in the mushroom body enhances learned odor discrimination

Sparse, decorrelated odor coding in the mushroom body enhances learned odor discrimination Sprse, decorrelted odor coding in the mushroom ody enhnces lerned odor discrimintion Andrew C. Lin, Alexei Bygrve, Alix de Clignon, Tzumin Lee, nd Gero Miesenöck Supplementry Mteril R64C08-GAL4 (γ) R35B12-GAL4

More information

UNCORRECTED PROOF. Materials and methods

UNCORRECTED PROOF. Materials and methods Exp rin Res (26) DOI.7/s22-6-456-2 3 RESEARCH ARTICLE 4 Giovnni Mirell Pierpolo Pni 5 Mrtin Pre Stefno Ferrin 6 Inhiitory control of reching movements in humns 7 8 Received: August 25 / Accepted: Mrch

More information

Lesson 8: Application Technology

Lesson 8: Application Technology The type of ppliction equipment used must suit the type of ppliction. In this module, you ll lern the prts of the most common types of ppliction equipment used by ssistnt pplictors, s well s how to properly

More information